C. W. Andrews, A description of new species of zeuglodont and of leathery turtle from the Eocene of southern Nigeria, Proceedings of the Zoological Society of London, vol.22, pp.309-319, 1920.

P. Antoine, Phylog enie et evolution des Elasmotheriina (Mammalia, Rhinocerotidae). M emoires du Mus eum National d'Histoire Naturelle, vol.188, pp.1-358, 2002.

P. Antoine, L. Marivaux, D. A. Croft, G. Billet, M. Ganerød et al., Middle Eocene rodents from Peruvian Amazonia reveal the pattern and timing of caviomorph origins and biogeography, Proceedings of the Royal Society. Series B, vol.279, pp.1319-1326, 2012.

S. Bajpai and J. G. Thewissen, Protocetid cetaceans (Mammalia) from the Eocene of India, Palaeontologia Electronica, vol.17, issue.3, pp.1-19, 2014.

J. Benoit, S. Adnet, E. El-mabrouk, H. Khayati, M. Ben-haj-ali et al., Cranial remain from Tunisia provides new clues for the origin and evolution of Sirenia (Mammalia, Afrotheria) in Africa, PloS ONE, vol.8, issue.1, pp.1-9, 2013.
URL : https://hal.archives-ouvertes.fr/hal-00807308

G. Bianucci and P. D. Gingerich, Aegyptocetus tarfa,n . gen. et sp. (Mammalia, Cetacea), from the middle Eocene of Egypt: Clinorhynchy, olfaction, and hearing in a protocetid whale, Journal of Vertebrate Paleontology, vol.31, pp.1173-1188, 2011.

K. Bremer, Branch support and tree stability, Cladistics, vol.10, pp.295-304, 1994.

H. Cappetta and M. Traverse, Une riche faune de s elaciens dans le bassin a phosphate de Kpogam e-Hahoto e ( Eoc ene moyen du Togo): note pr eliminaire et pr ecisions sur la structure et l'âge du gisement, Geobios, vol.21, pp.359-365, 1988.

A. L. Cione, M. A. Cozzuol, M. T. Dozo, and C. A. Hospitaleche, Marine vertebrate assemblages in the southwest Atlantic during Miocene, Biological Journal of the Linnean Society, vol.103, pp.423-440, 2011.

W. B. Dames, Ueber Zeuglodonten aus Aegypten und die Beziehungen der Archaeoceten zu den ? ubrigen Cetaceen, Geologische und Pala ?ontologische Abhandlungen. Jena, vol.5, pp.189-222, 1894.

V. De-buffr-enil, W. Dabin, and L. Zylberberg, Histology and growth of the cetacean petro-tympanic bone complex, Journal of Zoology, vol.262, pp.371-381, 2004.

R. Dehm and T. Oettingen-spielberg, Pal? aontologische und geologische Untersuchungen im Terti? ar von Pakistan. 2. Die mitteleoc? anen Sa? ugetierre von Ganda Kas bei Basal in Nordwest Pakistan, Abhandlungen der Bayerischen Akademie der Wissenschaften, vol.91, pp.1-54, 1958.

F. B. De-oliveira, E. C. Molina, and G. Marroig, Paleogeography of the South Atlantic: a route for primates and rodents into the New World? Pp. 55-68 in, South American Primates: comparative perspectives in the study of behavior, ecology, and conservation, 2009.

C. Springer,

E. G. Ekdale, A. Berta, and T. A. Dem-er-e, The comparative osteology of the petrotympanic complex (ear region) of extant baleen whales (Cetacea: Mysticeti), PloS ONE, vol.6, issue.6, pp.21311-21312, 2011.

R. M. Feldmann, K. L. Bice, C. Schweitzer-hopkins, E. W. Salva, and K. Pickford, Decapod crustaceans from the Eocene Castle Hayne limestone, North Carolina: Paleoceanographic implications, Journal of Paleontology, vol.48, pp.1-28, 1998.

W. H. Flower, On the arrangement of the orders and families of existing Mammalia, Proceedings of the Zoological Society of London, vol.1883, pp.178-186, 1883.

E. Fraas, Neue Zeuglodonten aus dem unteren Mitteleoc? an vom Mokattam bei Cairo. Mittheilungen aus dem Ko ?niglichen Naturalien-Cabinet in Stuttgart, vol.27, pp.1-24, 1904.

F. C. Fraser and P. E. Purves, Anatomy and function of the cetacean ear, Proceedings of the Royal Society of London. Series B, vol.152, pp.62-77, 1960.

F. C. Fraser and P. E. Purves, Hearing in cetaceans, evolution of the accessory air sacs and the structure and function, 1960.

M. J. Mourlam and M. J. Orliac-museum, Natural History), vol.7, pp.1-140

R. E. Fordyce, Simocetus rayi (Odonceti: Simocetidae, New family): A bizarre new archaic Oligocene dolphin from the Eastern North Pacific, Smithsonian Contributions to Paleobiology, vol.93, pp.185-222, 2002.

A. Galatius and P. E. Gol'din, Geographic variation of skeletal ontogeny and skull shape in the harbour porpoise (Phocoena phocoena), Canadian Journal of Zoology, vol.89, pp.869-879, 2011.

P. J. Gannon, A. R. Eden, and J. T. Laitman, The subarcuate fossa and cerebellum of extant Primates: comparative study of a skull-brain interface, American Journal of Physical Anthropology, vol.77, pp.143-164, 1988.

J. H. Geisler and Z. X. Luo, Relationships of Cetacea to terrestrial ungulates and the evolution of cranial vasculature in Cete, pp.163-212, 1998.

J. H. Geisler, A. E. Sanders, and Z. X. Luo, A new protocetid whale (Cetacea: Archaeoceti) from the late middle Eocene of South Carolina, American Museum Novitates, vol.3480, pp.1-65, 2005.

R. W. Gibbes, Description of the teeth of a new fossil animal found in the Green Sand of South Carolina, Proceedings of the Academy of Natural Sciences, vol.2, pp.254-256, 1845.

J. W. Gilpatrick and W. L. Jr-&-perryman, Geographic variation in external morphology of North Pacific and southern Hemisphere blue whales (Balaenoptera musculus), Journal of Cetacean Research and Management, vol.10, pp.9-21, 2008.

P. D. Gingerich and H. Cappetta, A new archaeocete and other marine mammals (Cetacea and Sirenia) from lower middle Eocene phosphate deposits of Togo, Journal of Paleontology, vol.88, pp.109-129, 2014.

P. D. Gingerich and D. E. Russell, Pakicetus inachus,a new archaeocete (Mammalia, Cetacea) from the earlymiddle Eocene Kuldana formation of Kohat (Pakistan). Contributions from the Museum of Paleontology, vol.25, pp.235-246, 1981.

P. D. Gingerich and S. Zouhri, New fauna of archaeocete whales (Mammalia, cetacean) from the Bartonian middle Eocene of southern Morocco, Journal of African Sciences, vol.111, pp.273-286, 2015.

P. D. Gingerich, S. M. Raza, M. Arif, M. Anwar, and X. Zhou, New whale from the Eocene of Pakistan and the origin of cetacean swimming, Nature, vol.368, pp.844-847, 1994.

P. D. Gingerich, M. Arif, and W. C. Clyde, New archaeocetes (Mammalia, Cetacea) from the middle Eocene Domanda formation of the Sulaiman range, Punjab (Pakistan), Contributions from the Museum of Paleontology, University of Michigan, vol.29, pp.291-330, 1995.

P. D. Gingerich, M. Arif, M. A. Bhatti, H. A. Raza, and S. M. Raza, Protosiren and Babiacetus (Mammalia, Sirenia and Cetacea) from the middle Eocene Drazinda formation, Sulaiman range, Contributions from the Museum of Paleontology, vol.29, pp.331-357, 1995.

P. D. Gingerich, M. Haq, W. Koenigswald, W. J. Sanders, B. H. Smith et al., New protocetid whale from the middle Eocene of Pakistan: birth on land, precocial development and sexual dimorphism, PloS ONE, vol.4, issue.2, pp.1-20, 2009.

P. E. Gol'din and E. Gladilina, Small dolphins in a small sea: age, growth and life-history aspects of the Black Sea common bottlenose dolphin Tursiops truncatus, Aquatic Biology, vol.23, pp.159-166, 2015.

P. E. Gol'din and E. Zvonok, Basilotritus uheni, a new cetacean (Cetacea, Basilosauridae) from the late middle Eocene of Eastern Europe, Journal of Paleontology, vol.87, pp.254-268, 2013.

J. E. Gray, On the cetaceous animals, Zoology of the Voyage of H, pp.13-53, 1846.

M. S. Erebus and . Terror, , vol.1

R. Harlan, Notice of fossil bones found in the Tertiary formation of the State of Louisiana, Transactions of the American Philosophical Society, vol.4, pp.397-403, 1834.

L. Hautier, R. Sarr, F. Lihoreau, R. Tabuce, and P. M. Hameh, First record of the family Protocetidae in the Lutetian of Senegal (West Africa), Palaeovertebrata, vol.38, issue.2, pp.1-7, 2014.
URL : https://hal.archives-ouvertes.fr/hal-01923037

D. Huchon and E. J. Douzery, From the Old World to the New World: a molecular chronicle of the phylogeny and biogeography of hystricognath rodents, Molecular Phylogenetics and Evolution, vol.20, pp.238-251, 2001.

R. C. Hulbert, . Jr, R. M. Petkewich, G. A. Bishop, D. Bukry et al., A new middle Eocene protocetid whale (Mammalia: Cetacea: Archaeoceti) and associated biota from Georgia, Journal of Paleontology, vol.72, pp.907-927, 1998.

R. Kellogg, A review of the Archaeoceti. Canergie Institution of Washington, vol.482, pp.1-366, 1936.

K. Kumar and A. Sahni, Remingtonocetus harudiensis, new combination, a middle Eocene archaeocete (Mammalia, Cetacea) from western Kutch, India. Journal of Vertebrate Paleontology, vol.6, pp.326-349, 1986.

D. R. Ketten, The Cetacean ear: form, frequency and evolution, Marine Mammal Sensory Systems, pp.53-75, 1992.

W. C. Lancaster, The middle ear of the Archaeoceti, Journal of Vertebrate Paleontology, vol.10, pp.117-127, 1990.

C. Linnaeus, Systema naturae per regna tria naturae, secundum classes, ordines, genera species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata), Systema Naturae, vol.824, p.pp, 1758.

Z. X. Luo, Homology and transformation of cetacean ectotympanic structures, The emergence of whales: evolutionary patterns in the origin of Cetacea, pp.269-301, 1998.

Z. X. Luo and E. R. Eastman, Petrosal and inner ear of a squalodontid whale: Implications for evolution of hearing in Odontocetes, Journal of Vertebrate Paleontology, vol.15, pp.431-442, 1995.

Z. X. Luo and P. D. Gingerich, Terrestrial Mesonychia to aquatic Cetacea: transformation of the basicranium and evolution of hearing in whales, vol.31, pp.1-98, 1999.

Z. X. Luo and K. Marsh, Petrosal (periotic) and inner ear of a Pliocene kogiine whale (Kogiinae, Odontoceti): implications on relationships and hearing evolution of toothed whales, Journal of Vertebrate Paleontology, vol.16, pp.328-348, 1996.

M. Mart-inez-c-aceres, O. Lambert, and C. D. Muizon, The anatomy and phylogenetic affinities of Cynthiacetus peruvianus, a large Dorudon-like basilosaurid (Cetacea, Mammalia) from the late Eocene of Peru, Geodiversitas, vol.39, issue.1, pp.7-163, 2017.

S. A. Mcleod and L. G. Barnes, A new genus and species of Eocene protocetid archaeocete whale (Mammalia, Cetacea) from the Atlantic Coastal Plain, Natural History Museum of Los Angeles County, vol.41, pp.73-98, 2008.

J. G. Mead and R. E. Fordyce, The therian skull: A lexicon with emphasis on the odontocetes, Smithsonian Contributions to Zoology, vol.627, pp.1-248, 2009.

G. Montagu, Description of a species of Delphinus, which appears to be new, Memoirs of the Wernerian Natural History Society, vol.3, pp.75-82, 1821.

M. J. Mourlam and M. J. Orliac, 3D models related to the publication: Protocetid (Cetacea, Artiodactyla) bullae and petrosals from the Middle Eocene locality of Kpogam e, Togo: new insights into the early history of cetacean hearing, MorphoMuseuM, vol.3, issue.1, 2017.

S. Nummela, T. W?-agar, S. Hemil?-a, and T. Reuter, Scaling of the cetacean middle ear, Hearing Research, vol.133, pp.71-81, 1999.

S. Nummela, J. G. Thewissen, S. Bajpai, S. T. Hussain, and K. Kumar, Eocene evolution of whale hearing, Nature, vol.430, pp.776-778, 2004.

S. Nummela, S. T. Hussain, and J. G. Thewissen, Cranial anatomy of Pakicetidae (Cetacea, Mammalia), Journal of Vertebrate Paleontology, vol.26, pp.746-759, 2006.

S. Nummela, J. G. Thewissen, S. Bajpai, S. T. Hussain, and K. Kumar, Sound transmission in archaic and modern whales: anatomical adaptations for underwater hearing, The Anatomical Record, vol.290, pp.716-733, 2007.

M. A. O'leary, An anatomical and phylogenetic study of the osteology of the petrosal of extant and extinct artiodactylans (Mammalia) and relatives, Bulletin of the American Museum of Natural History, vol.335, pp.1-226, 2010.

M. J. Orliac and M. A. O'leary, Comparative anatomy of the petrosal bone of Dichobunoids, early members of Artiodactylamorpha (Mammalia), Journal of Mammalian Evolution, vol.21, pp.299-320, 2014.
URL : https://hal.archives-ouvertes.fr/hal-01897864

M. J. Orliac, P. Antoine, and S. Ducrocq, Phylogenetic relationships of the Suidae (Mammalia, Cetartiodactyla): new insights on the relationships within Suoidea, Zoologica Scripta, vol.39, pp.315-330, 2010.

M. J. Orliac, P. Antoine, A. Charruault, S. Hervet, F. Prodeo et al., Specialization for amphibiosis in Brachyodus onoideus (Artiodactyla, Hippopotamoidea) from the Early Miocene of France, Swiss Journal of Geosciences, vol.106, pp.265-278, 2013.
URL : https://hal.archives-ouvertes.fr/hal-01902587

R. Owen, Description of teeth and portions of jaws of two extinct Anthracotherioid quadrupeds (Hyopotamus vectianus and Hyop. bovinus) discovered by the Marchioness of Hastings in the Eocene deposits on the NW coast of the Isle of Wight: with an attempt to develop Cuvier's idea of the classification of Pachyderms by the number of their toes, Quarterly Journal of the Geological Society of London, vol.4, pp.103-141, 1848.

W. F. Perrin, Variation of spotted and spinner porpoise (genus Stenella) in the eastern Pacific and Hawaii, Bulletin of the Scripps Institution of Oceanography, vol.21, pp.1-206, 1975.

W. F. Perrin, G. D. Schnell, D. J. Hough, J. W. Gilpatrick, and J. V. Jr-&-kashiwada, Reexamination of geographic variation in cranial morphology of the pantropical spotted dolphin, Stenella attenuate, in the eastern Pacific, Fishery Bulletin, vol.92, pp.324-346, 1994.

B. G. Pritchard, On the discovery of a fossil whale in the older Tertiaries of Toquay, Victoria. The Victorian Naturalist, vol.55, pp.151-159, 1939.

W. G. Ridewood, Observations on the skull in foetal specimens of whales of the genera Megaptera and Balaenoptera, Philosophical Transactions of the Royal Society of London. Series B, vol.121, pp.209-272, 1922.

A. Sahni and V. P. Mishra, Lower Tertiary vertebrates from western India. Monograph of the Paleontological Society of India, vol.3, pp.1-48, 1975.

N. Saitou and M. Nei, The neighbor-joining method: a new method for reconstructing phylogenetic trees, Molecular Biology and Evolution, vol.4, pp.406-425, 1987.

R. Sassu and B. Cozzi, The external and middle ear of the striped dolphin Stenella coeruleoalba (Meyen 1833). Anatomia, Histologia, Embryologia, vol.36, pp.197-201, 2007.

W. Segall, Morphological parallelisms of the bulla and auditory ossicles in some Insectivores and Marsupials, Fieldiana Zoology, vol.51, issue.16, pp.169-205, 1970.

P. H. Sneath and R. R. Sokal, Numerical taxonomy: the principles and practice of numerical classification, vol.573, p.pp, 1973.

E. Stromer, Die Archaeoceti des ? Agyptischen Eoz? ans. Beitra ?ge zur Pala ?ontologie von O ? sterreich, Mitteilungen des Geologischen und Pala ?ontologischen Institutes des Universita ?t Wien, vol.21, pp.106-178, 1908.

D. L. Swofford, . Associates, M. A. Sunderland, J. G. Thewissen, and S. Bajpai, New skeletal material of Andrewsiphius and Kutchicetus, two Eocenecetaceans from India, Journal of Paleontology, vol.83, pp.635-663, 2002.

A. N. Trivedy and P. P. Satsangi, A new archaeocete (whale) from the Eocene of India. Abstracts of 27 th International Geological Congress, vol.1, pp.322-323, 1984.

M. D. Uhen, New species of protocetid archaeocete whale, Eocetus wardii (Mammalia: Cetacea) from the middle Eocene of North Carolina, Journal of Paleontology, vol.73, pp.512-528, 1999.

M. D. Uhen, Form, function, and anatomy of Dorudon atrox (Mammalia, Cetacea): an archaeocete from the middle to late Eocene of Egypt. The University of Michigan Museum of Paleontology, Papers on Paleontology, vol.34, pp.1-222, 2004.

M. D. Uhen, A new genus and species of archaeocete whale from Mississippi, Southeastern Geology, vol.43, issue.3, pp.157-172, 2005.

M. D. Uhen, New protocetid whales from Alabama and Mississippi, and a new cetacean clade, Pelagiceti, Journal of Vertebrate Paleontology, vol.28, pp.589-593, 2008.

M. D. Uhen, New specimens of Protocetidae (Mammalia, Cetacea) from New Jersey and South Carolina, Journal of Vertebrate Paleontology, vol.34, pp.211-219, 2014.

M. D. Uhen, N. D. Pyenson, T. J. Devries, M. Urbina, P. R. Renne et al., The auditory bulla in some fossil mammals, with a general introduction to this region of the skull, Bulletin of the American Museum of Natural History, vol.85, pp.1-352, 1931.

J. R. Wible, Cranial circulation and relationships of the Colugo Cynocephalus (Dermoptera, Mammalia), American Museum Novitates, vol.3072, pp.1-27, 1993.

J. R. Wible, On the cranial osteology of the shorttailed opossum Monodelphis brevicaudata (Didelphidae, Marsupialia), vol.72, pp.137-202, 2003.

E. M. Williams, Synopsis of the earliest Cetaceans: Pakicetidae, Ambulocetidae, Remingtonocetidae and Protocetidae, The Emergence of whales: evolutionary patterns in the origin of Cetacea, pp.1-28, 1998.

D. S. Houser, D. A. Helweg, and P. W. Moore, A Bandpass filterbank model of auditory sensitivity in the humpback whale, Aquat. Mamm, vol.27, pp.82-91, 2001.

C. Erbe, Hearing Abilities of Baleen Whales (DRDC Atlantic, 2002.

S. E. Parks, D. R. Ketten, J. T. O'malley, A. , and J. , Anatomical predictions of hearing in the North Atlantic right whale, Anat. Rec. (Hoboken), vol.290, pp.734-744, 2007.

S. Hemil?-a, S. Nummela, and T. Reuter, Modeling whale audiograms: effects of bone mass on high-frequency hearing, Hear. Res, vol.151, pp.221-226, 2001.

D. R. Ketten, Marine mammal auditory systems: A summary of audiometric and anatomical data and implications for underwater acoustic impacts, Polarforschung, vol.72, pp.79-92, 2004.

V. V. Popov and A. Y. Supin, Analysis of auditory information in the brains of cetaceans, Neurosci. Behav. Physiol, vol.37, pp.285-291, 2007.

J. G. Thewissen, S. I. Madar, and S. T. Hussain, Ambulocetus natans, an Eocene cetacean (Mammalia) from Pakistan, CFS Cour. Forschungsinstitut Senckenb, vol.191, pp.1-86, 1996.

S. Nummela, J. G. Thewissen, S. Bajpai, S. T. Hussain, and K. Kumar, Eocene evolution of whale hearing, Nature, vol.430, pp.776-778, 2004.

S. Nummela, J. G. Thewissen, S. Bajpai, T. Hussain, and K. Kumar, Sound transmission in archaic and modern whales: anatomical adaptations for underwater hearing, Anat. Rec. (Hoboken), vol.290, pp.716-733, 2007.

M. D. Uhen, Form, Function, and Anatomy of Dorudon atrox (Mammalia, Cetacea): An Archaeocete from the Middle to Late Eocene of Egypt, Papers on Paleontology, vol.34, 2003.

E. G. Ekdale and R. A. Racicot, Anatomical evidence for low frequency sensitivity in an archaeocete whale: comparison of the inner ear of Zygorhiza kochii with that of crown Mysticeti, J. Anat, vol.226, pp.22-39, 2015.

T. Park, E. M. Fitzgerald, and A. R. Evans, Ultrasonic hearing and echolocation in the earliest toothed whales, Biol. Lett, vol.12, pp.1-4, 2016.

T. Park, A. R. Evans, S. J. Gallagher, and E. M. Fitzgerald, Lowfrequency hearing preceded the evolution of giant body size and filter feeding in baleen whales, Proc. Biol. Sci, vol.284, 2017.

D. R. Ketten, The cetacean ear: form, frequency, and evolution. In Marine Mammal Sensory Systems, pp.53-75, 1992.

M. Churchill, M. Martinez-caceres, C. De-muizon, J. Mnieckowski, and J. H. Geisler, The Origin of High-Frequency Hearing in Whales, Curr. Biol, vol.26, pp.2144-2149, 2016.
URL : https://hal.archives-ouvertes.fr/hal-02612270

M. J. Mourlam and M. J. Orliac, Protocetid (Cetacea, Artiodactyla) bullae and petrosals from the Middle Eocene locality of Kpogam e, Togo: new insights into the early history of cetacean hearing, J. Syst. Palaeontol, 2017.

G. Fleischer, Hearing in extinct cetaceans as determined by cochlear structure, J. Paleontol, vol.50, pp.133-152, 1976.

E. G. Ekdale, Morphological variation among the inner ears of extinct and extant baleen whales (Cetacea: Mysticeti), J. Morphol, vol.277, pp.1599-1615, 2016.

, Current Biology, vol.27, pp.1-6, 20175-06-19.

, Please cite this article in press as: Mourlam and Orliac, Infrasonic and Ultrasonic Hearing Evolved after the Emergence of Modern Whales, Current Biology, 2017.

C. Origine5, S. )-resoo&lt;-c, (. Oo, . Ocar, . Ogam et al., GamLA) PrO<-mahalanobis(ctrPro, Origine5, S) PrCar<-mahalanobis(ctrPro,Car, S) PrGam<-mahalanobis(ctrPro,Gam,S) PrPr<-mahalanobis(ctrPro, ctrPro, S) PrBa<-mahalanobis(ctrPro, ctrBa, S) PrAr<-mahalanobis(ctrPro, ctrAr, S) PrMy<-mahalanobis(ctrPro, ctrMy, S) PrOd<-mahalanobis(ctrPro, ctrOd, S) PrPel<-mahalanobis(ctrPro, ctrPel, S) PrLA<-mahalanobis(ctrPro, ctrLA, S) resPrPr<-c(PrO,PrCar,PrGam,PrPr,PrBa,PrAr,PrMy,PrOd,PrPel,PrLA) BaO<-mahalanobis(ctrBa, Origine5, S) BaCar<-mahalanobis(ctrBa,Car, S) BaGam<-mahalanobis(ctrBa,Gam,S) BaPr<-mahalanobis(ctrBa, ctrPro, S) BaBa<-mahalanobis(ctrBa, ctrBa, S) BaAr<-mahalanobis(ctrBa, ctrAr, S) BaMy<-mahalanobis(ctrBa, ctrMy, S) BaOd<-mahalanobis(ctrBa, ctrOd, S) BaPel<-mahalanobis(ctrBa, ctrPel, S) BaLA<-mahalanobis(ctrBa, ctrLA, S) resBaBa<-c, p.7, 2017.

, Please cite this article in press as: Mourlam and Orliac, Infrasonic and Ultrasonic Hearing Evolved after the Emergence of Modern Whales, Current Biology, 2017.

J. Thewissen and S. T. Hussain, Origin of underwater hearing in whales, Nature, vol.361, pp.444-445, 1993.

S. Nummela, J. Thewissen, S. Bajpai, S. T. Hussain, and K. Kumar, Eocene evolution of whale hearing, Nature, vol.430, pp.776-778, 2004.

S. Nummela, J. Thewissen, S. Bajpai, S. T. Hussain, and K. Kumar, Sound transmission in archaic and modern whales: anatomical adaptations for royalsocietypublishing.org/journal/rspb, Proc. R. Soc. B, vol.286, p.20191417, 2007.

, Behav, vol.150, pp.273-284

J. G. Mccormick, E. G. Wever, J. Palin, and S. H. Ridgway, Sound Conduction in the Dolphin Ear, J. Acoust. Soc. Am, vol.48, pp.1418-1428, 1970.

J. G. Mead and R. E. Fordyce, The therian skull : a lexicon with emphasis on the odontocetes, Smithson. Contrib. to Zool, pp.1-249, 2009.

A. R. Møller, Hearing: Anatomy, Physiology, and Disorders of the Auditory System, 2006.

T. A. Mooney, M. Yamato, and B. K. Branstetter, Hearing in Cetaceans: From Natural History to Experimental Biology, Adv. Mar. Biol, vol.63, pp.197-246, 2012.

T. A. Mooney, W. C. Yang, H. Yu, D. R. Ketten, and J. , Hearing abilities and sound reception of broadband sounds in an adult Risso's dolphin (Grampus griseus), J. Comp. Physiol. A, vol.201, pp.751-761, 2015.

M. J. Mourlam and M. J. Orliac, Infrasonic and Ultrasonic Hearing Evolved after the Emergence of Modern Whales, Curr. Biol, vol.27, pp.1776-1781, 2017.
URL : https://hal.archives-ouvertes.fr/hal-01894431

M. J. Mourlam and M. J. Orliac, Protocetid (Cetacea, Artiodactyla) bullae and petrosals from the middle Eocene locality of Kpogamé, Togo: new insights into the early history of cetacean hearing, J. Syst. Palaeontol, vol.16, pp.621-644, 2018.

M. J. Mourlam and M. J. Orliac, Early Evolution of the Ossicular Chain in Cetacea: Into the Middle Ear Gears of a Semi-Aquatic Protocetid Whale, vol.286, 2019.
URL : https://hal.archives-ouvertes.fr/hal-02367801

K. S. Norris, The evolution of acoustic mechanisms in odontocete cetaceans, pp.297-324, 1968.

K. S. Norris and G. W. Harvey, Sound transmission in the porpoise head, J. Acoust. Soc. Am, vol.56, pp.659-664, 1974.

S. Nummela, Hearing in Aquatic Mammals. Sensory Evolution on the Threshold Adaptations in Secondarily Aquatic Vertebrates, pp.210-224, 2008.

S. Nummela, T. Reuter, S. Hemilä, P. Holmberg, and P. Paukku, The anatomy of the killer whale middle ear (Orcinus orca), Hear. Res, vol.133, pp.61-70, 1999.

S. Nummela and J. G. Thewissen, The Physics of Sound in Air and Water. Sensory Evolution on the Threshold Adaptations in Secondarily Aquatic Vertebrates, pp.174-181, 2008.

S. Nummela, J. G. Thewissen, S. Bajpai, S. T. Hussain, and K. Kumar, Eocene evolution of whale hearing, Nature, vol.430, pp.776-778, 2004.

S. Nummela, J. G. Thewissen, S. Bajpai, S. T. Hussain, and K. Kumar, Sound transmission in archaic and modern whales: Anatomical adaptations for underwater hearing, Anat. Rec, vol.290, pp.716-733, 2007.

S. Nummela, T. Wägar, S. Hemilä, and T. Reuter, Scaling of the cetacean middle ear, Hear. Res, vol.133, pp.71-81, 1999.

S. Nummela and M. Yamato, Hearing. Encyclopedia of Marine Mammals, pp.462-470, 2018.

R. S. Payne and S. Mcvay, Songs of Humpback Whales. Science (80-. ), vol.173, pp.585-597, 1971.

V. V. Popov and A. Y. Supin, Analysis of auditory information in the brains of cetaceans, Neurosci. Behav. Physiol, vol.37, pp.285-291, 2007.

V. V. Popov, A. Y. Supin, D. I. Nechaev, A. A. Lemazina, and E. V. Sysueva, Position of an acoustic window in a beluga whale: Computation based on auditory evoked potential latencies, J. Acoust. Soc. Am, vol.145, pp.3578-3585, 2019.

V. V. Popov, E. V. Sysueva, D. I. Nechaev, A. A. Lemazina, and A. Y. Supin, Auditory sensitivity to local stimulation of the head surface in a beluga whale (Delphinapterus leucas), J. Acoust. Soc. Am, vol.140, pp.1218-1226, 2016.

M. A. Rauschmann, S. Huggenberger, L. S. Kossatz, and H. A. Oelschläger, Head morphology in perinatal dolphins: A window into phylogeny and ontogeny, J. Morphol, vol.267, pp.1295-1315, 2006.

S. J. Rehorek, R. Stimmelmayr, J. C. George, R. Suydam, D. M. Mcburney et al., Structure of the external auditory meatus of the Bowhead whale (Balaena mysticetus) and its relation to their seasonal migration, J. Anat, vol.234, pp.201-215, 2019.

L. E. Rendell and H. Whitehead, Vocal clans in sperm whales (<i>Physeter macrocephalus<i/>), Proc. R. Soc. London. Ser. B Biol. Sci, vol.270, pp.225-231, 2003.

F. W. Reysenbach-de-haan, Hearing in whales, Acta Otolaryngol. Suppl, vol.134, pp.1-114, 1957.

W. G. Ridewood, Observations on the Skull in Foetal Specimens of Whales of the Genera Megaptera and Balaenoptera, Philos. Trans. R. Soc. London. Ser. B, vol.211, pp.209-272, 1922.

M. R. Rossi-santos, The Spreading Message in a Noisy World: Song Behavior and Acoustic Ecology in Humpback Whales (Megaptera novaeangliae), from the Southwestern Atlantic Ocean, pp.407-440, 2018.

V. A. Ryabov, The effect of acoustic shielding the region of a dolphin's mental foramens on its hearing sensitivity, St. Petersbg. State Polytech. Univ. Journal. Phys. Math, vol.248, pp.102-110, 2016.

K. M. Stafford, C. Lydersen, Ø. Wiig, and K. Kovacs, Extreme diversity in the songs of Spitsbergen's bowhead whales, Biol. Lett, vol.14, 2018.

E. V. Sysueva, P. E. Nachtigall, T. W. Cranford, A. F. Pacini, J. L. Pawloski et al., Auditory sensitivity areas of head to local underwater stimulation in a bottlenose dolphin (Tursiops truncatus), Proc. Meet. Acoust, vol.30, p.10003, 2018.

J. G. Thewissen and S. T. Hussain, Origin of underwater hearing in whales, Nature, vol.361, pp.444-445, 1993.

G. Thiery, V. Lazzari, A. Ramdarshan, and F. Guy, Beyond the Map: Enamel Distribution Characterized from 3D Dental Topography, Front. Physiol, vol.8, pp.1-12, 2017.

S. W. Tinker, Whales of the World, 1988.

S. J. Trumble, S. A. Norman, D. D. Crain, F. Mansouri, Z. C. Winfield et al.,

D. C. Adams and E. Otárola-castillo, geomorph: an R package for the collection and analysis of geometric morphometric shape data, Methods Ecol. Evol, vol.4, pp.393-399, 2013.

D. C. Adams, F. J. Rohlf, and D. E. Slice, Geometric morphometrics: Ten years of progress following the 'revolution, Ital. J. Zool, vol.71, pp.5-16, 2004.

J. J. El-adli and T. A. Deméré, On the Anatomy of the Temporomandibular Joint and the Muscles That Act Upon It: Observations on the Gray Whale, Eschrichtius robustus, Anat. Rec, vol.298, pp.680-690, 2015.

J. A. Allen, E. C. Garland, R. A. Dunlop, and M. Noad, Cultural revolutions reduce complexity in the songs of humpback whales, Proc. R. Soc. B Biol. Sci, vol.285, 2018.

E. Amson, G. Billet, and C. De-muizon, Evolutionary adaptation to aquatic lifestyle in extinct sloths can lead to systemic alteration of bone structure, Proc. R. Soc. B Biol. Sci, vol.285, 2018.
URL : https://hal.archives-ouvertes.fr/hal-02611956

A. Barr and W. , Methods in Paleoecology: Reconstructing Cenozoic Terrestrial Environments and Ecological Communities, pp.339-349, 2018.

C. W. Andrews, A description of new species of zeuglodont and of leathery turtle from the Eocene of southern Nigeria, Proc. Zool. Soc. London, vol.1919, pp.309-319, 1920.

Ú. Árnason, A. Gullberg, S. Gretarsdottir, B. Ursing, and A. Janke, The Mitochondrial Genome of the Sperm Whale and a New Molecular Reference for Estimating Eutherian Divergence Dates, J. Mol. Evol, vol.50, pp.569-578, 2000.

Ú. Árnason, F. Lammers, V. Kumar, M. A. Nilsson, and A. Janke, Whole-genome sequencing of the blue whale and other rorquals finds signatures for introgressive gene flow, Sci. Adv, vol.4, p.9873, 2018.

W. J. Ary, T. W. Cranford, A. Berta, and P. Krysl, Functional Morphology and Symmetry in the Odontocete Ear Complex, pp.57-64, 2016.

R. J. Asher and K. M. Helgen, Nomenclature and placental mammal phylogeny, BMC Evol. Biol, vol.10, p.102, 2010.

A. Assemat and N. Brualla, The petrosal bone of Prodremotherium sp, 2015.

. Morphomuseum,

S. Bajpai and P. D. Gingerich, A new Eocene archaeocete (Mammalia, Cetacea) from India and the time of origin of whales, Proc. Natl. Acad. Sci, vol.95, pp.15464-15468, 1998.

S. Bajpai and J. G. Thewissen, Protocetid cetaceans (Mammalia) from the Eocene of India, 2014.

S. Bajpai, J. G. Thewissen, and A. Sahni, The origin and early evolution of whales: Macroevolution documented on the Indian Subcontinent, J. Biosci, vol.34, pp.673-686, 2009.

L. T. Ballance, Cetacean Ecology, Encyclopedia of Marine Mammals. Second Edition, pp.196-201, 2008.

W. E. Barklow, Some Underwater Sounds of the Hippopotamus (Hippopotamus amphibius), Mar. Freshw. Behav. Physiol, vol.29, pp.237-249, 1997.

W. E. Barklow, Amphibious communication with sound in hippos, Hippopotamus amphibius, Anim. Behav, vol.68, pp.1125-1132, 2004.

W. E. Barklow, Low-frequency sounds and amphibious communication in Hippopotamus amphibius, J. Acoust. Soc. Am, vol.115, pp.2555-2555, 2004.

R. Barone, Anatomie comparee des mammiferes domestiques. Tome Premier Ostéologie, Fascicule, vol.2, 1976.

W. A. Barr, Functional morphology of the bovid astragalus in relation to habitat: Controlling phylogenetic signal in ecomorphology, J. Morphol, vol.275, pp.1201-1216, 2014.

C. Barroso, T. W. Cranford, and A. Berta, Shape analysis of odontocete mandibles: Functional and evolutionary implications, J. Morphol, vol.273, pp.1021-1030, 2012.

R. M. Bebej, I. S. Zalmout, A. A. El-aziz, M. S. Antar, P. D. Gingerich et al., First remingtonocetid archaeocete (Mammalia, Cetacea) from the middle Eocene of Egypt with implications for biogeography and locomotion in early cetacean evolution, J. Paleontol, vol.89, p.191, 2015.

A. Berta, Return to the sea. The Life and Evolutionary Times of Marine Mammals, 2012.

A. Berta, J. L. Sumich, and K. M. Kovacs, Marine Mammals, 2015.

G. Bianucci and P. D. Gingerich, Aegyptocetus tarfa, n. gen. et sp. (Mammalia, Cetacea), from the middle Eocene of Egypt: Clynorhynchy, olfaction, and hearing in protocetid whale, J. Vertebr. Paleontol, vol.31, pp.1173-1188, 2011.

G. Bianucci, O. Lambert, R. Salas-gismondi, J. Tejada, F. Pujos et al., A Miocene relative of the Ganges River dolphin (Odontoceti, Platanistidae) from the Amazonian Basin, J. Vertebr. Paleontol, vol.33, pp.741-745, 2013.
URL : https://hal.archives-ouvertes.fr/hal-01922388

F. Bibi, A multi-calibrated mitochondrial phylogeny of extant Bovidae (Artiodactyla, Ruminantia) and the importance of the fossil record to systematics, BMC Evol. Biol, vol.13, p.166, 2013.

M. Bisconti, O. Lambert, and M. E. Bosselaers, Taxonomic revision of Isocetus depauwi (Mammalia, Cetacea, Mysticeti) and the phylogenetic relationships of archaic 'cetothere' mysticetes, Palaeontology, vol.56, pp.95-127, 2013.

L. C. Bishop, Cenozoic Mammals of Africa, pp.821-842, 2010.

C. C. Black, Anthracotheriidae, Evolution of African Mammals, pp.423-434, 1978.

S. P. Blomberg and T. Garland, Tempo and mode in evolution: phylogenetic inertia, adaptation and comparative methods, J. Evol. Biol, vol.15, pp.899-910, 2002.

J. F. Blumenbach, Handbuch der Naturgeschichte, 1779.

W. J. Bock, From Biologische Anatomie To Ecomorphology, Netherlands J. Zool, vol.40, pp.254-277, 1989.

W. J. Bock, Concepts and methods in ecomorphology, J. Biosci, vol.19, pp.403-413, 1994.

W. J. Bock and G. Von-wahlert, Adaptation and the form-function complex, Evolution (N. Y), vol.19, pp.269-299, 1965.

R. W. Boessenecker, A new marine vertebrate assemblage from the Late Neogene Purisima Formation in Central California, part II: Pinnipeds and Cetaceans, Geodiversitas, vol.35, pp.815-940, 2013.

R. W. Boessenecker and R. E. Fordyce, A new Eomysticetid (Mammalia: Cetacea) from the Late Oligocene of New Zealand and a re-evaluation of 'Mauicetus' waitakiensis, Pap. Palaeontol, vol.1, pp.107-140, 2015.

J. Boisserie, Systématique phylogénétique des Hippopotamidae fossiles d'Afrique : problèmes actuels. Ann. d'Ethiopie, vol.17, pp.185-204, 2001.

J. Boisserie, The Evolution of Artiodactyls, pp.106-119, 2007.

J. Boisserie, R. E. Fisher, F. Lihoreau, and E. M. Weston, Evolving between land and water: key questions on the emergence and history of the Hippopotamidae (Hippopotamoidea, Cetancodonta, Cetartiodactyla), Biol. Rev, vol.86, pp.601-625, 2011.
URL : https://hal.archives-ouvertes.fr/hal-00586344

J. Boisserie, F. Lihoreau, and M. Brunet, The position of Hippopotamidae within Cetartiodactyla, Proc. Natl. Acad. Sci, vol.102, pp.1537-1541, 2005.
URL : https://hal.archives-ouvertes.fr/hal-02886595

J. Boisserie, F. Lihoreau, M. J. Orliac, R. E. Fisher, E. M. Weston et al., Morphology and phylogenetic relationships of the earliest known hippopotamids (Cetartiodactyla, Hippopotamidae, Kenyapotaminae), Zool. J. Linn. Soc, vol.158, pp.325-366, 2010.

J. Boisserie, G. Suwa, B. Asfaw, F. Lihoreau, R. L. Bernor et al., Basal hippopotamines from the upper Miocene of Chorora, Ethiopia. J. Vertebr. Paleontol, vol.37, 2017.
URL : https://hal.archives-ouvertes.fr/hal-02880163

M. Boivin, M. J. Orliac, M. Telles-antunes, M. Godinot, Y. Laurent et al., New material of Diacodexis (Mammalia, Artiodactyla) from the early Eocene of Southern Europe, Geobios, vol.51, pp.285-306, 2018.

F. L. Bookstein, Morphometric tools for landmark data, 1992.

F. L. Bookstein, Landmark methods for forms without landmarks: localizing group differences in outline shape, Proc. Work. Math. Methods Biomed. Image Anal, vol.1, pp.225-243, 1996.

E. Bourdon and H. Cappetta, Pseudo-toothed birds (Aves, Odontopterygiformes) from the eocene phosphate deposits of Togo, Africa. J. Vertebr. Paleontol, vol.32, pp.965-970, 2012.

T. E. Bowdich, An analysis of the Natural Classifications of Mammalia, for the use of Students and Travellers. Paris: Smith, J. Boyden A. 1935. Genetics and Homology, Q. Rev. Biol, vol.10, pp.448-451

A. Boyden and D. Gemeroy, The relative position of the Cetacea among orders of Mammalia as indicated by precipitin tests, Zoologica, vol.35, pp.145-151, 1950.

K. Bremer, Branch Support and Tree Stability, Cladistics, vol.10, pp.295-304, 1994.

R. L. Brill, P. W. Moore, D. A. Helweg, and L. A. Dankiewicz, Investigating the Dolphin's Peripheral Hearing System: Acoustic Sensitivity About the Head and Lower Jaw, 2001.

R. L. Brill, M. L. Sevenich, T. J. Sullivan, J. D. Sustman, and R. E. Witt, Behavioral evidence for hearing through the lower jaw by an echolocating dolphin (Tursiops truncatus), Mar. Mammal Sci, vol.4, pp.223-230, 1988.

M. Brisson, Regnum animale in classes IX. distributum, sive, Synopsis methodica : sistens generalem animalium distributionem in classes IX, & duarum primarum classium, quadrupedum scilicet & cetaceorum, particularem divisionem in ordines, sectiones, genera &a, Lugduni Batavorum (Leyde): Apud Theodorum Haak, 1762.

T. H. Bullock, A. D. Grinnell, E. Ikezono, K. Kameda, Y. Katsuki et al., Electrophysiological studies of central auditory mechanisms in cetaceans, Z. Vgl. Physiol, vol.59, pp.117-156, 1968.

M. R. Buono, M. T. Dozo, F. G. Marx, and R. E. Fordyce, A Late Miocene potential neobalaenine mandible from Argentina sheds light on the origins of the living pygmy right whale, Acta Palaeontol. Pol, vol.59, pp.787-793, 2014.

C. J. Burgin, J. P. Colella, P. L. Kahn, and N. S. Upham, How many species of mammals are there?, J. Mammal, vol.99, pp.1-14, 2018.

J. A. Buytaert, W. H. Salih, M. Dierick, P. Jacobs, and J. J. Dirckx, Realistic 3D Computer Model of the Gerbil Middle Ear, Featuring Accurate Morphology of Bone and Soft Tissue Structures, J. Assoc. Res. Otolaryngol, vol.12, pp.681-696, 2011.

H. Cappetta and M. Traverse, Une riche faune de Sélaciens dans le bassin à phosphatede Kpogamé-Hahotoé (Éocène moyen du Togo): Note préliminaire et précisions sur la structure et l'âge du gisement, Geobios, vol.21, pp.359-365, 1988.

M. M. Churchill, M. Martínez-cáceres, C. Muizon, . De, J. Mnieckowski et al., The Origin of High-Frequency Hearing in Whales, Curr. Biol, vol.26, pp.2144-2149, 2016.
URL : https://hal.archives-ouvertes.fr/hal-02612270

R. Clarke, Hearing in Cetacea, Nature, vol.161, pp.979-980, 1948.

M. T. Clementz, A. Goswami, P. D. Gingerich, and P. L. Koch, Isotopic records from early whales and sea cows: Contrasting patterns of ecological transition, J. Vertebr. Paleontol, vol.26, pp.355-370, 2006.

M. T. Clementz, P. A. Holroyd, and P. L. Koch, Identifying Aquatic Habits Of Herbivorous Mammals Through Stable Isotope Analysis, Palaios, vol.23, pp.574-585, 2008.

K. M. Cohen, S. C. Finney, P. L. Gibbard, and J. Fan, The ICS International Chronostratigraphic Chart. Episodes, vol.36, pp.199-204, 2018.

G. C. Conroy and M. W. Vannier, Noninvasive Three-Dimensional Computer Imaging of Matrix-Filled Fossil Skulls by High-Resolution Computed Tomography. Science (80-. ), vol.226, pp.456-458, 1984.

L. N. Cooper, Evolution and development of cetacean appendages -A dissertation submitted to Kent State University in partial fulfillment of the requirements for the degree of Doctor of Philosophy, 2009.

L. N. Cooper, M. T. Clementz, S. Usip, S. Bajpai, S. T. Hussain et al., Aquatic habits of cetacean ancestors: Integrating bone microanatomy and stable isotopes, Integr. Comp. Biol, vol.56, pp.1370-1384, 2016.

L. N. Cooper, J. G. Thewissen, S. Bajpai, and B. N. Tiwari, Postcranial morphology and locomotion of the Eocene raoellid Indohyus (Artiodactyla: Mammalia), Hist. Biol, vol.24, pp.279-310, 2012.

E. D. Cope, The oldest artiodactyl, Am. Nat, vol.16, p.61, 1882.

E. D. Cope, The Phylogeny of the Camelidae, Am. Nat, vol.20, pp.611-624, 1886.

S. M. Cote, Cenozoic Mammals of Africa, pp.731-740, 2010.

T. W. Cranford and P. Krysl, Fin whale sound reception mechanisms: Skull vibration enables lowfrequency hearing, PLoS One, vol.10, pp.1-17, 2015.

T. W. Cranford and P. Krysl, Sound Paths, Cetaceans. Encyclopedia of Marine Mammals, pp.901-904, 2018.

T. W. Cranford, P. Krysl, and M. Amundin, A new acoustic portal into the odontocete ear and vibrational analysis of the tympanoperiotic complex, PLoS One, vol.5, pp.11927-193, 2010.

T. W. Cranford, P. Krysl, and J. A. Hildebrand, Acoustic pathways revealed: Simulated sound transmission and reception in Cuvier's beaked whale (Ziphius cavirostris), Bioinspiration and Biomimetics, vol.3, pp.1-10, 2008.

W. C. Cummings and P. O. Thompson, Underwater Sounds from the Blue Whale, Balaenoptera musculus, J. Acoust. Soc. Am, vol.50, pp.1193-1198, 1971.

S. C. Curran, Expanding ecomorphological methods: geometric morphometric analysis of Cervidae post-crania, J. Archaeol. Sci, vol.39, pp.1172-1182, 2012.

S. C. Curran, Methods in Paleoecology: Reconstructing Cenozoic Terrestrial Environments and Ecological Communities, pp.319-337, 2018.

F. Cuvier, Du Sanglier à masque et des Phacochoeres. Mémoires du Muséum d'Histoire Nat, vol.8, pp.447-455

G. Cuvier, Recherches sur les Ossemens Fossiles, où l'on rétablit les caractères de plusieurs animaux dont les révolutions du globe ont détruit les espèces. Nouvelle Edition. Tome Troisième

G. Cuvier, Recherches sur les Ossemens Fossiles, où l'on rétablit les caractères de plusieurs animaux dont les révolutions du globe ont détruit les espèces. Quatrième Edition. Tome Cinquième, 1835.

P. Darlu and P. Tassy, La Reconstruction phylogénétique, 1993.

C. Darwin and A. R. Wallace, On the Tendency of Species to form Varieties; and on the Perpetuation of Varieties and Species by Natural Means of Selection, pp.45-62, 1858.

R. David, A. Stoessel, A. Berthoz, F. Spoor, and D. Bennequin, Assessing morphology and function of the semicircular duct system: introducing new in-situ visualization and software toolbox, Sci. Rep, vol.6, p.32772, 2016.
URL : https://hal.archives-ouvertes.fr/hal-01375786

B. V. Dickson, E. Sherratt, J. B. Losos, and S. E. Pierce, Semicircular canals in Anolis lizards: ecomorphological convergence and ecomorph affinities of fossil species, R. Soc. Open Sci, vol.4, p.170058, 2017.

D. P. Domning and V. De-buffrénil, Hydrostasis in the Sirenia: Quantitative data and functional interpretations, Mar. Mammal Sci, vol.7, pp.331-368, 1991.

A. H. Doran, 1878. XVIII. Morphology of the Mammalian Ossicula auditûs, Trans. Linn. Soc. London. 2nd Ser. Zool, vol.1, pp.371-497

S. Ducrocq, A. N. Soe, A. K. Aung, M. Benammi, B. Bo et al., A new anthracotheriid artiodactyl from Myanmar, and the relative ages of the eocene anthropoid primate-bearing localities of Thailand (Krabi) and Myanmar (Pondaung), J. Vertebr. Paleontol, vol.20, pp.755-760, 2001.

W. F. Eddy, A New Convex Hull Algorithm for Planar Sets, ACM Trans. Math. Softw, vol.3, pp.398-403, 1977.

G. Ehret and M. C. Göpfert, Neurosciences -From Molecule to Behavior: a university textbook, pp.337-362, 2013.

E. G. Ekdale, Comparative Anatomy of the Bony Labyrinth (Inner Ear) of Placental Mammals, PLoS One, vol.8, p.100, 2013.

E. G. Ekdale, Morphological variation among the inner ears of extinct and extant baleen whales (Cetacea: Mysticeti), J. Morphol, vol.277, pp.1599-1615, 2016.

E. G. Ekdale, The Ear of Mammals: From Monotremes to Humans, Evolution of the Vertebrate Ear, pp.175-206, 2016.

E. G. Ekdale, Form and function of the mammalian inner ear, J. Anat, vol.228, pp.324-337, 2016.

E. G. Ekdale, A. Berta, and T. A. Deméré, The Comparative Osteology of the Petrotympanic Complex (Ear Region) of Extant Baleen Whales (Cetacea: Mysticeti), PLoS One, vol.6, p.21311, 2011.

E. G. Ekdale and R. A. Racicot, Anatomical evidence for low frequency sensitivity in an archaeocete whale: Comparison of the inner ear of Zygorhiza kochii with that of crown Mysticeti, J. Anat, vol.226, pp.22-39, 2015.

E. S. Jansson, A. Meloro, C. Louys, J. Plummer, T. Bishop et al., Exploring morphological generality in the Old World monkey postcranium using an ecomorphological framework, J. Anat, vol.228, pp.534-560, 2016.

C. Erbe, Hearing Abilities of Baleen Whales, 2002.

J. Erfurt, . Métais-g-;-cebochoeridae, . Choeropotamidae, . Mixtotheriidae, . Cainotheriidae et al., Endemic European Paleogene Artiodactyls, pp.59-84, 2007.

N. Eriksen, L. A. Miller, J. Tougaard, and D. A. Helweg, Cultural change in the songs of humpback whales (Megaptera novaeangliae) from Tonga, Behaviour, vol.142, pp.305-328, 2005.

J. S. Farris, Methods for Computing Wagner Trees, Syst. Biol, vol.19, pp.83-92, 1970.

J. S. Farris, The Retention Index and the Rescaled Consistency Index, Cladistics, vol.5, pp.417-419, 1989.

D. L. Felten, M. K. O&apos;banion, and M. S. Maida, Sensory Systems. Netter's Atlas of Neuroscience, pp.353-389, 2016.

S. Ferreira-cardoso, R. Araújo, N. E. Martins, G. G. Martins, S. Walsh et al., Floccular fossa size is not a reliable proxy of ecology and behaviour in vertebrates, Sci. Rep, vol.7, 2005.

E. M. Fitzgerald, The morphology and systematics of Mammalodon colliveri (Cetacea: Mysticeti), a toothed mysticete from the Oligocene of Australia, Zool. J. Linn. Soc, vol.158, pp.367-476, 2010.

G. Fleischer, Structural Analysis of the Tympanicum Complex in the Bottlenosed Dolphin (Tursiops truncatus), J. Aud. Res, vol.13, pp.178-190, 1973.

G. Fleischer, Hearing in Extinct Cetaceans as Determined by Cochlear Structure, J. Paleontol, vol.50, pp.133-152, 1976.

G. Fleischer, Evolutionary Principles of the Mammalian Middle Ear, Adv. Anatomy, Embryol. Cell Biol, vol.55, pp.1-70, 1978.

W. H. Flower, Description of the skeleton of Inia geoffrensis and the skull of Pontoporia blainvillii, with remarks on the systematic position of these animals in the Order Cetacea, Trans. Zool. Soc. London, vol.6, pp.87-116, 1867.

W. H. Flower, On whales, past and present and their probable origin. A discourse, Sci. Am. Suppl, vol.399, pp.6372-6374, 1883.

W. H. Flower and H. Gadow, An Introduction to the Osteology of the Mammalia, 1885.

R. E. Fordyce, Cetacean Evolution and Eocene/Oligocene Environments, Eocene-Oligocene Climatic and Biotic Evolution, pp.368-381, 1992.

R. E. Fordyce, Waipatia maerewhenua, new genus and new species (Waipatiidae, new family), an archaic Late Oligocene dolphin (Cetacea: Odontoceti: Platanistoidea) from New Zealand. Proc. San Diego Soc, vol.29, pp.147-176, 1994.

R. E. Fordyce, Simocetus rayi (Odontoceti: Simocetidae) (new species, new genus, new family), a bizarre new archaic Oligocene dolphin from the eastern North Pacific, pp.185-222, 2002.

R. E. Fordyce, The transition from Archaeoceti to Neoceti: Oligocene archaeocetes in the Southwest Pacific, J. Vertebr. Paleontol, vol.24, pp.1-139, 2004.

R. E. Fordyce, C. Muizon, and . De, Evolutionary history of cetaceans: a review, Secondary Adaptation of Tetrapods to lLfe in Water. Proceedings of the international meeting Poitiers, pp.169-233, 1996.

S. E. Foss and D. R. Prothero, The Evolution of Artiodactyls, pp.1-3, 2007.

E. Fraas, Neue Zeuglodonten aus dem unteren Mitteleocän vom Mokattam bei Cairo, Geol. und Palaontologische Abhandlungen, vol.6, pp.197-220, 1904.

F. C. Fraser and H. W. Parker, Handbook of R.H. Burne's Cetacean dissections, 1952.

F. C. Fraser and P. E. Purves, Hearing in Cetaceans. Bull. Br. Museum (Natural Hist. Zool, vol.7, pp.1-140, 1960.

F. C. Fraser and P. E. Purves, Anatomy and Function of the Cetacean Ear, Proc. R. Soc. London. Ser. B, Biol. Sci, vol.152, pp.62-77, 1960.

T. Frisch, M. V. Sørensen, S. Overgaard, M. Lind, and P. Bretlau, Volume-Referent Bone Turnover Estimated From the Interlabel Area Fraction After Sequential Labeling, Bone, vol.22, pp.677-682, 1998.

A. Galik, E. Mohandesan, G. Forstenpointner, U. M. Scholz, E. Ruiz et al., A Sunken Ship of the Desert at the River Danube in Tulln, Austria. PLoS One, vol.10, p.121235, 2015.

M. Garcia, M. Wondrak, L. Huber, and W. T. Fitch, Honest signaling in domestic piglets (Sus scrofa domesticus): vocal allometry and the information content of grunt calls, J. Exp. Biol, vol.219, pp.1913-1921, 2016.

J. Gatesy, J. H. Geisler, J. Chang, C. Buell, A. Berta et al., A phylogenetic blueprint for a modern whale, Mol. Phylogenet. Evol, vol.66, pp.479-506, 2013.

J. Gatesy, C. Hayashi, M. A. Cronin, and P. Arctander, Evidence from milk casein genes that cetaceans are close relatives of hippopotamid artiodactyls, Mol. Biol. Evol, vol.13, pp.954-963, 1996.

J. Gatesy, M. C. Milinkovitch, V. Waddell, and M. Stanhope, Stability of Cladistic Relationships between Cetacea and Higher-Level Artiodactyl Taxa, Syst. Biol, vol.48, pp.6-20, 1999.

J. Gatesy and M. A. O&apos;leary, Deciphering whale origins with molecules and fossils, Trends Ecol. Evol, vol.16, pp.562-570, 2001.

E. W. Gaupp, Die Reichertsche theorie. (Hammer-, amboss-und kieferfrage), Arch. für Anat. und entwickelungsgeschichte, pp.1-416, 1913.

C. L. Gazin, A review of the upper Eocene Artiodactyla of North America, Smithson. Misc. Collect, vol.128, pp.1-96, 1955.

J. H. Geisler, New morphological evidence for the phylogeny of Artiodactyla Cetacea and Mesonychidae, Am. Museum Novit, vol.40, p.53, 2001.

J. H. Geisler and Z. Luo, Relationships of Cetacea to Terrestrial Ungulates and the Evolution of Cranial Vasculature in Cete, pp.163-212, 1998.

J. H. Geisler, A. E. Sanders, and Z. Luo, A New Protocetid Whale (Cetacea: Archaeoceti) from the Late Middle Eocene of South Carolina, Am. Museum Novit, vol.3480, pp.1-65, 2005.

J. H. Geisler and J. M. Theodor, Hippopotamus and whale phylogeny, Nature, vol.458, pp.1-1, 2009.

J. H. Geisler, J. M. Theodor, M. D. Uhen, and S. E. Foss, Phylogenetic Relationships of Cetaceans to Terrestrial Artiodactyls, pp.19-31, 2007.

J. H. Geisler and M. D. Uhen, Morphological support for a close relationship between hippos and whales, J. Vertebr. Paleontol, vol.23, pp.991-996, 2003.

J. H. Geisler and M. D. Uhen, Phylogenetic relationships of extinct cetartiodactyls: Results of simultaneous analyses of molecular, morphological, and stratigraphic data, J. Mamm. Evol, vol.12, pp.145-160, 2005.

A. W. Gentry and . Bovidae, Cenozoic Mammals of Africa, pp.741-796

A. W. Gentry and . Cervidae, Cenozoic Mammals of Africa, pp.813-814

A. W. Gentry and J. J. Hooker, The phylogeny and classification of the tetrapods, vol.2, pp.235-272, 1988.

D. Geraads, Cenozoic Mammals of Africa, pp.723-730, 2010.

S. Gero, A. Bøttcher, H. Whitehead, and P. T. Madsen, Socially segregated, sympatric sperm whale clans in the, Atlantic Ocean. R. Soc. Open Sci, vol.3, p.160061, 2016.

J. J. Gibson, The Senses Considered as Perceptual Systems, 1966.

M. L. Gibson, J. Mnieckowski, and J. H. Geisler, Tupelocetus palmeri, a new species of protocetid whale (Mammalia, Cetacea) from the middle Eocene of South Carolina, J. Vertebr. Paleontol, vol.38, p.1555165, 2019.

P. D. Gingerich, Land-to-sea transition in early whales: evolution of Eocene Archaeoceti (Cetacea) in relation to skeletal proportions and locomotion of living semiaquatic mammals, 2003.

P. D. Gingerich, Cenozoic Mammals of Africa, pp.873-900, 2010.

P. D. Gingerich, Evolution of Whales from Land to Sea, Proceedings of the American Philosophical SOciety, p.196, 2012.

P. D. Gingerich, M. S. Antar, and I. S. Zalmout, Skeleton of a new Protocetid (Cetacea, Archaeoceti) from the lower Gehannam formation of Wadi Al Hitan in Egypt: Survival of a Protocetid into the Priabonian Late Eocene, Soc. Vertebr. Paleontol, p.138, 2014.

P. D. Gingerich, M. Arif, and W. C. Clyde, New archaeocetes (Mammalia, Cetacea) from the Middle Eocene Domanda Formation of the Sulaiman Range, 1995.

P. D. Gingerich and H. Cappetta, A new archaeocete and other marine mammals (Cetacea and Sirenia) from lower middle Eocene phosphate deposits of Togo, J. Paleontol, vol.88, pp.109-129, 2014.

P. D. Gingerich, H. Cappetta, and M. Traverse, Marine mammals (Cetacea and Sirenia) from the middle Eocene of Kpogamé-Hahotoé in Togo, J. Vertebr. Paleontol, vol.12, p.29, 1992.

P. D. Gingerich, S. M. Raza, M. Arif, M. Anwar, and X. Zhou, New whale from the Eocene of Pakistan and the origin of cetacean swimming, Nature, vol.368, pp.844-847, 1994.

P. D. Gingerich and D. E. Russell, Pakicetus inachus, a new archaeocete (Mammalia, Cetacea) from the early-middle Eocene Kuldana formation of Kohat (Pakistan), Contrib. from Museum Contrib. from Museum Paleontol. Univ. Michigan, vol.25, pp.235-246, 1981.

P. D. Gingerich and D. E. Russell, Dentition of early Eocene Pakicetus (Mammalia, Cetacea), Contrib. from Museum Paleontol. Univ. Michigan, vol.28, pp.1-20, 1990.

P. D. Gingerich, M. Ul-haq, W. Von-koenigswald, W. J. Sanders, B. H. Smith et al., New protocetid whale from the middle eocene of pakistan: birth on land, precocial development, and sexual dimorphism, PLoS One, vol.4, p.4366, 2009.

P. D. Gingerich, M. Ul-haq, I. S. Zalmout, I. H. Khan, and M. S. Malkani, Origin of Whales from Early Artiodactyls: Hands and Feet of Eocene Protocetidae from Pakistan. Science (80-. ), vol.293, pp.2239-2242, 2001.

P. D. Gingerich and S. Zouhri, New fauna of archaeocete whales (Mammalia, Cetacea) from the Bartonian middle Eocene of southern Morocco, J. African Earth Sci, 2015.

A. Goëffon, J. Richer, and J. Hao, Heuristic Methods for Phylogenetic Reconstruction with Maximum Parsimony, Algorithms in Computational Molecular Biology, pp.579-597, 2010.

A. S. Gohar, M. S. Antar, S. El-sayed, and H. Sallam, Egyptian Eocene archaeocetes Mammalia Cetacea Review and new discoveries. 56th, Annu. Sci. Meet. Geol. Soc. Egypt, pp.31-32, 2018.

P. E. Gol&apos;din, New Paratethyan dwarf baleen whales mark the origin of cetotheres. PeerJ. 6:e5800. Gol'din P.E., Zvonok E. 2013. Basilotritus uheni, a New Cetacean (Cetacea, Basilosauridae) from the Late Middle Eocene of Eastern Europe, J. Paleontol, vol.87, pp.254-268, 2018.

D. J. Golz, Eocene Artiodactyla of Southern California, Nat. Hist. Museum Los Angeles Cty. -Sci. Bull, vol.26, pp.1-85, 1976.

G. Rodrigues, H. Lihoreau, F. Orliac, M. J. Thewissen, J. G. Boisserie et al., Unexpected evolutionary patterns of dental ontogenetic traits in cetartiodactyl mammals, Proc. R. Soc. B Biol. Sci, vol.286, 2019.
URL : https://hal.archives-ouvertes.fr/hal-02098192

S. J. Gould and E. S. Vrba, Exaptation-a Missing Term in the Science of Form, Paleobiology, vol.8, pp.4-15, 1982.

J. C. Gower, Generalized Procrustes Analysis, Psychometrika, vol.40, pp.33-51, 1975.

W. Graf and F. Klam, Le système vestibulaire : anatomie fonctionnelle et comparée, évolution et développement, Comptes Rendus -Palevol, 2006.

K. E. Graham, C. Hobaiter, J. Ounsley, T. Furuichi, and R. W. Byrne, Bonobo and chimpanzee gestures overlap extensively in meaning, PLOS Biol, vol.16, p.2004825, 2018.

D. Graur and D. G. Higgins, Molecular evidence for the inclusion of Cetaceans within the order Artiodactyla, Mol. Biol. Evol, vol.11, pp.357-364, 1994.

H. Gray, Anatomy of the Human Body, 1918.

J. E. Gray, On the natural arrangement of vertebrose animals, London Med. Repos, vol.15, pp.297-310, 1821.

J. E. Gray, The zoology of the voyage of the H.M.S. Erebus & Terror

J. E. Gray, On the Cetacea which have been observed in the seas surrounding the British Islands, Proc. Zool. Soc. London, pp.195-248, 1864.

W. D. Green, The thin-plate spline and images with curving features, Proc. Image Fusion Shape Var. Tech, pp.79-87, 1996.

W. K. Gregory, Mammalian Orders Part II. Genetic Relations of the Mammalian Orders, Bull. Am. Museum Nat. Hist, vol.27, pp.105-524, 1910.

C. Grohé, Z. J. Tseng, R. Lebrun, R. Boistel, and J. J. Flynn, Bony labyrinth shape variation in extant Carnivora: a case study of Musteloidea, J. Anat, vol.228, pp.366-383, 2016.

C. P. Groves and P. Grubb, Ungulate Taxonomy, 2011.

B. Grzimek, Grzimek's, Animal Life Encyclopedia, vol.15, 2003.

P. Gunz and P. Mitteroecker, Semilandmarks: A method for quantifying curves and surfaces, Hystrix, vol.24, pp.103-109, 2013.

P. Gunz, P. Mitteroecker, and F. L. Bookstein, Modern Morphometrics in Physical Anthropology, pp.73-98, 2005.

P. Gunz, M. Ramsier, M. Kuhrig, J. Hublin, and F. Spoor, The mammalian bony labyrinth reconsidered, introducing a comprehensive geometric morphometric approach, J. Anat, vol.220, pp.529-543, 2012.

K. Habel, R. Grasman, R. B. Gramacy, A. Stahel, and D. C. Sterratt, geometry: Mesh Generation and Surface Tesselation, 2015.

E. Haeckel, Generelle Morphologie der Organismen: allgemeine Grundzüge der organischen Formen-Wissenschaft, mechanisch begründet durch die von Charles Darwin reformirte Descendenz-Theorie II Allgemeine Entwicklungsgeschichte der Organismen, 1866.

J. M. Harris, D. Geraads, N. 2. Solounias, and . Camelidae, Cenozoic Mammals of Africa, pp.815-820

J. M. Harris and L. Li-ping, The Evolution of Artiodactyls, pp.130-150, 2007.

J. M. Harris, N. Solounias, D. 2. Geraads, and . Giraffoidea, Cenozoic Mammals of Africa, pp.797-812

D. Hartnagel, La perception de l'espace multisensoriel appréhendée par l'étude de la fusion visuoauditive, 2007.

A. Hassanin, F. Delsuc, A. Ropiquet, C. Hammer, B. Jansen-van-vuuren et al., Pattern and timing of diversification of Cetartiodactyla (Mammalia, Laurasiatheria), as revealed by a comprehensive analysis of mitochondrial genomes, C. R. Biol, vol.335, pp.32-50, 2012.

L. Hautier, R. Sarr, F. Lihoreau, R. Tabuce, M. Hameh et al., First record of the family Protocetidae in the Lutetian of Senegal (West Africa), Palaeovertebrata, vol.38, pp.1-7, 2014.
URL : https://hal.archives-ouvertes.fr/hal-01923037

J. Hector, Notes on New Zealand Cetacea, recent and fossil, Trans. Proc. New Zeal. Inst, vol.13, pp.434-437, 1881.

K. M. Helgen, Major mammalian clades: A review under consideration of molecular and palaeontological evidence, Mamm. Biol, vol.68, pp.1-15, 2003.

S. Hemilä, S. Nummela, and T. Reuter, What middle ear parameters tell about impedance matching and high frequency hearing, Hear. Res, vol.85, pp.31-44, 1995.

S. Hemilä, S. Nummela, and T. Reuter, A model of the odontocete middle ear, Hear. Res, vol.133, pp.82-97, 1999.

S. Hemilä, S. Nummela, and T. Reuter, Modeling whale audiograms: Effects of bone mass on highfrequency hearing, Hear. Res, vol.151, pp.221-226, 2001.

S. Hemilä, S. Nummela, and T. Reuter, Anatomy and physics of the exceptional sensitivity of dolphin hearing (Odontoceti: Cetacea), J. Comp. Physiol. A Neuroethol. Sensory, Neural, Behav. Physiol, vol.196, pp.165-179, 2010.

W. Hennig, Phylogenetic Systematics, Annu. Rev. Entomol, vol.10, pp.97-116, 1965.

A. E. Hernández-cisneros and C. Tsai, A possible enigmatic kekenodontid (Cetacea, Kekenodontidae) from the Oligocene of Mexico, Paleontol. Mex, vol.5, pp.147-155, 2016.

T. Hetherington, Comparative Anatomy and Function of Hearing in Aquatic Amphibians, Reptiles, and Birds, Sensory Evolution on the Threshold Adaptations in Secondarily Aquatic Vertebrates, pp.182-209, 2008.

P. A. Holroyd, F. Lihoreau, G. F. Gunnell, and E. R. Miller, Cenozoic Mammals of Africa, pp.843-852, 2010.

J. G. Honey, J. A. Harrison, D. R. Prothero, and M. S. Stevens, Evolution of Tertiary Mammals of North America, Terrestrial Carnivores, Ungulates, and Ungulate like Mammals, vol.1, pp.439-462, 1998.

J. J. Hooker and K. M. Thomas, A new species of Amphirhagatherium (Choeropotamidae, Artiodactyla, Mammalia) from the late Eocene Headeon Hill formation of Southern England and phylogeny of endemic European "Anthracotherioids, Palaeontology, vol.44, pp.827-853, 2001.

G. N. Hounsfield, Computerized transverse axial scanning (tomography): Part I. Description of system, Br. J. Radiol, vol.46, pp.1016-1022, 1973.

D. S. Houser, D. A. Helweg, and P. W. Moore, A Bandpass filter-bank model of auditory sensitivity in the humpback whale, Aquat. Mamm, vol.27, pp.82-91, 2001.

A. Houssaye, Pachyostosis" in aquatic amniotes: a review, Integr. Zool, vol.4, pp.325-340, 2009.
URL : https://hal.archives-ouvertes.fr/hal-00548784

G. B. Howes, Some Points in the Anatomy of the Porpoise (Phocoena communis), J. Anat. Physiol, vol.14, pp.467-74, 1880.

R. C. Hulbert, R. M. Petkewich, G. A. Bishop, D. Bukry, and D. P. Aleshire, A new middle Eocene protocetid whale (Mammalia: Cetacea: Archaeoceti) and associated biota from Georgia, J. Paleontol, 1998.

J. Hunter, XXXVIII. Observations on the structure and oeconomy of whales, Philos. Trans. R. Soc. London, vol.77, pp.371-450, 1787.

T. Ichihara, Formation Mechanism of Ear Plug in Baleen Whales in Relation To Glove-Finger, Sci. Reports Whales Res. Inst, vol.14, pp.107-135, 1959.

T. Ichihara, Prenatal Development of Ear Plug in Baleen Whales, Sci. Reports Whales Res, vol.18, pp.29-48, 1964.

J. K. Illiger, International Code of Zoological Nomenclature -Fourth Edition. London: The International Trust for Zoological Nomenclature, 1811.

D. M. Irwin, T. D. Kocher, and A. C. Wilson, Evolution of the Cytochrome b Gene of Mammals, J. Mol. Evol, vol.32, pp.128-144, 1991.

J. Rohlf, F. Slice, and D. , Extensions of the Procrustes Method for the Optimal Superimposition of Landmarks, Syst. Zool, vol.39, pp.40-5940, 1990.

C. M. Janis, K. M. Scott, and L. L. Jacobs, Evolution of Tertiary Mammals of North America, Terrestrial Carnivores, Ungulates, and Ungulate like Mammals, vol.1, 1998.

R. M. Joeckel and J. M. Stavas, Basicranial anatomy of Syndyoceras cooki (Artiodactyla, Protoceratidae) and the need for a reappraisal of tylopod relationships, J. Vertebr. Paleontol, vol.16, pp.320-327, 1996.

J. Josse and F. Husson, missMDA: A Package for Handling Missing Values in Multivariate Data Analysis, J. Stat. Softw, vol.70, pp.1-31, 2016.
URL : https://hal.archives-ouvertes.fr/hal-01314945

M. Kadwell, M. Fernandez, H. F. Stanley, R. Baldi, J. C. Wheeler et al., Genetic analysis reveals the wild ancestors of the llama and the alpaca, Proc. R. Soc. B Biol. Sci, vol.268, pp.2575-2584, 2001.

J. Kassewitz, M. T. Hyson, J. S. Reid, and R. L. Barrera, A Phenomenon Discovered While Imaging Dolphin Echolocation Sounds, J. Mar. Sci. Res. Dev, vol.6, 2016.

T. Kasuya, Systematic consideration of recent toothed whales based on the morphology of the tympano-periotic bone, Sci. Reports Whales Res. Inst, pp.1-103, 1973.

R. Kellogg, A Review of the Archaeoceti, Carnegie Inst. Washingt, vol.482, pp.1-366, 1936.

D. R. Ketten, The Marine Mammal Ear: Specializations for Aquatic Audition and Echolocation, The Evolutionary Biology of Hearing, pp.717-750, 1992.

D. R. Ketten, Functional Analyses of Whale Ears: Adaptations for Underwater Hearing, I.E.E.E Proc. Underw. Acoust, vol.1, pp.264-270, 1994.

D. R. Ketten, Structure and function in whale ears, Bioacoustics, vol.8, pp.103-135, 1997.

D. R. Ketten, Cetacean Ears, Hearing by whales and dolphins, pp.43-108, 2000.

D. R. Ketten, Marine mammal auditory systems: A summary of audiometric and anatomical data and implications for underwater acoustic impacts, Polarforschung, vol.72, pp.79-92, 2004.

D. R. Ketten and D. Wartzok, Three-dimensional reconstructions of the dolphin ear, Sensory Abilities of Cetaceans: Laboratory and Field Evidence, pp.81-105, 1990.

K. Zaw, S. Meffre, M. Takai, H. Suzuki, C. Burrett et al., The oldest anthropoid primates in SE Asia: Evidence from LA-ICP-MS U-Pb zircon age in the Late Middle Eocene Pondaung Formation, Myanmar. Gondwana Res, vol.26, pp.122-131, 2014.

S. L. Kim, J. G. Thewissen, M. M. Churchill, R. S. Suydam, D. R. Ketten et al., Unique Biochemical and Mineral Composition of Whale Ear Bones, Physiol. Biochem. Zool, vol.87, pp.576-584, 2014.

S. L. King and V. M. Janik, Bottlenose dolphins can use learned vocal labels to address each other, Proc. Natl. Acad. Sci, vol.110, pp.13216-13221, 2013.

M. D. Kinkel, J. G. Thewissen, and H. A. Oelschläger, Rotation of middle ear ossicles during cetacean development, J. Morphol, vol.249, pp.126-131, 2001.

C. J. Klaauw and . Van-der, The Auditory Bulla in Some Fossil Mammals. With a General Introduction to this Region of the Skull, Bull. Americ, vol.62, pp.1-352, 1931.

A. G. Kluge and J. S. Farris, Quantitative Phyletics and the Evolution of Anurans, Syst. Biol, vol.18, pp.1-32, 1969.

R. H. Kröger, Sensory Evolution on the Threshold Adaptations in Secondarily Aquatic Vertebrates, pp.113-120, 2008.

K. Kumar and A. Sahni, Eocene Mammals from the Upper Subathu Group, India. J. Vertebr. Paleontol, vol.5, pp.153-168, 1985.

K. Kumar and A. Sahni, Remingtonocetus harudiensis, new combination, a middle eocene archaeocete (mammalia, cetacea) from Western Kutch, India. J. Vertebr. Paleontol, 1986.

M. Ladegaard, J. Mulsow, D. S. Houser, F. H. Jensen, M. Johnson et al., Dolphin echolocation behaviour during active long-range target approaches, J. Exp. Biol, vol.222, p.189217, 2019.

S. Ladevèze, P. Missiaen, and T. Smith, Condylarthra") from the late Paleocene of Berru (France) and phylogenetic affinities of the enigmatic European family Pleuraspidotheriidae, J. Vertebr. Paleontol, vol.30, pp.1559-1578, 2010.

T. Lafarge, B. Pateiro-lópez, A. Possolo, and J. P. Dunkers, R Implementation of a Polyhedral Approximation to a 3D Set of Points Using the ? -Shape, J. Stat. Softw, vol.56, pp.1-19, 2014.

A. Lahunta, . De, E. Glass, and M. Kent, Veterinary Neuroanatomy and Clinical Neurology, 2008.

O. Lambert, G. Bianucci, and K. Post, A new beaked whale (Odontoceti, Ziphiidae) from the middle Miocene of Peru, J. Vertebr. Paleontol, vol.29, pp.910-922, 2009.

O. Lambert, G. Bianucci, R. Salas-gismondi, C. Di-celma, E. Steurbaut et al., An Amphibious Whale from the Middle Eocene of Peru Reveals Early South Pacific Dispersal of Quadrupedal Cetaceans, Curr. Biol, vol.29, pp.1352-1359, 2019.
URL : https://hal.archives-ouvertes.fr/hal-02611923

W. C. Lancaster, The Middle Ear of the Archaeoceti, J. Vertebr. Paleontol, vol.10, pp.117-127, 1990.

R. Lebrun, MorphoDig, an open-source 3d freeware dedicated to biology, Int. Paleontol. Congr, p.399, 2018.
URL : https://hal.archives-ouvertes.fr/hal-01876987

R. Lebrun, M. Godinot, S. Couette, P. Tafforeau, and C. Zollikofer, The labyrinthine morphology of Pronycticebus gaudryi (Primates, Adapiformes). Palaeobiodiversity and Palaeoenvironments, vol.92, pp.527-537, 2012.
URL : https://hal.archives-ouvertes.fr/hal-00765877

R. Lebrun, M. P. De-león, P. Tafforeau, and C. Zollikofer, Deep evolutionary roots of strepsirrhine primate labyrinthine morphology, J. Anat, vol.216, pp.368-380, 2010.
URL : https://hal.archives-ouvertes.fr/hal-01874240

S. Lees, D. B. Hanson, and E. A. Page, Some acoustical properties of the otic bones of a fin whale, J. Acoust. Soc. Am, vol.99, pp.2421-2427, 1996.

J. Leidy, The extinct mammalian fauna of Dakota and Nebraska : Including an account of some allied forms from other localities, together with a synopsis of the mammalian remains of North America. Philadelphia : Published for the Academy, 1869.

P. L. Li- and P. Helsinki, Chinese fossil Suoidea Systematics, Evolution, 2003.

D. R. Lide, CRC Handbook of Chemistry and Physics, 2005.

F. Lihoreau, Systématique et paléoécologie des Anthracotheriidae [Artiodactyla, 2003.

]. Suiformes and . Du, Mio-Pliocène de l'ancien monde: implications paléobiogéographiques

F. Lihoreau, L. Alloing-séguier, P. Antoine, J. Boisserie, L. Marivaux et al., Enamel microstructure defines a major Paleogene hippopotamoid clade: the Merycopotamini (Cetartiodactyla, Hippopotamoidea), Hist. Biol, vol.29, pp.947-957, 2017.
URL : https://hal.archives-ouvertes.fr/hal-01813199

F. Lihoreau, J. Barry, C. Blondel, and M. Brunet, A new species of Anthracotheriidae, Merycopotamus medioximus nov. sp. from the Late Miocene of the Potwar Plateau, Comptes Rendus Palevol, vol.3, pp.653-662, 2004.

F. Lihoreau, J. Barry, C. Blondel, Y. Chaimanee, J. Jaeger et al., Anatomical revision of the genus Merycopotamus (Artiodactyla; Anthracotheriidae): Its significance for Late Miocene Mammal Dispersal in Asia, Palaeontology, vol.50, pp.503-524, 2007.
URL : https://hal.archives-ouvertes.fr/hal-00243096

F. Lihoreau, J. Boisserie, C. Blondel, L. Jacques, A. Likius et al., Description and palaeobiology of a new species of Libycosaurus (Cetartiodactyla, Anthracotheriidae) from the Late Miocene of Toros-Menalla, northern Chad, J. Syst. Palaeontol, vol.12, pp.761-798, 2014.
URL : https://hal.archives-ouvertes.fr/hal-01893836

F. Lihoreau, J. Boisserie, F. K. Manthi, and S. Ducrocq, Hippos stem from the longest sequence of terrestrial cetartiodactyl evolution in Africa, Nat. Commun, vol.6, 2015.
URL : https://hal.archives-ouvertes.fr/hal-01893828

F. Lihoreau, J. Boisserie, L. Viriot, Y. Coppens, A. Likius et al., Anthracothere dental anatomy reveals a late Miocene Chado-Libyan bioprovince, Proc. Natl. Acad. Sci, vol.103, pp.8763-8767, 2006.
URL : https://hal.archives-ouvertes.fr/hal-00162938

F. Lihoreau and S. Ducrocq, The Evolution of Artiodactyls, pp.89-105, 2007.

F. Lihoreau, E. M. Essid, H. Khayati-ammar, L. Marivaux, W. Marzougui et al., The Libycosaurus (Hippopotamoidea, Artiodactyla) intercontinental dispersal event at the early late Miocene revealed by new fossil remains from Kasserine area, Tunisia. Hist. Biol, vol.00, pp.1-13, 2019.
URL : https://hal.archives-ouvertes.fr/hal-02082905

F. Lihoreau, L. Hautier, and M. Mahboubi, The new Algerian locality of Bir el Ater 3: validity of Libycosaurus algeriensis (Mammalia, Hippopotamoidea) and the age of the Nementcha Formation, pp.1-11, 2015.
URL : https://hal.archives-ouvertes.fr/hal-01893840

F. Lihoreau, F. Martin, G. Daver, S. Ducrocq, H. Gomes-rodrigues et al., A hippopotamoid adaptative suite: the case study of Bothriodon velaunum (Cuvier, 1824), Mammalia, 2018.
URL : https://hal.archives-ouvertes.fr/hal-02328477

. Int, . Paleontol, and . Congr, , p.66

D. G. Lillie, Observations on the Anatomy and General Biology of some Members of the Larger Cetacea, Proc. Zool. Soc. London, vol.80, pp.769-792, 1910.

D. G. Lillie, Terra Nova") Expedition. 1910. Br. Museum (Natural Hist, vol.3, pp.85-124, 1915.

C. V. Linnaeus, Systema naturae per regna tria naturae: secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Holmiae: Impensis Direct, 1758.

C. Lockyer, Age determination by means of the earplug in baleen whales, Rep. Int. Whal. Comm, vol.34, pp.692-696, 1984.

M. Logez and G. Terraz, Calculer la surface d'un polygone avec ADE4, 2015.

J. López-marulanda, O. Adam, T. Blanchard, M. Vallée, D. Cazau et al., First Results of an Underwater 360° HD Audio-Video Device for Etho-Acoustical Studies on Bottlenose Dolphins (Tursiops truncatus), Aquat. Mamm, vol.43, pp.162-176, 2017.

W. P. Luckett and N. Hong, Phylogenetic relationships between the orders artiodactyla and cetacea: A combined assessment of morphological and molecular evidence, J. Mamm. Evol, vol.5, pp.127-182, 1998.

Z. Luo, Homology and transformation of cetacean ectotympanic structures, pp.269-301, 1998.

Z. Luo, Transformation and diversification in early mammal evolution, Nature, vol.450, pp.1011-1019, 2007.

Z. Luo, Developmental Patterns in Mesozoic Evolution of Mammal Ears, Annu. Rev. Ecol. Evol. Syst, vol.42, pp.355-380, 2011.

Z. Luo, P. Chen, G. Li, and M. Chen, A new eutriconodont mammal and evolutionary development in early mammals, Nature, vol.446, pp.288-293, 2007.

Z. Luo and E. R. Eastman, Petrosal and Inner Ear of a Squalodontoid Whale : Implications for Evolution of Hearing in Odontocetes, J. Vertebr. Paleontol, vol.15, pp.431-442, 1995.

Z. Luo and P. D. Gingerich, Terrestrial Mesonychia to Aquatic Cetacea: Transformation of the Basicranium and Evolution of Hearing in Whales, Univ. Michigan -Pap. Paleontol, vol.31, pp.1-98, 1999.

Z. Luo and D. R. Ketten, CT scanning and computerized reconstructions of the inner ear of multituberculate mammals, J. Vertebr. Paleontol, vol.11, pp.220-228, 1991.

Z. Luo and K. Marsh, Petrosal (Periotic) and Inner Ear of a Pliocene Kogiine Whale, Implications on Relationships and Hearing Evolution of Toothed Whales, vol.16, pp.328-348, 1996.

Z. Luo, Q. Meng, D. M. Grossnickle, D. Liu, A. I. Neander et al., New evidence for mammaliaform ear evolution and feeding adaptation in a Jurassic ecosystem, Nature, vol.548, pp.326-329, 2017.

Z. Luo, I. Ruf, J. A. Schultz, and T. Martin, Fossil evidence on evolution of inner ear cochlea in Jurassic mammals, Proc. R. Soc. B Biol. Sci, vol.278, pp.28-34, 2011.

D. Lusseau, The emergence of cetaceans: Phylogenetic analysis of male social behaviour supports the Cetartiodactyla clade, J. Evol. Biol, vol.16, pp.531-535, 2003.

J. N. Lythgoe, The Ecology of Vision, 1979.

G. T. Macintyre, The Trisulcate Petrosal Pattern of Mammals, Evolutionary Biology, vol.6, pp.275-303, 1972.

N. Macleod, Generalizing and extending the eigenshape method of shape space visualization and analysis, Paleobiology, vol.25, pp.107-138, 1999.

W. P. Maddison, Squared-Change Parsimony Reconstructions of Ancestral States for Continuous-Valued Characters on a Phylogenetic Tree, Syst. Biol, vol.40, pp.304-314, 1991.

P. C. Mahalanobis, On the generalized distance in statistics, Proc. Natl. Inst. Sci. India, vol.2, pp.49-55, 1936.

W. Maier, The entotympanic in late fetal artiodactyla (Mammalia), J. Morphol, vol.274, pp.926-939, 2013.

S. Mansour, J. Magnan, H. Haidar, K. Nicolas, and S. Louryan, Comprehensive and Clinical Anatomy of the Middle Ear, 2013.

J. D. Marcot, Molecular Phylogeny of Terrestrial Artiodactyls: Conflicts and Resolution, pp.4-18, 2007.

M. J. Martin, S. H. Elwen, R. Kassanjee, and T. Gridley, To buzz or burst-pulse? The functional role of Heaviside's dolphin, Cephalorhynchus heavisidii, rapidly pulsed signals, Anim. Behav, vol.150, pp.273-284, 2019.

M. Martínez-cáceres, O. Lambert, C. Muizon, and . De, The anatomy and phylogenetic affinities of Cynthiacetus peruvianus, a large Dorudon-like basilosaurid (Cetacea, Mammalia) from the late Eocene of Peru, Geodiversitas, vol.39, pp.7-163, 2017.

J. Martinez and J. Sudre, The astragalus of Paleogene artiodactyls: comparative morphology, variability and prediction of body mass, Lethaia, vol.28, pp.197-209, 1995.

E. P. Martins, Estimation of Ancestral States of Continuous Characters: A Computer Simulation Study, Syst. Biol, vol.48, pp.642-650, 1999.

F. G. Marx, O. Lambert, and M. D. Uhen, Cetacean Paleobiology, 2016.

F. G. Marx and M. D. Uhen, Climate, Critters, and Cetaceans: Cenozoic Drivers of the Evolution of Modern Whales. Science (80-. ), vol.327, pp.993-996, 2010.

C. A. Matthee, J. D. Burzlaff, J. F. Taylor, and S. K. Davis, Mining the Mammalian Genome for Artiodactyl Systematics, Syst. Biol, vol.50, pp.367-390, 2001.

W. D. Matthew and W. Granger, New Mammals From the Shara Murun Eocene of Mongolia, Am. Museum Novit, vol.9, pp.415-424, 1925.

M. Maust-mohl, J. Soltis, and D. Reiss, Acoustic and behavioral repertoires of the hippopotamus (Hippopotamus amphibius ), J. Acoust. Soc. Am, vol.138, pp.545-554, 2015.

M. Maust-mohl, J. Soltis, and D. Reiss, Underwater click train production by the hippopotamus (Hippopotamus amphibius) suggests an echo-ranging function, Behaviour, vol.155, pp.231-251, 2018.

J. G. Mccormick, E. G. Wever, J. Palin, and S. H. Ridgway, Sound Conduction in the Dolphin Ear, J. Acoust. Soc. Am, vol.48, pp.1418-1428, 1970.

S. A. Mcleod and L. G. Barnes, A new genus and species of Eocene protocetid archaeocete whale (Mammalia, Cetacea) from the Atlantic Coastal Plain, Nat. Hist. Museum Los Angeles Cty. -Sci. Ser, vol.41, pp.73-98, 2008.

J. G. Mead and R. E. Fordyce, The therian skull : a lexicon with emphasis on the odontocetes, Smithson. Contrib. to Zool, pp.1-249, 2009.

R. W. Meredith, J. E. Janecka, J. Gatesy, O. A. Ryder, C. A. Fisher et al., Impacts of the Cretaceous Terrestrial Revolution and KPg Extinction on Mammal Diversification, vol.334, pp.521-524, 2011.

G. Métais and I. A. Vislobokova, The Evolution of Artiodactyls, pp.189-212, 2007.

M. C. Milinkovitch and J. G. Thewissen, Evolutionary biology. Even-toed fingerprints on whale ancestry, Nature, vol.388, pp.622-624, 1997.

E. D. Mitchell, A New Cetacean from the Late Eocene La Meseta Formation Seymour Island, Antarctic Peninsula, Can. J. Fish. Aquat. Sci, vol.46, pp.2219-2235, 1989.

G. I. Molina, Saggio sulla storia naturale del Chili, 1782.

A. R. Møller, Handbook of Sensory Physiology, pp.491-518, 1974.

A. R. Møller, Hearing: Anatomy, Physiology, and Disorders of the Auditory System, 2006.

C. Montgelard, F. M. Catzeflis, and E. Douzery, Phylogenetic relationships of Artiodactyls and Cetaceans as deduced from the comparison of cytochrome b and 12S rRNA mitochondrial sequences, Mol. Biol. Evol, vol.14, pp.550-559, 1997.

T. A. Mooney, M. Yamato, and B. K. Branstetter, Hearing in Cetaceans: From Natural History to Experimental Biology, Adv. Mar. Biol, vol.63, pp.197-246, 2012.

S. G. Morton, Additional observations on a new living species of Hippopotamus, of Western Africa (H. liberiensis), J. Acad. Nat. Sci. Philadelphia, vol.2, pp.231-239

P. J. Motta, K. B. Clifton, P. Hernandez, and B. T. Eggold, Ecomorphological correlates in ten species of subtropical seagrass fishes: diet and microhabitat utilization, Environ. Biol. Fishes, vol.44, pp.37-60, 1995.

P. J. Motta, K. B. Clifton, P. Hernandez, B. T. Eggold, S. D. Giordano et al., Feeding relationships among nine species of seagrass fishes of Tampa Bay, Florida. Bull. Mar. Sci, vol.56, pp.185-200, 1995.

P. J. Motta, S. F. Norton, and J. J. Luczkovich, Perspectives on the ecomorphology of bony fishes, Environ. Biol. Fishes, vol.44, pp.11-20, 1995.

M. J. Mourlam and M. J. Orliac, Infrasonic and Ultrasonic Hearing Evolved after the Emergence of Modern Whales, Curr. Biol, vol.27, pp.1776-1781, 2017.
URL : https://hal.archives-ouvertes.fr/hal-01894431

M. J. Mourlam and M. J. Orliac, Protocetid (Cetacea, Artiodactyla) bullae and petrosals from the middle Eocene locality of Kpogamé, Togo: new insights into the early history of cetacean hearing, J. Syst. Palaeontol, vol.16, pp.621-644, 2018.

C. D. Muizon, Nouvelles données sur le diphylétisme des Dauphins de rivière (Odontoceti, Cetacea, Mammalia), Academie des Sci. Série, vol.2, pp.359-362, 1985.

C. Muizon, . De, G. Billet, C. Argot, S. Ladevèze et al., Alcidedorbignya inopinata, a basal pantodont (Placentalia, Mammalia) from the early Palaeocene of Bolivia: Anatomy, phylogeny and palaeobiology, Geodiversitas, vol.37, pp.397-631, 2015.
URL : https://hal.archives-ouvertes.fr/hal-01455642

C. A. Müller, K. Schmitt, A. L. Barber, and L. Huber, Dogs Can Discriminate Emotional Expressions of Human Faces, Curr. Biol, vol.25, pp.601-605, 2015.

G. J. Naylor, D. C. Adams, and . O&apos;leary, Total Evidence Versus Relevant Evidence: A Response to, Syst. Biol, vol.52, pp.864-865, 2003.

J. M. Neenan, T. Reich, S. W. Evers, P. S. Druckenmiller, D. F. Voeten et al., Evolution of the Sauropterygian Labyrinth with Increasingly Pelagic Lifestyles, Curr. Biol, vol.27, pp.3852-3858, 2017.

S. V. Nelson, Isotopic reconstructions of habitat change surrounding the extinction of Sivapithecus, a Miocene hominoid, in the Siwalik Group of Pakistan, Palaeogeogr. Palaeoclimatol. Palaeoecol, vol.243, pp.204-222, 2007.

M. Nikaido, A. P. Rooney, and N. Okada, Phylogenetic relationships among cetartiodactyls based on insertions of short and long interpersed elements: Hippopotamuses are the closest extant relatives of whales, Proc. Natl. Acad. Sci. U. S. A, vol.96, pp.10261-10266, 1999.

K. S. Norris, Some problems of echolocation in cetaceans, Marine bioacoustics, pp.316-336, 1964.

K. S. Norris, The echolocation of marine mammals, pp.391-423, 1969.

K. S. Norris and G. W. Harvey, Sound transmission in the porpoise head, J. Acoust. Soc. Am, vol.56, pp.659-664, 1974.

K. S. Norris and W. N. Mcfarland, A New Harbor Porpoise of the Genus Phocoena from the Gulf of California, J. Mammal, vol.39, p.22, 1958.

S. F. Norton, J. J. Luczkovich, and P. J. Motta, The role of ecomorphological studies in the comparative biology of fishes, Environ. Biol. Fishes, vol.44, pp.287-304, 1995.

M. J. Novacek, Aspects of the problem of variation, origin and evolution of the eutherian auditory bulla, Mamm. Rev, vol.7, pp.131-150, 1977.

S. Nummela, Scaling of the mammalian middle ear, Hear. Res, vol.85, pp.18-30, 1995.

S. Nummela, Sensory Evolution on the Threshold Adaptations in Secondarily Aquatic Vertebrates, pp.210-224, 2008.

S. Nummela, The Primate Peripheral Auditory System and the Evolution of Primate Hearing, pp.13-45, 2017.

S. Nummela, S. T. Hussain, and J. G. Thewissen, Cranial anatomy of Pakicetidae (Cetacea, Mammalia), J. Vertebr. Paleontol, vol.26, pp.746-759, 2006.

S. Nummela, T. Reuter, S. Hemilä, P. Holmberg, and P. Paukku, The anatomy of the killer whale middle ear (Orcinus orca), Hear. Res, vol.133, pp.61-70, 1999.

S. Nummela and M. R. Sánchez-villagra, Scaling of the marsupial middle ear and its functional significance, J. Zool, vol.270, pp.256-267, 2006.

S. Nummela and J. G. Thewissen, The Physics of Sound in Air and Water, Sensory Evolution on the Threshold Adaptations in Secondarily Aquatic Vertebrates, pp.174-181, 2008.

S. Nummela, J. G. Thewissen, S. Bajpai, S. T. Hussain, and K. Kumar, Eocene evolution of whale hearing, Nature, vol.430, pp.776-778, 2004.

S. Nummela, J. G. Thewissen, S. Bajpai, S. T. Hussain, and K. Kumar, Sound transmission in archaic and modern whales: Anatomical adaptations for underwater hearing, Anat. Rec, vol.290, pp.716-733, 2007.

S. Nummela, T. Wägar, S. Hemilä, and T. Reuter, Scaling of the cetacean middle ear, Hear. Res, vol.133, pp.71-81, 1999.

S. Nummela and M. Yamato, Hearing. Encyclopedia of Marine Mammals, pp.462-470, 2018.

H. D. O&apos;brien, Cranial arterial pattern of the Sri Lankan spotted chevrotain, Moschiola memmina, and comparative basicranial osteology of the Tragulidae, PeerJ, vol.3, p.1451, 2015.

H. D. O&apos;brien, Cranial arterial patterns of the alpaca (Camelidae: Vicugna pacos), R. Soc. Open Sci, vol.4, p.160967, 2017.

H. O&apos;brien, From Anomalous Arteries to Selective Brain Cooling: Parallel Evolution of the Artiodactyl Carotid Rete, Anat. Rec, 2018.

M. A. O&apos;leary, An anatomical and phylogenetic study of the osteology of the petrosal of extant and extinct artiodactylans (Mammalia) and relatives, Bull. Am. Museum Nat. Hist, vol.335, pp.1-206, 2010.

M. A. O&apos;leary, Comparative Basicranial Anatomy of Extant Terrestrial and Semiaquatic Artiodactyla, Bull. Am. Museum Nat. Hist, vol.409, pp.1-55, 2016.

M. A. O&apos;leary, J. I. Bloch, J. J. Flynn, T. J. Gaudin, A. Giallombardo et al., The Placental Mammal Ancestor and the Post-K-Pg Radiation of Placentals, Science, vol.339, pp.662-667, 2013.

M. A. O&apos;leary and J. Gatesy, Impact of increased character sampling on the phylogeny of Cetartiodactyla (Mammalia): Combined analysis including fossils, Cladistics, vol.24, pp.397-442, 2008.

M. A. O&apos;leary, J. Gatesy, and M. J. Novacek, Are the Dental Data Really at Odds with the Molecular Data? Morphological Evidence for Whale Phylogeny (Re)Reexamined, Syst. Biol, vol.52, pp.853-864, 2003.

M. A. O&apos;leary and J. H. Geisler, The position of Cetacea within Mammalia: Phylogenetic analysis of morphological data from extinct and extant taxa, Syst. Biol, vol.48, pp.455-490, 1999.

M. A. O&apos;leary, B. A. Patel, and M. N. Coleman, Endocranial petrosal anatomy of Bothriogenys (Mammalia, Artiodactyla, Anthracotheriidae), and petrosal volume and density comparisons among aquatic and terrestrial artiodactyls and outgroups, J. Paleontol, vol.86, pp.44-50, 2012.

M. J. Orliac, The petrosal bone of extinct Suoidea (Mammalia, Artiodactyla), J. Syst. Palaeontol, vol.11, pp.925-945, 2013.

M. J. Orliac, P. Antoine, A. Charruault, S. Hervet, F. Prodeo et al., Specialization for amphibiosis in Brachyodus onoideus (Artiodactyla, Hippopotamoidea) from the Early Miocene of France, Swiss J. Geosci, vol.106, pp.265-278, 2013.
URL : https://hal.archives-ouvertes.fr/hal-01902587

M. J. Orliac, P. Antoine, and S. Ducrocq, Phylogenetic relationships of the Suidae (Mammalia, Cetartiodactyla): New insights on the relationships within Suoidea, vol.39, pp.315-330, 2010.

M. J. Orliac, R. Araújo, and F. Lihoreau, The petrosal and bony labyrinth of Diplobune minor, an enigmatic Artiodactyla from the Oligocene of Western Europe, J. Morphol, vol.278, pp.1168-1184, 2017.
URL : https://hal.archives-ouvertes.fr/hal-01942305

M. J. Orliac, C. Argot, and E. Gilissen, Condylarthra"): A Case of Paleogene Terrestrial Echolocation?, PLoS One, vol.7, p.30000, 2012.
URL : https://hal.archives-ouvertes.fr/hal-00721456

M. J. Orliac, J. Benoit, and M. A. O&apos;leary, The inner ear of Diacodexis, the oldest artiodactyl mammal, J. Anat, vol.221, pp.417-426, 2012.

M. J. Orliac and G. Billet, Fallen in a dead ear: intralabyrinthine preservation of stapes in fossil artiodactyls, Palaeovertebrata, vol.40, pp.1-10, 2016.
URL : https://hal.archives-ouvertes.fr/hal-01897786

M. J. Orliac, J. Boisserie, L. Maclatchy, and F. Lihoreau, Early Miocene hippopotamids (Cetartiodactyla) constrain the phylogenetic and spatiotemporal settings of hippopotamid origin, Proc. Natl. Acad. Sci. U. S. A, vol.107, pp.11871-11876, 2010.
URL : https://hal.archives-ouvertes.fr/hal-00591979

M. J. Orliac and S. Ducrocq, Eocene raoellids (Mammalia, Cetartiodactyla) outside the Indian Subcontinent: palaeogeographical implications, Geol. Mag, vol.149, pp.80-92, 2012.
URL : https://hal.archives-ouvertes.fr/hal-02328226

M. J. Orliac and E. Gilissen, Virtual endocranial cast of earliest Eocene Diacodexis (Artiodactyla, Mammalia) and morphological diversity of early artiodactyl brains, Proc. R. Soc. B Biol. Sci, vol.279, pp.3670-3677, 2012.

M. J. Orliac, F. Guy, and R. Lebrun, Osteological connections of the petrosal bone of the extant Hippopotamidae Hippopotamus amphibius and Choeropsis liberiensis, MorphoMuseuM, vol.1, p.1, 2014.
URL : https://hal.archives-ouvertes.fr/hal-01902601

M. J. Orliac and M. A. O&apos;leary, Comparative Anatomy of the Petrosal Bone of Dichobunoids, Early Members of Artiodactylamorpha (Mammalia), J. Mamm. Evol, vol.21, pp.299-320, 2014.
URL : https://hal.archives-ouvertes.fr/hal-01897864

M. J. Orliac and M. A. O&apos;leary, The inner ear of Protungulatum (Pan-Euungulata, Mammalia), J. Mamm. Evol, vol.23, 2016.
URL : https://hal.archives-ouvertes.fr/hal-01925070

R. Owen, Description of teeth and portions of jaw of two extinct anthraco-therioid quadrupeds (Hyopotamus vectianus and Hyop. bovinus) discovered by the Marchioness of Hastings in the Eocene deposits of the N.W. coast of the Isle of Wight: with an at, J. Geol. Soc. London, vol.4, pp.103-141, 1848.

R. D. Page, NDE (Nexus Data Editor for Windows), 2001.

O. C. Paine, J. N. Leichliter, N. Avenant, D. Codron, A. Lawrence et al., The ecomorphology of southern African rodent incisors: Potential applications to the hominin fossil record, PLoS One, vol.14, p.205476, 2019.

E. Paradis, Analysis of Phylogenetics and Evolution with R, 2006.

E. Paradis, J. Claude, and K. Strimmer, APE: Analyses of Phylogenetics and Evolution in R language, Bioinformatics, vol.20, pp.289-290, 2004.
URL : https://hal.archives-ouvertes.fr/ird-01887318

T. Park, E. M. Fitzgerald, and A. R. Evans, The Tympanal Recess of the Cetacean Cochlea: Function and Evolution, Acoust. Aust, vol.45, pp.273-278, 2017.

S. E. Parks, D. R. Ketten, J. T. O&apos;malley, and J. Arruda, Anatomical predictions of hearing in the North Atlantic right whale, Anat. Rec, vol.290, pp.734-744, 2007.

R. S. Payne and S. Mcvay, Songs of Humpback Whales. Science (80-. ), vol.173, pp.585-597, 1971.

R. S. Payne and D. Webb, Orientation By Means of Long Range Acoustic Signaling in Baleen Whales, Ann. N. Y. Acad. Sci, vol.188, pp.110-141, 1971.

H. S. Pearson, On the skulls of early tertiary Suidae together with an account of the otic region in some primitive Artiodactyla, Philos. Trans. R. Soc. London. Ser. B, vol.215, pp.389-460, 1927.

C. M. Peredo, N. D. Pyenson, M. D. Uhen, and C. D. Marshall, Alveoli, teeth, and tooth loss: Understanding the homology of internal mandibular structures in mysticete cetaceans, PLoS One, vol.12, p.178243, 2017.

S. I. Perez, V. Bernal, and P. N. Gonzalez, Differences between sliding semi-landmark methods in geometric morphometrics, with an application to human craniofacial and dental variation, J. Anat, vol.208, pp.769-784, 2006.

A. Perier, R. Lebrun, and L. Marivaux, Different Level of Intraspecific Variation of the Bony Labyrinth Morphology in Slow-Versus Fast-Moving Primates, J. Mamm. Evol, vol.23, pp.353-368, 2016.
URL : https://hal.archives-ouvertes.fr/hal-01813241

M. Pickford, On the origins of Hippopotamidae togetherwith descriptions of two new species, a new genus and a new subfamily from the Miocene of Kenya, Geobios, vol.16, pp.193-217, 1983.

M. Pickford, Revision of the Neogene Anthracotheriidae of Africa, The Geology of Libya, pp.1491-1525, 1991.

H. Pihlström, Sensory Evolution on the Threshold Adaptations in Secondarily Aquatic Vertebrates, pp.95-110, 2008.

G. E. Pilgrim, Middle Eocene Mammals from North-West India, Proc. Zool. Soc. London, vol.110, pp.127-152, 1940.

R. Pinhasi, D. Fernandes, K. Sirak, M. Novak, S. Connell et al., Optimal Ancient DNA Yields from the Inner Ear Part of the Human Petrous Bone, vol.10, p.129102, 2015.

P. D. , Adaptive Zones and the Pinniped Ankle: A Three-Dimensional Quantitative Analysis of Carnivoran Tarsal Evolution, pp.167-196, 2009.

V. V. Popov and A. Y. Supin, Analysis of auditory information in the brains of cetaceans, Neurosci. Behav. Physiol, vol.37, pp.285-291, 2007.

V. V. Popov, A. Y. Supin, V. O. Klishin, M. B. Tarakanov, and M. G. Pletenko, Evidence for double acoustic windows in the dolphin, Tursiops truncatus, J. Acoust. Soc. Am, vol.123, pp.552-560, 2008.

V. V. Popov, A. Y. Supin, D. I. Nechaev, A. A. Lemazina, and E. V. Sysueva, Position of an acoustic window in a beluga whale: Computation based on auditory evoked potential latencies, J. Acoust. Soc. Am, vol.145, pp.3578-3585, 2019.

V. V. Popov, E. V. Sysueva, D. I. Nechaev, A. A. Lemazina, and A. Y. Supin, Auditory sensitivity to local stimulation of the head surface in a beluga whale (Delphinapterus leucas), J. Acoust. Soc. Am, vol.140, pp.1218-1226, 2016.

S. A. Price, O. R. Bininda-emonds, and J. L. Gittleman, A complete phylogeny of the whales, dolphins and even-toed hoofed mammals (Cetartiodactyla), Biol. Rev. Camb. Philos. Soc, vol.80, pp.445-473, 2005.

U. Pritchard, 1881. II. The Cochlea of the Ornithorhynchus platypus compared with that of ordinary Mammals and of Birds, Philos. Trans. R. Soc. London, vol.172, pp.267-282

D. R. Prothero and S. E. Foss, The Evolution of Artiodactyls, 2007.

R. Team, R: A language and environment for statistical computing, 2015.

R. A. Racicot, S. A. Darroch, and N. Kohno, Neuroanatomy and inner ear labyrinths of the narwhal, Monodon monoceros, and beluga, Delphinapterus leucas (Cetacea: Monodontidae), J. Anat, vol.233, pp.421-439, 2018.

R. A. Racicot, W. Gearty, N. Kohno, and J. J. Flynn, Comparative anatomy of the bony labyrinth of extant and extinct porpoises (Cetacea: Phocoenidae), Biol. J. Linn. Soc, vol.119, pp.831-846, 2016.

T. S. Raffles, Descriptive Catalogue of a Zoological Collection, made on account of the Honourable East India Company, in the Island of Sumatra and its Vicinity, under the Direction of Sir Thomas Stamford Raffles, Lieutenant-Governor of Fort Marlborough, Trans. Linn. Soc. London, vol.13, pp.239-274

M. Rakotovao, Y. Lignereux, M. J. Orliac, F. Duranthon, and P. Antoine, Hippopotamus lemerlei, 2014.
URL : https://hal.archives-ouvertes.fr/hal-01904986

. Grandidier, 1868 et Hippopotamus madagascariensis Guldberg, 1883 (Mammalia, Hippopotamidae) : anatomie crânio-dentaire et révision systématique, Geodiversitas, vol.36, pp.117-161

R. Rao and A. , New Mammals from Murree (Kalakot Zone) of the Himalayan Foot Hills Near Kalakot, J. Geol. Soc. India, vol.12, pp.125-134, 1971.

M. A. Rauschmann, S. Huggenberger, L. S. Kossatz, and H. A. Oelschläger, Head morphology in perinatal dolphins: A window into phylogeny and ontogeny, J. Morphol, vol.267, pp.1295-1315, 2006.

S. J. Rehorek, R. Stimmelmayr, J. C. George, R. Suydam, D. M. Mcburney et al., Structure of the external auditory meatus of the Bowhead whale (Balaena mysticetus) and its relation to their seasonal migration, J. Anat, vol.234, pp.201-215, 2019.

K. B. Reichert, Ueber die Visceralbogen der Wirbelthiere im Allgemeinen und deren Metamorphosen bei den Vögeln und Säugethieren, Arch. für Anat. Physiol. und Wissenschaftliche Med, pp.120-222, 1837.

J. S. Reidenberg, Terrestrial, semiaquatic, and fully aquatic mammal sound production mechanisms, Acoutics Today, vol.13, pp.35-43, 2017.

J. S. Reidenberg and J. T. Laitman, Cetacean Prenatal Development, Encyclopedia of Marine Mammals. Second Edition, pp.220-238, 2008.

L. E. Rendell and H. Whitehead, Vocal clans in sperm whales (<i>Physeter macrocephalus<i/>), Proc. R. Soc. London. Ser. B Biol. Sci, vol.270, pp.225-231, 2003.

L. J. Revell, phytools: an R package for phylogenetic comparative biology (and other things), Methods Ecol. Evol, vol.3, pp.217-223, 2012.

F. W. Reysenbach-de-haan, Reysenbach de Haan F.W. 1960. Some aspects of mammalian hearing under water, Proc. R. Soc. Lond. B. Biol. Sci, vol.134, pp.54-62, 1957.

W. G. Ridewood, Observations on the Skull in Foetal Specimens of Whales of the Genera Megaptera and Balaenoptera, Philos. Trans. R. Soc. London. Ser. B, vol.211, pp.209-272, 1922.

P. T. Robinson, The status of the pygmy hippopotamus and other wildlife in West Africa, 1970.

P. T. Robinson, G. L. Flacke, and K. M. Hentschel, The Pygmy Hippo Story: West Africa's Enigma of the Rainforest, 2017.

L. J. Roe, J. G. Thewissen, J. Quade, J. R. O&apos;neil, S. Bajpai et al., Isotopic Approaches to Understanding the Terrestrial-to-Marine Transition of the Earliest Cetaceans, pp.399-422, 1998.

L. Rojas-bracho and B. L. Taylor, Vaquita. IUCN Red List Threat. Species. e.T17028A5, p.12, 2017.

G. Rondelet, Libri de piscibus marinis, quibus verae piscium effigies expressae sunt. Lugduni,: apud Matthiam Bonhomme, 1554.

K. D. Rose, Skeleton of Diacodexis, oldest known artiodactyl. Science (80-. ), vol.216, pp.621-623, 1982.

K. D. Rose, On the origin of the order Artiodactyla, Proc. Natl. Acad. Sci. U. S. A, vol.93, pp.1705-1709, 1996.

K. D. Rose, The Beginning of the Age of Mammals, pp.271-305, 2006.

J. J. Rosowski, Hearing in Transitional Mammals: Predictions from the Middle-Ear Anatomy and Hearing Capabilities of Extant Mammals, The Evolutionary Biology of Hearing, pp.615-631, 1992.

J. J. Rosowski, The Middle and External Ears of Terrestrial Vertebrates as Mechanical and Acoustic Transducers, Sensors and Sensing in Biology and Engineering, pp.59-70, 2003.

M. R. Rossi-santos, The Spreading Message in a Noisy World: Song Behavior and Acoustic Ecology in Humpback Whales (Megaptera novaeangliae), from the Southwestern Atlantic Ocean, pp.407-440, 2018.

G. E. Rössner, The Evolution of Artiodactyls, pp.213-220, 2007.

P. Roudié, Une richesse nationale africaine. Les phosphates du Togo. Cah. d'outre-mer, vol.30, pp.313-318, 1977.

G. W. Rougier and J. R. Wible, Major Changes in the Ear Region and Basicranium of Early Mammals, Amniote paleobiology: Perspectives on the evolution of mammals, birds, and reptiles, pp.269-311, 2006.

T. Rowe, Definition, Diagnosis, and Origin of Mammalia, J. Vertebr. Paleontol, vol.8, pp.241-264, 1988.

V. Ryabov, Role of the mental foramens in dolphin hearing, Nat. Sci, vol.02, pp.646-653, 2010.

A. Sahni, S. Bhatia, J. Hartenberger, J. Jaeger, K. Kumar et al., Vertebrates from Subathu Formation and comments on the biogeography of Indian subcontinent during early Paleogene, Bull. la Soc. Geol. Fr. S7-XXIII, pp.689-695, 1981.

A. Sahni and S. K. Khare, Three new Eocene mammals from Rajauri District, Jammu and Kashmir. J. Palaeontol. Soc. India, vol.16, pp.41-53, 1971.

R. Sassu and B. Cozzi, The External and Middle Ear of the Striped Dolphin Stenella coeruleoalba (Meyen 1833), Anat. Histol. Embryol. J. Vet. Med. Ser. C, vol.36, pp.197-201, 2007.

B. Schaeffer, Note on the origin and function of the artiodactyl tarsus, Am. Museum Novit, vol.1356, pp.1-24, 1947.

L. Scherler, F. Lihoreau, and D. Becker, To split or not to split Anthracotherium? A phylogeny of Anthracotheriinae (Cetartiodactyla: Hippopotamoidea) and its palaeobiogeographical implications, Zool. J. Linn. Soc, pp.1-24, 2018.
URL : https://hal.archives-ouvertes.fr/hal-02131130

J. Schindelin, I. Arganda-carreras, E. Frise, V. Kaynig, M. Longair et al., Fiji: an open-source platform for biological-image analysis, Nat. Methods, vol.9, pp.676-682, 2012.
URL : https://hal.archives-ouvertes.fr/pasteur-02616466

D. Schluter, T. Price, A. Ø. Mooers, and D. Ludwig, Likelihood of ancestor states in adaptive radiation, Evol. Int. J. Org. Evol, vol.51, pp.1699-1711, 1997.

F. N. Schmidt, M. M. Delsmann, K. Mletzko, T. A. Yorgan, M. Hahn et al., Ultra-high matrix mineralization of sperm whale auditory ossicles facilitates high sound pressure and high-frequency underwater hearing, Proc. R. Soc. B Biol. Sci, vol.285, 2018.

A. V. Schweizer, R. Lebrun, L. A. Wilson, L. Costeur, T. Schmelzle et al., Size Variation under Domestication: Conservatism in the inner ear shape of wolves, dogs and dingoes, Sci. Rep, vol.7, p.13330, 2017.
URL : https://hal.archives-ouvertes.fr/hal-01874750

G. A. Scopoli, Introductio ad historiam naturalem sistens genera lapidum, plantarum, et animalium :hactenus detecta, caracteribus essentialibus donata, tribus divisa, subinde ad leges naturae /Joannis Ant. Scopoli. Prague: Apud Wolfgangum Gerle, 1777.

W. B. Scott, The mammalian fauna of the White River Oligocene. Transaction of the American Philosophical Society, pp.363-746, 1940.

W. Segall, Morphological parallelisms of the bulla and auditory ossicles in some insectivores and marsupials, Fieldiana Zool, vol.51, pp.169-205, 1970.

G. E. Shambaugh, The distribution of blood-vessels in the labyrinth of the ear of Sus scrofa domesticus, pp.137-153, 1903.

M. Shimamura, H. Yasue, K. Ohshima, H. Abe, H. Kato et al., Molecular evidence from retroposons that whales form a clade within even-toed ungulates, Nature, vol.388, pp.666-670, 1997.

A. H. Sileem, H. M. Sallam, A. G. Hewaidy, E. R. Miller, and G. F. Gunnell, A new anthracothere (Artiodactyla) from the early Oligocene, Fayum, Egypt, and the mystery of African "Rhagatherium" solved, J. Paleontol, vol.90, pp.170-181, 2016.

G. G. Simpson, The Principles of Classification and a Classification of Mammals, Bull. Am. Museum Nat. Hist, vol.85, p.350, 1945.

J. S. Sipla and F. Spoor, Sensory Evolution on the Threshold Adaptations in Secondarily Aquatic Vertebrates, pp.227-232, 2008.

S. Sisson, A Text-Book of Veterinary Anatomy, 1910.

G. J. Slater, J. A. Goldbogen, and N. D. Pyenson, Independent evolution of baleen whale gigantism linked to Plio-Pleistocene ocean dynamics, Proc. R. Soc. B Biol. Sci, vol.284, 2017.

A. S. Smith and R. Araújo, Thaumatodracon wiedenrothi, a morphometrically and stratigraphically intermediate new rhomaleosaurid plesiosaurian from the Lower Jurassic (Sinemurian) of Lyme Regis, Palaeontogr. Abteilung A, vol.308, pp.89-125, 2017.

A. N. Soe, O. Chavasseau, Y. Chaimanee, C. Sein, J. Jaeger et al., New remains of Siamotherium pondaungensis (Cetartiodactyla, Hippopotamoidea) from the Eocene of Pondaung, Myanmar: Paleoecologic and phylogenetic implications, J. Vertebr. Paleontol, vol.37, 2017.
URL : https://hal.archives-ouvertes.fr/hal-02328380

G. N. Solntseva, The middle ear in the ontogenesis of mammals, Russ. J. Dev. Biol, vol.42, pp.412-425, 2011.

G. N. Solntseva and V. A. Rodionov, Structural and functional organization of sound generation and sound perception organs in dolphins, Dokl. Biol. Sci, vol.417, pp.476-479, 2007.

M. Spaulding, M. A. O&apos;leary, and J. Gatesy, Relationships of Cetacea (Artiodactyla) Among Mammals: Increased Taxon Sampling Alters Interpretations of Key Fossils and Character Evolution, PLoS One, vol.4, p.7062, 2009.

F. Spoor, S. Bajpai, S. T. Hussain, K. Kumar, and J. G. Thewissen, Vestibular evidence for the evolution of aquatic behaviour in early cetaceans, Nature, vol.417, pp.163-166, 2002.

F. Spoor and J. G. Thewissen, Comparative and Functional Anatomy of Balance in Aquatic Mammals, Sensory Evolution on the Threshold Adaptations in Secondarily Aquatic Vertebrates, pp.257-284, 2008.

M. S. Springer and J. Gatesy, Evolution of the MC5R gene in placental mammals with evidence for its inactivation in multiple lineages that lack sebaceous glands, Mol. Phylogenet. Evol, vol.120, pp.364-374, 2018.

K. M. Stafford, C. Lydersen, Ø. Wiig, and K. Kovacs, Extreme diversity in the songs of Spitsbergen's bowhead whales, Biol. Lett, vol.14, 2018.

M. E. Steeman, A new baleen whale from the Late Miocene of Denmark and early mysticete hearing, Palaeontology, vol.52, pp.1169-1190, 2009.

H. G. Stehlin, Remarques sur les faunules de Mammifères des couches éocènes et oligocènes du Bassin de Paris, vol.4, pp.488-520, 1910.

A. S. Stoeger and A. Baotic, Male African elephants discriminate and prefer vocalizations of unfamiliar females, Sci. Rep, vol.7, p.46892, 2017.

E. Stromer, Die Archaeoceti des Ägyptischen Eozäns, 1908.

J. Sudre, Les Artiodactyles de l'Éocène moyen et supérieur d'Europe occidentale (Systématique et évolution), 1978.

V. Suteethorn, E. Buffetaut, R. Helmcke-ingavat, J. Jaeger, and Y. Jongkanjanasoontorn, Oldest known Tertiary mammals from South East Asia : middle Eocene primate and anthracotheres from Thailand, Neues Jahrb. für Geol. und Paläontologie, Monatshefte, vol.9, pp.563-570, 1988.

D. L. Swofford, PAUP (Phylogenetic Analysis Using Parsimony), Version 2.4. User's manual, Illinois Natural History Survey, 1985.

D. L. Swofford, PAUP*. Phylogenetic Analysis Using Parsimony (*and Other Methods), 2002.

D. L. Swofford and G. J. Olsen, Phylogeny reconstruction, Molecular systematics, pp.411-501, 1990.

E. V. Sysueva, P. E. Nachtigall, T. W. Cranford, A. F. Pacini, J. L. Pawloski et al., Auditory sensitivity areas of head to local underwater stimulation in a bottlenose dolphin (Tursiops truncatus), Proc. Meet. Acoust, vol.30, p.10003, 2018.

J. V. Tejada-lara, R. Salas-gismondi, F. Pujos, P. Baby, M. Benammi et al., Life in proto-Amazonia: Middle Miocene mammals from the Fitzcarrald Arch (Peruvian Amazonia), Palaeontology, vol.58, pp.341-378, 2015.
URL : https://hal.archives-ouvertes.fr/hal-01922735

J. M. Theodor, J. Erfurt, and G. Métais, The Earliest Artiodactyls: Diacodexeidae, Dichobunidae, Homacodontidae, Leptochoeridae, and Raoellidae, pp.32-58, 2007.

J. M. Theodor and S. E. Foss, Deciduous dentitions of Eocene cebochoerid artiodactyls and cetartiodactyl relationships, J. Mamm. Evol, vol.12, pp.161-181, 2005.

J. G. Thewissen, Phylogenetic aspects of Cetacean origins: A morphological perspective, J. Mamm. Evol, vol.2, pp.157-184, 1994.

J. G. Thewissen, The walking whales: from land to water in eight million years, 2014.

J. G. Thewissen, L. N. Cooper, M. T. Clementz, S. Bajpai, and B. N. Tiwari, Whales originated from aquatic artiodactyls in the Eocene epoch of India, Nature, vol.450, pp.1190-1194, 2007.

J. G. Thewissen, L. N. Cooper, J. C. George, and S. Bajpai, From Land to Water: The Origin of Whales, Dolphins, and Porpoises, Evol. Educ. Outreach, vol.2, pp.272-288, 2009.

J. G. Thewissen and D. P. Domning, The role of phenacodontids in the origin of the modern orders of ungulate mammals, J. Vertebr. Paleontol, vol.12, pp.494-504, 1992.

J. G. Thewissen, P. D. Gingerich, and D. E. Russell, Artiodactyla and Perissodactyla (Mammalia) from the early-middle Eocene Kuldana formation of Kohat (Pakistan), Contrib. from Museum Paleontol. Univ. Michigan, vol.27, pp.247-274, 1987.

J. G. Thewissen and S. T. Hussain, Postcranial Osteology of the most Primitive Artiodactyl: Diacodexis pakistanensis (Dichobunidae), Anat. Histol. Embryol, vol.19, pp.37-48, 1990.

J. G. Thewissen and S. T. Hussain, Origin of underwater hearing in whales, Nature, vol.361, pp.444-445, 1993.

J. G. Thewissen, S. T. Hussain, and M. Arif, Fossil Evidence for the Origin of Aquatic Locomotion in Archaeocete Whales, Science, vol.263, pp.210-212, 1994.

J. G. Thewissen and S. I. Madar, Ankle morphology of the earliest cetaceans and its implications for the phylogenetic relations among ungulates, Syst. Biol, vol.48, pp.21-30, 1999.

J. G. Thewissen, S. I. Madar, and S. T. Hussain, Ambulocetus natans, an Eocene cetacean (Mammalia) from Pakistan, CFS Cour. Forschungsinstitut Senckenb, pp.1-86, 1996.

J. G. Thewissen and S. Nummela, Sensory Evolution on the Threshold Adaptations in Secondarily Aquatic Vertebrates, 2008.

J. G. Thewissen, L. J. Roe, J. R. O&apos;neil, S. T. Hussain, A. Sahni et al., Evolution of cetacean osmoregulation, Nature, vol.381, pp.379-380, 1996.

J. G. Thewissen, J. D. Sensor, M. T. Clementz, and S. Bajpai, Evolution of dental wear and diet during the origin of whales, Paleobiology, vol.37, pp.655-669, 2011.

J. G. Thewissen and E. M. Williams, The Early Radiations of Cetacea (Mammalia): Evolutionary Pattern and Developmental Correlations, Annu. Rev. Ecol. Syst, vol.33, pp.73-90, 2002.

J. G. Thewissen, E. M. Williams, and S. T. Hussain, Eocene mammal faunas from nothern Indo-Pakistan, J. Vertebr. Paleontol, vol.21, pp.347-366, 2001.

J. G. Thewissen, E. M. Williams, L. J. Roe, and S. T. Hussain, Skeletons of terrestrial cetaceans and the relationship of whales to artiodactyls, Nature, vol.413, pp.277-281, 2001.

S. W. Tinker, Whales of the World, 1988.

J. Tonndorf, A New Concept of Bone Conduction, Arch. Otolaryngol, vol.87, pp.595-600, 1968.

S. J. Trumble, S. A. Norman, D. D. Crain, F. Mansouri, Z. C. Winfield et al., Baleen whale cortisol levels reveal a physiological response to 20th century whaling, Nat. Commun, vol.9, p.4587, 2018.

S. J. Trumble, E. M. Robinson, M. Berman-kowalewski, C. W. Potter, and S. Usenko, Blue whale earplug reveals lifetime contaminant exposure and hormone profiles, Proc. Natl. Acad. Sci, vol.110, pp.16922-16926, 2013.

G. Tsagkogeorga, M. R. Mcgowen, K. T. Davies, S. Jarman, A. Polanowski et al., A phylogenomic analysis of the role and timing of molecular adaptation in the aquatic transition of cetartiodactyl mammals, R. Soc. Open Sci, vol.2, p.150156, 2015.

C. Tsai and N. Kohno, Multiple origins of gigantism in stem baleen whales, Sci. Nat, vol.103, p.89, 2016.

T. Tsubamoto, N. Egi, M. Takai, and . Thaung-htike, Zin-Maung-Maung-Thein. 2013. A New Genus and Species of Bunodont Artiodactyl from the Eocene Pondaung Formation, vol.17, pp.297-311

A. A. Tubelli, A. Zosuls, D. R. Ketten, and D. C. Mountain, A model and experimental approach to the middle ear transfer function related to hearing in the humpback whale, 2018.

, J. Acoust. Soc. Am, vol.144, pp.525-535

P. L. Tyack, Communication and cognition, Biology of Marine Mammals. Washington and London, pp.287-323, 1999.

M. D. Uhen, Middle to Late Eocene Basilosaurines and Dorudontines, pp.29-62, 1998.

M. D. Uhen, New protocetid (Mammalia, Cetacea) from the late middle Eocene Cook Mountain Formation of Louisiana, J. Vertebr. Paleontol, vol.18, pp.664-668, 1998.

M. D. Uhen, Form, Function, and Anatomy of Dorudon atrox (Mammalia, Cetacea): An Archaeocete from the Middle to Late Eocene of Egypt, Univ. Michigan -Pap. Paleontol, vol.34, pp.1-222, 2004.

M. D. Uhen, Evolution of Tertiary Mammals of North America, Small Mammals, Xenarthrans, and Marine Mammals, vol.2, pp.557-565, 2007.

M. D. Uhen, New material of Natchitochia jonesi and a comparison of the innominata and locomotor capabilities of Protocetidae, Mar. Mammal Sci, vol.30, pp.1029-1066, 2014.

M. D. Uhen, New specimens of Protocetidae (Mammalia, Cetacea) from New Jersey and South Carolina, J. Vertebr. Paleontol, vol.34, pp.211-219, 2014.

N. Vandenberghe, F. J. Hilgen, R. P. Speijer, J. G. Ogg, F. M. Gradstein et al., The Paleogene Period, The Geologic Time Scale, pp.855-921, 2012.

U. Varanasi and D. C. Malins, Unique lipids of the porpoise (Tursiops gilli): Differences in triacyl glycerols and wax esters of acoustic (mandibular canal and melon) and blubber tissues, Biochim. Biophys. Acta -Lipids Lipid Metab, vol.231, pp.415-418, 1971.

U. Varanasi and D. C. Malins, Triacylglycerols characteristic of porpoise acoustic tissues: molecular structures of diisovaleroylglycerides. Science (80-. ), vol.176, pp.926-934, 1972.

Q. Vautrin, F. Lihoreau, B. Sambou, M. Thiam, J. E. Martin et al., From limb to fin: an Eocene protocetid forelimb from Senegal sheds new light on the early locomotor evolution of cetaceans, 2019.
URL : https://hal.archives-ouvertes.fr/hal-02302367

M. Viglino, M. R. Buono, C. S. Gutstein, M. Cozzuol, and J. Cuitiño, A new dolphin from the early Miocene of Patagonia (Argentina): insights into the evolution of Platanistoidea in the Southern Hemisphere, Acta Palaeontol. Pol, vol.63, 2018.

I. A. Vislobokova and B. A. Trofimov, Archaeomeryx (Archaeomerycidae, Ruminantia): Morphology, Ecology, and Role in the Evolution of the Artiodactyla, Paleontol. J, vol.36, 2002.

, Avizo: 3D Analysis Software for Scientific and Industrial Data, 2018.

S. Vreese, . De, M. Doom, J. Haelters, and P. Cornillie, Heeft de uitwendige gehoorgang van walvisachtigen nog enige functie? Vlaams Diergeneeskd, Tijdschr, vol.83, pp.284-292, 2014.

S. Vreese, . De, S. Mazzariol, and M. André, Comparative morphology of the external ear canal in several species of odontocetes, Eff. Noise Aquat. Life Fifth Int, p.81, 2019.

P. J. Waddell, N. Okada, and M. Hasegawa, Towards resolving the interordinal relationships of placental mammals, Syst. Biol, vol.48, pp.1-5, 1999.

A. R. Wallace, On the Law which has Regulated the Introduction of New Species, Ann. Mag. Nat. Hist, vol.16, pp.184-196, 1855.

D. Wartzok and D. R. Ketten, Marine Mammal Sensory Systems, Biology of Marine Mammals. Washington and London, pp.117-136, 1999.

S. D. Webb and B. E. Taylor, The phylogeny of hornless ruminants and a description of the cranium of Archaeomeryx, Bull. Am. Museum Nat. Hist, vol.167, pp.117-158, 1980.

C. D. West, The relationship of the spiral turns of the cochlea and the length of the basilar membrane to the range of audible frequencies in ground dwelling mammals, J. Acoust. Soc. Am, vol.77, pp.1091-1101, 1985.

R. M. West, Middle Eocene Large Mammal Assemblage with Tethyan Affinities , Ganda Kas Region , Pakistan, J. Paleontol, vol.54, pp.508-533, 1980.

E. M. Weston and J. Boisserie, Cenozoic Mammals of Africa, pp.853-872, 2010.

E. G. Wever, J. G. Mccormick, J. Palin, and S. H. Ridgway, Cochlea of the Dolphin, Tursiops truncatus: The Basilar Membrane, Proc. Natl. Acad. Sci, vol.68, pp.2708-2711, 1971.

E. G. Wever, J. G. Mccormick, J. Palin, and S. H. Ridgway, The Cochlea of the Dolphin, Tursiops truncatus: General Morphology, Proc. Natl. Acad. Sci, vol.68, pp.2381-2385, 1971.

E. G. Wever, J. G. Mccormick, J. Palin, and S. H. Ridgway, The Cochlea of the Dolphin, Tursiops truncatus: Hair Cells and Ganglion Cells, Proc. Natl. Acad. Sci, vol.68, pp.2908-2912, 1971.

J. R. Wible, Cranial Circulation and Relationships of the Colugo Cynocephalus (Dermoptera, Mammalia), Am. Museum Novit, vol.3072, pp.1-27, 1993.

J. R. Wible, On the cranial osteology of the short-tailed opossum Monodelphis brevicaudata (Didelphidae, Marsupialia), Ann. Carnegie Museum, vol.72, pp.137-202, 2003.

H. C. Wilkie, The Auditory Organ of the Ox (Bos taurus ), Proc. Zool. Soc. London, vol.106, pp.985-1009, 1936.

E. M. Williams, Synopsis of the Earliest Cetaceans Pakicetidae, Ambulocetidae, Remingtonocetidae, and Protocetidae, pp.1-28, 1998.

X. Wu and L. A. Schepartz, Application of computed tomography in paleoanthropological research, Prog. Nat. Sci, vol.19, pp.913-921, 2009.

M. Yamada, Contribution to the anatomy of the organ of hearing of whales, Sci. Reports Whales Res. Inst, vol.8, pp.3-79, 1953.

M. Yamada and F. Yoshizaki, Osseous labyrinth of Cetacea. Sci. Reports Whales Res. Inst, vol.14, pp.291-304, 1959.

M. Yamato, D. R. Ketten, J. Arruda, S. R. Cramer, and K. Moore, The Auditory Anatomy of the Minke Whale (Balaenoptera acutorostrata): A Potential Fatty Sound Reception Pathway in a Baleen Whale, Anat. Rec, vol.295, pp.991-998, 2012.

M. Yamato, K. Khidas, N. D. Pyenson, R. E. Fordyce, and J. G. Mead, Extensively remodeled, fractured cetacean tympanic bullae show that whales can survive traumatic injury to the ears, J. Anat, vol.228, pp.125-136, 2016.

M. Yamato, H. Koopman, M. Niemeyer, and D. R. Ketten, Characterization of lipids in adipose depots associated with minke and fin whale ears: Comparison with "acoustic fats" of toothed whales, Mar. Mammal Sci, vol.30, pp.1549-1563, 2014.

M. Yamato and N. D. Pyenson, Early development and orientation of the acoustic funnel provides insight into the evolution of sound reception pathways in cetaceans, PLoS One, vol.10, pp.1-15, 2015.

M. L. Zelditch, D. L. Swiderski, H. D. Sheets, and W. L. Fink, Geometric Morphometrics for Biologists, 2004.

X. Zhou, S. Xu, Y. Yang, K. Zhou, and G. Yang, Phylogenomic analyses and improved resolution of Cetartiodactyla, Mol. Phylogenet. Evol, vol.61, pp.255-264, 2011.

J. P. Zurano, F. M. Magalhães, A. E. Asato, G. Silva, C. J. Bidau et al., Cetartiodactyla: Updating a time-calibrated molecular phylogeny, Mol. Phylogenet. Evol, vol.133, pp.256-262, 2019.

, Références bibliographiques

C. W. Andrews, A description of new species of zeuglodont and of leathery turtle from the Eocene of southern Nigeria, Proc. Zool. Soc. London, vol.1919, pp.309-319, 1920.

S. Bajpai and J. G. Thewissen, Middle Eocene Cetaceans from the Harudi and Subathu Formations of India, pp.213-234, 1998.

S. Bajpai and J. G. Thewissen, Protocetid cetaceans (Mammalia) from the Eocene of India, 2014.

P. Batik and O. Fejfar, Les vertébrés du Lutétien du Miocène et du Pliocène de Tunisie centrale, Notes Serv. Géologique Tunisie, vol.56, pp.69-80, 1990.

G. Bianucci and P. D. Gingerich, Aegyptocetus tarfa, n. gen. et sp. (Mammalia, Cetacea), from the middle Eocene of Egypt: Clynorhynchy, olfaction, and hearing in protocetid whale, J. Vertebr. Paleontol, vol.31, pp.1173-1188, 2011.

C. J. Burgin, J. P. Colella, P. L. Kahn, and N. S. Upham, How many species of mammals are there?, J. Mammal, vol.99, pp.1-14, 2018.

E. Fraas, Neue Zeuglodonten aus dem unteren Mitteleocän vom Mokattam bei Cairo, Geol. und Palaontologische Abhandlungen, vol.6, pp.197-220, 1904.

J. H. Geisler, A. E. Sanders, and Z. Luo, A New Protocetid Whale (Cetacea: Archaeoceti) from the Late Middle Eocene of South Carolina, Am. Museum Novit, vol.3480, pp.1-65, 2005.

M. L. Gibson, J. Mnieckowski, and J. Geisler, Tupelocetus palmeri, a new species of protocetid whale (Mammalia, Cetacea) from the middle Eocene of South Carolina, J. Vertebr. Paleontol, vol.38, p.1555165, 2018.

P. D. Gingerich, Partial skeleton of a new archaeocete from the earliest middle Eocene Habib Rahi limestone, Pakistan. J. Vertebr. Paleontol, vol.11, p.31, 1991.

P. D. Gingerich, Cetacea. Cenozoic Mammals of Africa, pp.873-900, 2010.

P. D. Gingerich, M. Arif, and W. C. Clyde, New archaeocetes (Mammalia, Cetacea) from the Middle Eocene Domanda Formation of the Sulaiman Range, 1995.

P. D. Gingerich and H. Cappetta, A new archaeocete and other marine mammals (Cetacea and Sirenia) from lower middle Eocene phosphate deposits of Togo, J. Paleontol, vol.88, pp.109-129, 2014.

P. D. Gingerich, S. M. Raza, M. Arif, M. Anwar, and X. Zhou, New whale from the Eocene of Pakistan and the origin of cetacean swimming, Nature, vol.368, pp.844-847, 1994.

P. D. Gingerich, M. Ul-haq, W. Von-koenigswald, W. J. Sanders, B. H. Smith et al., New protocetid whale from the middle eocene of pakistan: birth on land, precocial development, and sexual dimorphism, PLoS One, vol.4, p.4366, 2009.

P. D. Gingerich, M. Ul-haq, I. S. Zalmout, I. H. Khan, and M. S. Malkani, Origin of Whales from Early Artiodactyls: Hands and Feet of Eocene Protocetidae from Pakistan. Science (80-. ), vol.293, pp.2239-2242, 2001.

P. D. Gingerich, I. S. Zalmout, M. Ul-haq, A. Bhatti, and M. , Makaracetus bidens, a new protocetid archaeocete (Mammalia, Cetacea) from the early middle Eocene of, 2005.

, Contrib. from Museum Paleontol. Univ. Michigan, vol.31, pp.197-210

P. D. Gingerich and S. Zouhri, New fauna of archaeocete whales (Mammalia, Cetacea) from the Bartonian middle Eocene of southern Morocco, J. African Earth Sci, 2015.

S. J. Godfrey, J. Geisler, and E. M. Fitzgerald, On the Olfactory Anatomy in an Archaic Whale (Protocetidae, Cetacea) and the Minke Whale Balaenoptera acutorostrata (Balaenopteridae, Cetacea), Anat. Rec, vol.296, pp.257-272, 2013.

A. S. Gohar, M. S. Antar, S. El-sayed, and H. Sallam, Egyptian Eocene archaeocetes Mammalia Cetacea Review and new discoveries. 56th, Annu. Sci. Meet. Geol. Soc. Egypt, pp.31-32, 2018.

L. Hautier, R. Sarr, F. Lihoreau, R. Tabuce, M. Hameh et al., First record of the family Protocetidae in the Lutetian of Senegal (West Africa), Palaeovertebrata, vol.38, pp.1-7, 2014.
URL : https://hal.archives-ouvertes.fr/hal-01923037

R. C. Hulbert, R. M. Petkewich, G. A. Bishop, D. Bukry, and D. P. Aleshire, A new middle Eocene protocetid whale (Mammalia: Cetacea: Archaeoceti) and associated biota from Georgia, J. Paleontol, 1998.

O. Lambert, C. Muizon, . De, E. Malinverno, C. Celma et al., A new odontocete (toothed cetacean) from the Early Miocene of Peru expands the morphological disparity of extinct heterodont dolphins, J. Syst. Palaeontol, vol.16, pp.981-1016, 2018.
URL : https://hal.archives-ouvertes.fr/hal-02612079

J. Leidy, Contributions to the extinct vertebrate fauna of the Western Territories, Rep. U.S. Geol. Surv. Territ, p.337, 1873.

S. A. Mcleod and L. G. Barnes, A new genus and species of Eocene protocetid archaeocete whale (Mammalia, Cetacea) from the Atlantic Coastal Plain, Nat. Hist. Museum Los Angeles Cty. -Sci. Ser, vol.41, pp.73-98, 2008.

A. Sahni and V. P. Mishra, Lower Tertiary Vertebrates from Western India, Monogr. Paleontol. Soc. India, vol.3, pp.1-48, 1975.

A. N. Trivedy and P. P. Satsangi, A new archaeocete (whale) from the Eocene of India, Abstr. 27th Intern. Geol. Congr. Moscow, pp.322-323, 1984.

M. D. Uhen, New protocetid (Mammalia, Cetacea) from the late middle Eocene Cook Mountain Formation of Louisiana, J. Vertebr. Paleontol, vol.18, pp.664-668, 1998.

M. D. Uhen, New specimens of Protocetidae (Mammalia, Cetacea) from New Jersey and South Carolina, J. Vertebr. Paleontol, vol.34, pp.211-219, 2014.

A. W. Wight, Paleogene Vertebrae Fauna and Regressive Sediment of Dur at Talhah Southern Sirt Basin Libya, pp.309-325, 1980.

, Remingtonocetus harudiensis VPL/K 15001

&. Kumar and . Sahni, , 1986.

G. Bianucci and P. D. Gingerich, Aegyptocetus tarfa, n. gen. et sp. (Mammalia, Cetacea), from the middle Eocene of Egypt: Clinorhynchy, olfaction, and hearing in a protocetid whale, Journal of Vertebrate Paleontology, vol.31, issue.6, pp.1173-1188, 2011.

E. M. Fitzgerald, The morphology and systematics of Mammalodon colliveri (Cetacea: Mysticeti), a toothed mysticete from the Oligocene of Australia, Zoological Journal of the Linnean Society, vol.158, pp.367-476, 2010.

R. E. Fordyce, Simocetus rayi (Odontoceti: Simocetidae, New Family): A bizarre new archaic Oligocene Dolphin from the Eastern North Pacific, Smithsonian Contributions to Paleobiology, vol.93, pp.185-222, 2002.

J. H. Geisler, A. E. Sanders, and Z. X. Luo, A new protocetid whale (Cetacea: Archaeoceti) from the late middle Eocene of South Carolina, American Museum Novitates, vol.3480, pp.1-65, 2005.

P. D. Gingerich, M. Arif, M. A. Bhatti, H. A. Raza, and S. M. Raza, Protosiren and Babiacetus (Mammalia, Sirenia and Cetacea) from the middle Eocene Drazinda formation, Sulaiman range, Contributions from the Museum of Paleontology, vol.29, p.357, 1995.

P. D. Gingerich, M. Arif, and W. C. Clyde, New archaeocetes (Mammalia, Cetacea) from the middle Eocene Domanda formation of the Sulaiman range, Contributions from the Museum of Paleontology, vol.29, pp.291-330, 1995.

P. D. Gingerich, M. Haq, I. H. Khan, and I. S. Zalmout, Eocene stratigraphy and archaeocete whales (Mammalia, Cetacea) of Drug Lahar in the eastern Sulaiman range, Contributions from the Museum of Paleontology, vol.30, pp.269-319, 2001.

P. D. Gingerich, M. Haq, W. Koenigswald, W. J. Sanders, B. H. Smith et al., New protocetid whale from the middle Eocene of Pakistan: birth on land, precocial development and sexual dimorphism, PloS ONE, vol.4, issue.2, pp.4366-4367, 2009.

P. D. Gingerich, S. M. Raza, M. Arif, M. Anwar, and X. Zhou, Partial skeletons of Indocetus ramani (Mammalia, Cetacea) from the lower middle Eocene Domanda shale in the Sulaiman range, Contributions from the Museum of Paleontology, vol.28, pp.393-416, 1993.

P. D. Gingerich, S. M. Raza, M. Arif, M. Anwar, and X. Zhou, New whale from the Eocene of Pakistan and the origin of cetacean swimming, Nature, vol.368, pp.844-847, 1994.

P. D. Gingerich and M. D. Uhen, Ancalecetus simonsi, a new dorudontine archaeocete (Mammalia, Cetacea) from the early late Eocene of Wadi Hitan, Egypt. Contributions from the Museum of Paleontology, vol.29, pp.359-401, 1996.

P. Gol&apos;din and E. Zvonok, Basilotritus uheni, a new cetacean (Cetacea, Basilosauridae) from the late middle Eocene of Eastern Europe, Journal of Paleontology, vol.87, issue.2, pp.254-268, 2013.

J. Hector, Note on New Zealand Cetacea, recent and fossil, Transactions and Proceedings of the New Zealand Institute, vol.13, pp.434-437, 1881.

R. C. Hulbert, R. M. Petkewich, G. A. Bishop, D. Bukry, and D. P. Aleshire, A new middle Eocene protocetid whale (Mammalia: Cetacea: Archaeoceti) and associated biota from Georgia, Journal of Paleontology, vol.72, issue.5, pp.907-927, 1998.

R. Kellogg, A review of the Archaeoceti. Canergie Institution of Washington, vol.482, pp.1-366, 1936.

R. Köhler and R. E. Fordyce, An archaeocete whale (Cetacea: Archaeoceti) from the Eocene Waihao Greensand, Journal of Vertebrate Paleontology, vol.17, issue.3, pp.574-583, 1997.

K. Kumar and A. Sahni, Remingtonocetus harudiensis, new combination, a middle Eocene archaeocete (Mammalia, Cetacea) from western Kutch, India. Journal of Vertebrate Paleontology, vol.6, issue.4, pp.326-349, 1986.

W. C. Lancaster, The middle ear of the Archaeoceti, Journal of Vertebrate Paleontology, vol.10, issue.1, pp.117-127, 1990.

Z. X. Luo, Homology and transformation of cetacean ectotympanic structures, The Emergence of Whales: evolutionary patterns in the origin of Cetacea, pp.269-301, 1998.

Z. X. Luo and P. D. Gingerich, Terrestrial Mesonychia to aquatic Cetacea: transformation of the basicranium and evolution of hearing in whales, vol.31, pp.1-98, 1999.

M. Martínez-cáceres and C. D. Muizon, A new basilosaurid (Cetacea, Pelagiceti) from the Late Eocene to Early Oligocene Otuma Formation of Peru, Comptes Rendus Palevol, vol.10, pp.517-526, 2011.

M. Martínez-cáceres, L'enracinement des cétacés modernes (Neoceti : Mysticeti + Odontoceti) au sein des archéocètes : les derniers archéocètes (Basilosauridae), les premiers néocètes, 2013.

. Paléontologie and C. D. De-muizon, Muséum national d'Histoire naturelle, vol.603

S. Nummela, S. Taseer-hussain, and J. G. Thewissen, Cranial anatomy of Pakicetidae (Cetacea, Mammalia), Journal of Vertebrate Paleontology, vol.26, issue.3, pp.746-759, 2006.

M. A. O&apos;leary, An anatomical and phylogenetic study of the osteology of the petrosal of extant and extinct artiodactylans (Mammalia) and relatives, Bulletin of the American Museum of Natural History, vol.335, pp.1-226, 2010.

J. G. Thewissen and S. Bajpai, New skeletal material of Andrewsiphius and Kutchicetus, two Eocene cetaceans from India, Journal of Paleontology, vol.83, issue.5, pp.635-663, 2009.

M. D. Uhen, Form, function, and anatomy of Dorudon atrox (Mammalia, Cetacea): an archaeocete from the middle to late Eocene of Egypt. The University of Michigan Museum of Paleontology Papers on Paleontology, vol.34, pp.1-222, 2004.

M. D. Uhen and P. D. Gingerich, New genus of dorudontine archaeocete (Cetacea) from the middle-to-late Eocene of South Carolina, Marine Mammal Science, vol.17, issue.1, pp.1-34, 2001.

M. D. Uhen, N. D. Pyenson, T. J. Devries, M. Urbina, and P. R. Renne, New middle Eocene whales from the Pisco Basin of Peru, Journal of Paleontology, vol.85, issue.5, pp.955-969, 2011.

E. G. Ekdale, A. Berta, and T. A. Deméré, The comparative osteology of the petrotympanic complex (ear region) of extant baleen whales (Cetacea: Mysticeti), PloS ONE, vol.6, issue.6, pp.21311-21312, 2011.

J. H. Geisler and Z. X. Luo, The petrosal and inner ear of Herpetocetus sp. (Mammalia: Cetacea) and their implications for the phylogeny and hearing of archaic mysticetes, Journal of Paleontology, vol.70, issue.6, pp.1045-1066, 1996.

J. H. Geisler, A. E. Sanders, and Z. X. Luo, A new protocetid whale (Cetacea: Archaeoceti) from the late middle Eocene of South Carolina, American Museum Novitates, vol.3480, pp.1-65, 2005.

T. Kasuya, Systematic consideration of recent toothed whales based on the morphology of tympano-periotic bone, Scientific Reports of the Whale Research Institute, vol.25, pp.1-103, 1973.

R. Kellogg, The history of whales-their adaptation to the life in the water, Quarterly Review of Biology, vol.3, pp.174-208, 1928.

R. Kellogg, A review of the Archaeoceti. Canergie Institution of Washington, vol.482, pp.1-366, 1936.

W. C. Lancaster, The middle ear of the Archaeoceti, Journal of Vertebrate Paleontology, vol.10, issue.1, pp.117-127, 1990.

Z. X. Luo, Homology and transformation of cetacean ectotympanic structures, The Emergence of Whales: evolutionary patterns in the origin of Cetacea, pp.269-301, 1998.

Z. X. Luo and E. R. Eastman, Petrosal and inner ear of a Squalodontid whale: Implications for evolution of hearing in Odontocetes, Journal of Vertebrate Paleontology, vol.15, issue.2, pp.431-442, 1995.

Z. X. Luo and P. D. Gingerich, Terrestrial Mesonychia to aquatic Cetacea: transformation of the basicranium and evolution of hearing in whales, vol.31, pp.1-98, 1999.

Z. X. Luo and K. Marsh, Petrosal (periotic) and inner ear of a pliocene kogiine whale (Kogiinae, Odontoceti): Implications on relationships and hearing evolution of toothed whales, Journal of Vertebrate Paleontology, vol.16, issue.2, pp.328-348, 1996.

M. Martínez-cáceres, L'enracinement des cétacés modernes (Neoceti : Mysticeti + Odontoceti) au sein des archéocètes : les derniers archéocètes (Basilosauridae), les premiers néocètes, 2013.

M. A. O&apos;leary, An anatomical and phylogenetic study of the osteology of the petrosal of extant and extinct artiodactylans (Mammalia) and relatives, Bulletin of the American Museum of Natural History, vol.335, pp.1-226, 2010.

G. Pilleri, M. Gihr, and C. Kraus, The organ of Corti in cetaceans -1. Recent Species. Investigations on Ceatacea, vol.20, pp.43-125, 1987.

J. R. Wible, On the cranial osteology of the short-tailed opossum Monodelphis brevicaudata (Didelphidae, Marsupialia), vol.72, pp.137-202, 2003.

G. Bianucci and P. D. Gingerich, Aegyptocetus tarfa, n. gen. et sp. (Mammalia, Cetacea), from the middle Eocene of Egypt: Clinorhynchy, olfaction, and hearing in a protocetid whale, Journal of Vertebrate Paleontology, vol.31, issue.6, pp.1173-1188, 2011.

E. M. Fitzgerald, The morphology and systematics of Mammalodon colliveri (Cetacea: Mysticeti), a toothed mysticete from the Oligocene of Australia, Zoological Journal of the Linnean Society, vol.158, pp.367-476, 2010.

R. E. Fordyce, Simocetus rayi (Odontoceti: Simocetidae, New Family): A bizarre new archaic Oligocene Dolphin from the Eastern North Pacific, Smithsonian Contributions to Paleobiology, vol.93, pp.185-222, 2002.

J. H. Geisler, A. E. Sanders, and Z. X. Luo, A new protocetid whale (Cetacea: Archaeoceti) from the late middle Eocene of South Carolina, American Museum Novitates, vol.3480, pp.1-65, 2005.

P. D. Gingerich, M. Haq, W. Koenigswald, W. J. Sanders, B. H. Smith et al., New protocetid whale from the middle Eocene of Pakistan: birth on land, precocial development and sexual dimorphism, PloS ONE, vol.4, issue.2, pp.4366-4367, 2009.

P. D. Gingerich and M. D. Uhen, Ancalecetus simonsi, a new dorudontine archaeocete (Mammalia, Cetacea) from the early late Eocene of Wadi Hitan, Egypt. Contributions from the Museum of Paleontology, vol.29, pp.359-401, 1996.

P. Gol&apos;din and E. Zvonok, Basilotritus uheni, a new cetacean (Cetacea, Basilosauridae) from the late middle Eocene of Eastern Europe, Journal of Paleontology, vol.87, issue.2, pp.254-268, 2013.

R. C. Hulbert, R. M. Petkewich, G. A. Bishop, D. Bukry, and D. P. Aleshire, A new middle Eocene protocetid whale (Mammalia: Cetacea: Archaeoceti) and associated biota from Georgia, Journal of Paleontology, vol.72, issue.5, pp.907-927, 1998.

R. Kellogg, A review of the Archaeoceti. Canergie Institution of Washington, vol.482, pp.1-366, 1936.

R. Köhler and R. E. Fordyce, An archaeocete whale (Cetacea: Archaeoceti) from the Eocene Waihao Greensand, Journal of Vertebrate Paleontology, vol.17, issue.3, pp.574-583, 1997.

K. Kumar and A. Sahni, Remingtonocetus harudiensis, new combination, a middle Eocene archaeocete (Mammalia, Cetacea) from western Kutch, India. Journal of Vertebrate Paleontology, vol.6, issue.4, pp.326-349, 1986.

W. C. Lancaster, The middle ear of the Archaeoceti, Journal of Vertebrate Paleontology, vol.10, issue.1, pp.117-127, 1990.

Z. X. Luo and P. D. Gingerich, Terrestrial Mesonychia to aquatic Cetacea: transformation of the basicranium and evolution of hearing in whales, vol.31, pp.1-98, 1999.

M. Martínez-cáceres and C. D. Muizon, A new basilosaurid (Cetacea, Pelagiceti) from the Late Eocene to Early Oligocene Otuma Formation of Peru, Comptes Rendus Palevol, vol.10, pp.517-526, 2011.

M. Martínez-cáceres, L'enracinement des cétacés modernes (Neoceti : Mysticeti + Odontoceti) au sein des archéocètes : les derniers archéocètes (Basilosauridae), les premiers néocètes, 2013.

S. Nummela, S. Taseer-hussain, and J. G. Thewissen, Cranial anatomy of Pakicetidae (Cetacea, Mammalia), Journal of Vertebrate Paleontology, vol.26, issue.3, pp.746-759, 2006.

M. A. O&apos;leary, An anatomical and phylogenetic study of the osteology of the petrosal of extant and extinct artiodactylans (Mammalia) and relatives, Bulletin of the American Museum of Natural History, vol.335, pp.1-226, 2010.

M. J. Orliac, F. Guy, and R. Lebrun, Osteological connections of the petrosal bone of the extant Hippopotamidae Hippopotamus amphibious and Choeropsis liberiensis, MorphoMuseuM, vol.1, issue.1, pp.1-6, 2014.

M. J. Orliac, The petrosal bone of extinct Suoidea (Mammalia, Artiodactyla), Journal of Systematic Palaeontology, vol.11, issue.8, pp.925-945, 2013.

J. G. Thewissen and S. Bajpai, New skeletal material of Andrewsiphius and Kutchicetus, two Eocene cetaceans from India, Journal of Paleontology, vol.83, issue.5, pp.635-663, 2009.

M. D. Uhen, Form, function, and anatomy of Dorudon atrox (Mammalia, Cetacea): an archaeocete from the middle to late Eocene of Egypt. The University of Michigan Museum of Paleontology Papers on Paleontology, vol.34, pp.1-222, 2004.

M. D. Uhen and P. D. Gingerich, New genus of dorudontine archaeocete (Cetacea) from the middle-to-late Eocene of South Carolina, Marine Mammal Science, vol.17, issue.1, pp.1-34, 2001.

M. D. Uhen, N. D. Pyenson, T. J. Devries, M. Urbina, and P. R. Renne, New middle Eocene whales from the Pisco Basin of Peru, Journal of Paleontology, vol.85, issue.5, pp.955-969, 2011.

P. D. Gingerich, M. Arif, M. A. Bhatti, H. A. Raza, and S. M. Raza, Protosiren and Babiacetus (Mammalia, Sirenia and Cetacea) from the middle Eocene Drazinda formation, Sulaiman range, Contributions from the Museum of Paleontology, vol.29, p.357, 1995.

P. D. Gingerich, M. Arif, and W. C. Clyde, New archaeocetes (Mammalia, Cetacea) from the middle Eocene Domanda formation of the Sulaiman range, Contributions from the Museum of Paleontology, vol.29, pp.291-330, 1995.

P. D. Gingerich, M. Haq, I. H. Khan, and I. S. Zalmout, Eocene stratigraphy and archaeocete whales (Mammalia, Cetacea) of Drug Lahar in the eastern Sulaiman range, Contributions from the Museum of Paleontology, vol.30, pp.269-319, 2001.

P. D. Gingerich, M. Haq, W. Koenigswald, W. J. Sanders, B. H. Smith et al., New protocetid whale from the middle Eocene of Pakistan: birth on land, precocial development and sexual dimorphism, PloS ONE, vol.4, issue.2, pp.4366-4367, 2009.

P. D. Gingerich, S. M. Raza, M. Arif, M. Anwar, and X. Zhou, New whale from the Eocene of Pakistan and the origin of cetacean swimming, Nature, vol.368, pp.844-847, 1994.

R. C. Hulbert, R. M. Petkewich, G. A. Bishop, D. Bukry, and D. P. Aleshire, A new middle Eocene protocetid whale (Mammalia: Cetacea: Archaeoceti) and associated biota from Georgia, Journal of Paleontology, vol.72, issue.5, pp.907-927, 1998.

C. W. Andrews, A description of new species of zeuglodont and of leathery turtle from the Eocene of southern Nigeria, Proceedings of the Zoological Society of London, vol.22, pp.309-319, 1920.

S. Bajpai and J. G. Thewissen, Protocetid cetaceans (Mammalia) from the Eocene of India, Palaeontologia Electronica, vol.17, issue.3, pp.1-19, 2014.

G. Bianucci and P. D. Gingerich, Aegyptocetus tarfa, n. gen. et sp. (Mammalia, Cetacea), from the middle Eocene of Egypt: Clinorhynchy, olfaction, and hearing in a protocetid whale, Journal of Vertebrate Paleontology, vol.31, issue.6, pp.1173-1188, 2011.

E. Fraas, Neue Zeuglodonten aus dem unteren Mitteleocän vom Mokattam bei Cairo. Mittheilungen aus dem Königlichen Naturalien-Cabinet in Stuttgart, vol.27, pp.1-24, 1904.

J. H. Geisler, A. E. Sanders, and Z. X. Luo, A new protocetid whale (Cetacea: Archaeoceti) from the late middle Eocene of South Carolina, American Museum Novitates, vol.3480, pp.1-65, 2005.

P. D. Gingerich, M. Arif, and W. C. Clyde, New archaeocetes (Mammalia, Cetacea) from the middle Eocene Domanda formation of the Sulaiman range, Contributions from the Museum of Paleontology, vol.29, pp.291-330, 1995.

P. D. Gingerich and H. Cappetta, A new Archaeocete and other marine mammals (Cetacea and Sirenia) from lower middle Eocene phosphate deposits of Togo, Journal of Paleontology, vol.88, issue.1, pp.109-129, 2014.

P. D. Gingerich, M. Haq, I. H. Khan, and I. S. Zalmout, Eocene stratigraphy and archaeocete whales (Mammalia, Cetacea) of Drug Lahar in the eastern Sulaiman range, Contributions from the Museum of Paleontology, vol.30, pp.269-319, 2001.

P. D. Gingerich, M. Haq, W. Koenigswald, W. J. Sanders, B. H. Smith et al., New protocetid whale from the middle Eocene of Pakistan: birth on land, precocial development and sexual dimorphism, PloS ONE, vol.4, issue.2, pp.4366-4367, 2009.

P. D. Gingerich, M. Haq, I. S. Zalmout, I. H. Khan, and M. S. Malkani, Origin of whales from early Artiodactyls: hand and feet of Eocene Protocetidae from Pakistan, Science, vol.293, pp.2239-2242, 2001.

P. D. Gingerich, S. M. Raza, M. Arif, M. Anwar, and X. Zhou, New whale from the Eocene of Pakistan and the origin of cetacean swimming, Nature, vol.368, pp.844-847, 1994.

P. D. Gingerich, I. S. Zalmout, M. Haq, and M. A. Bhatti, Makaracetus bidens, a new protocetid archaeocete (Mammalia, Cetacea) from the early middle Eocene of Balochistan (Pakistan), vol.31, pp.197-210, 2005.

L. Hautier, R. Sarr, F. Lihoreau, R. Tabuce, and P. M. Hameh, First record of the family Protocetidae in the Lutetian of Senegal (West Africa), Palaeovertebrata, vol.38, issue.2, pp.1-7, 2014.
URL : https://hal.archives-ouvertes.fr/hal-01923037

R. C. Hulbert, R. M. Petkewich, G. A. Bishop, D. Bukry, and D. P. Aleshire, A new middle Eocene protocetid whale (Mammalia: Cetacea: Archaeoceti) and associated biota from Georgia, Journal of Paleontology, vol.72, issue.5, pp.907-927, 1998.

S. A. Mcleod and L. G. Barnes, A new genus and species of Eocene protocetid archaeocete whale (Mammalia, Cetacea) from the Atlantic Coastal Plain, Natural History Museum of Los Angeles County, vol.41, pp.73-98, 2008.

A. Sahni and V. P. Mishra, Lower tertiary vertebrates from western India, vol.3, pp.1-48, 1975.

A. N. Trivedy and P. P. Satsangi, A new archaeocete (whale) from the Eocene of India. Abstracts of 27 th International Geological Congress, vol.1, pp.322-323, 1984.

M. D. Uhen, New protocetid (Mammalia, Cetacea) from the late middle Eocene Cook Mountain formation of Louisiana, Journal of Vertebrate Paleontology, vol.18, issue.3, pp.664-668, 1998.

S. Figure, 3D models of the casts of the bony labyrinths of ?Carolinacetus sp. (UM-KPG-M 164; A-D) and of the protocetid indeterminate (morphotype ?

E. , dorsal (D, H) views. Related to Figure 1. Scale bar = 5 mm. Abbreviations: aa, anterior ampulla; asc, anterior semicircular canal; br, basal ridge (imprint); ca, cochlear aqueduct; cc, common crus; co, cochlear canal; er, elliptical recess; fc, fenestra cochleae; fv, fenestra vestibuli; la, lateral ampulla; lsc, lateral semicircular canal; pa, posterior ampulla; psc, posterior semicircular canal; scc, secondary common crus, Kpogamé illustrated in medial (A, E), anterior (B, F), lateral (C, G)

S. Data, List of taxa and measurments included in the PCA and in ancestral character state Supplemental References S1, J. Morphol, vol.277, pp.1599-1615, 2016.

G. Fleischer, Hearing in Extinct Cetaceans as Determined by Cochlear Structure, J. Paleontol, vol.50, pp.133-152, 1976.

. S3, E. G. Ekdale, R. , and T. , Morphology and variation within the bony labyrinth of zhelestids (Mammalia, Eutheria) and other therian mammals, J. Vertebr. Paleontol, vol.31, pp.658-675, 2011.

. S4, E. G. Ekdale, R. A. Racicot, . S5, F. Spoor et al., Anatomical evidence for low frequency sensitivity in an archaeocete whale: Comparison of the inner ear of Zygorhiza kochii with that of crown Mysticeti, Am. J. Phys. Anthropol, vol.226, pp.211-251, 1998.

E. G. Ekdale, Ontogenetic Variation in the Bony Labyrinth of Monodelphis domestica (Mammalia: Marsupialia) Following Ossification of the Inner Ear Cavities, Anat. Rec, vol.293, pp.1896-1912, 2010.

M. Churchill, M. Martinez-caceres, C. De-muizon, J. Mnieckowski, and J. H. Geisler, The Origin of High-Frequency Hearing in Whales, Curr. Biol, vol.26, pp.2144-2149, 2016.
URL : https://hal.archives-ouvertes.fr/hal-02612270

. S8, M. J. Orliac, J. Benoit, and M. A. Leary, The inner ear of Diacodexis, the oldest artiodactyl mammal, J. Anat, vol.221, pp.417-426, 2012.

, Literature cited

Z. Luo and P. D. Gingerich, Terrestrial Mesonychia to Aquatic Cetacea: Transformation of the Basicranium and Evolution of Hearing in Whales, Univ. Michigan -Pap. Paleontol, vol.31, pp.1-98, 1999.

. S2 and M. A. O&apos;leary, An anatomical and phylogenetic study of the osteology of the petrosal of extant and extinct artiodactylans (Mammalia) and relatives, Bull. Am. Museum Nat. Hist, vol.335, pp.1-206, 2010.

. S3, M. Martínez-cáceres, O. Lambert, and . Muizon-c-de, The anatomy and phylogenetic affinities of Cynthiacetus peruvianus, a large Dorudon-like basilosaurid (Cetacea, Mammalia) from the late Eocene of Peru, Geodiversitas, vol.39, pp.7-163, 2017.

. S4, F. C. Fraser, and P. E. Purves, Hearing in Cetaceans. Bull. Br. Museum (Natural Hist. Zool, vol.7, pp.1-140, 1960.

. S5 and W. C. Lancaster, The Middle Ear of the Archaeoceti, J. Vertebr. Paleontol, vol.10, pp.117-127, 1990.

. S6 and G. Fleischer, Evolutionary Principles of the Mammalian Middle Ear, Adv. Anatomy, Embryol. Cell Biol, vol.55, 1978.

. S7 and . Pompeckj, Das Ohrskelett von Zeuglodon. Senckenbergiana, vol.3, pp.44-100, 1922.

J. S8.-thewissen and S. T. Hussain, Origin of underwater hearing in whales, Nature, vol.361, pp.444-445, 1993.

. S9.-uhen and . Md, Form, Function, and Anatomy of Dorudon atrox (Mammalia, Cetacea): An Archaeocete from the Middle to Late Eocene of Egypt, Univ. Michigan -Pap. Paleontol, vol.34, pp.1-222, 2004.

. S10, S. Nummela, J. Thewissen, S. Bajpai, S. T. Hussain et al., Eocene evolution of whale hearing, Nature, vol.430, pp.776-778, 2004.

. S11, S. Nummela, J. Thewissen, S. Bajpai, S. T. Hussain et al., Sound transmission in archaic and modern whales: Anatomical adaptations for underwater hearing, Anat. Rec, vol.290, pp.716-733, 2007.

V. Bouetel and . Muizon-c-de, The anatomy and relationships of Piscobalaena nana (Cetacea, Mysticeti), a Cetotheriidae s.s. from the early Pliocene of Peru, Geodiversitas, vol.28, pp.319-395, 2006.
URL : https://hal.archives-ouvertes.fr/hal-00092430

. S13 and O. Henson, Some morphological and functional aspects of certain structures of the middle ear in bats and insectivores, Univ. Kansas Sci. Bull, vol.42, pp.151-249, 1961.

. S14, J. Buytaert, W. Salih, M. Dierick, P. Jacobs et al., Realistic 3D Computer Model of the Gerbil Middle Ear, Featuring Accurate Morphology of Bone and Soft Tissue Structures, J. Assoc. Res. Otolaryngol, vol.12, pp.681-696, 2011.

. S15, J. G. Mead, and R. E. Fordyce, The therian skull : a lexicon with emphasis on the odontocetes, Smithson. Contrib. to Zool. 1-249, 2009.

J. Boisserie, The Evolution of Artiodactyls, pp.106-119, 2007.

A. W. Gentry and J. J. Hooker, The phylogeny and classification of the tetrapods, vol.2, pp.235-272, 1988.

J. E. Gray, On the natural arrangement of vertebrose animals, London Med. Repos, vol.15, pp.297-310, 1821.

F. Lihoreau, L. Alloing-séguier, P. Antoine, J. Boisserie, L. Marivaux et al., Enamel microstructure defines a major Paleogene hippopotamoid clade: the Merycopotamini (Cetartiodactyla, Hippopotamoidea), Hist. Biol, vol.29, pp.947-957, 2017.
URL : https://hal.archives-ouvertes.fr/hal-01813199

F. Lihoreau, J. Boisserie, F. K. Manthi, and S. Ducrocq, Hippos stem from the longest sequence of terrestrial cetartiodactyl evolution in Africa, Nat. Commun, vol.6, 2015.
URL : https://hal.archives-ouvertes.fr/hal-01893828

F. Lihoreau and S. Ducrocq, The Evolution of Artiodactyls, pp.89-105, 2007.

L. Scherler, F. Lihoreau, and D. Becker, To split or not to split Anthracotherium? A phylogeny of Anthracotheriinae (Cetartiodactyla: Hippopotamoidea) and its palaeobiogeographical implications, Zool. J. Linn. Soc, pp.1-24, 2018.
URL : https://hal.archives-ouvertes.fr/hal-02131130

A. H. Sileem, H. M. Sallam, A. G. Hewaidy, E. R. Miller, and G. F. Gunnell, A new anthracothere (Artiodactyla) from the early Oligocene, Fayum, Egypt, and the mystery of African "Rhagatherium" solved, J. Paleontol, vol.90, pp.170-181, 2016.

T. Tsubamoto, Z. Maung-thein, N. Egi, T. Nishimura, T. Htike et al., A new Anthracotheriid Artiodactyl from the Eocene Pondaung Formation of Myanmar, Vertebr. Palasiat, pp.85-113, 2011.

;. Cas, C. Academy, S. Sciences, C. A. Francisco, M. Usa;-ccnhm et al., LACM, Natural History Museum Los Angeles County, vol.IRSNB

S. Sdnhm/sdsnh, . Diego-museum-natural-history, S. Museum, . Diego, . Ca et al., UPPal (IPHEP), Institute of Paleoprimatology, Human Paleontology: Evolution and Paleoenvironments

M. Churchill, M. Martínez-cáceres, C. Muizon, . De, J. Mnieckowski et al., The Origin of High-Frequency Hearing in Whales, Curr. Biol, vol.26, pp.2144-2149, 2016.
URL : https://hal.archives-ouvertes.fr/hal-02612270

E. G. Ekdale, Comparative Anatomy of the Bony Labyrinth (Inner Ear) of Placental Mammals, PLoS One, vol.8, p.100, 2013.

E. G. Ekdale, Morphological variation among the inner ears of extinct and extant baleen whales (Cetacea: Mysticeti), J. Morphol, vol.277, pp.1599-1615, 2016.

E. G. Ekdale and R. A. Racicot, Anatomical evidence for low frequency sensitivity in an archaeocete whale: Comparison of the inner ear of Zygorhiza kochii with that of crown Mysticeti, J. Anat, vol.226, pp.22-39, 2015.

G. Fleischer, Hearing in Extinct Cetaceans as Determined by Cochlear Structure, J. Paleontol, vol.50, pp.133-152, 1976.

P. J. Gannon, A. R. Eden, and J. T. Laitman, The Subarcuate Fossa and Cerebellum o f Extant Primates : Comparative Study of a Skull-Brain Interface, Am. J. Phys. Anthropol, vol.77, pp.143-164, 1988.

J. H. Geisler and Z. Luo, The Petrosal and Inner Ear of Herpetocetus sp (Mammalia Cetacea) and Their Implications for the Phylogeny and Hearing of Archaic Mysticetes, J. Paleontol, vol.70, pp.1045-1066, 1996.

J. H. Geisler, A. E. Sanders, and Z. Luo, A New Protocetid Whale (Cetacea: Archaeoceti) from the Late Middle Eocene of South Carolina, Am. Museum Novit, vol.3480, pp.1-65, 2005.

Z. Luo and E. R. Eastman, Petrosal and Inner Ear of a Squalodontoid Whale : Implications for Evolution of Hearing in Odontocetes, J. Vertebr. Paleontol, vol.15, pp.431-442, 1995.

Z. Luo and P. D. Gingerich, Terrestrial Mesonychia to Aquatic Cetacea: Transformation of the Basicranium and Evolution of Hearing in Whales, Univ. Michigan -Pap. Paleontol, vol.31, pp.1-98, 1999.

R. D. Macphee, Auditory Regions of Primates and Eutherian Insectivores: Morphology, Ontogeny, and Character Analysis, Contrib. Primatol, vol.18, p.282, 1981.

M. J. Mourlam and M. J. Orliac, Infrasonic and Ultrasonic Hearing Evolved after the Emergence of Modern Whales, Curr. Biol, vol.27, pp.1776-1781, 2017.
URL : https://hal.archives-ouvertes.fr/hal-01894431

C. Muizon, . De, G. Billet, C. Argot, S. Ladevèze et al., Alcidedorbignya inopinata, a basal pantodont (Placentalia, Mammalia) from the early Palaeocene of Bolivia: Anatomy, phylogeny and palaeobiology, Geodiversitas, vol.37, pp.397-631, 2015.
URL : https://hal.archives-ouvertes.fr/hal-01455642

M. A. O&apos;leary, An anatomical and phylogenetic study of the osteology of the petrosal of extant and extinct artiodactylans (Mammalia) and relatives, Bull. Am. Museum Nat. Hist, vol.335, pp.1-206, 2010.

M. J. Orliac, R. Araújo, and F. Lihoreau, The petrosal and bony labyrinth of Diplobune minor, an enigmatic Artiodactyla from the Oligocene of Western Europe, J. Morphol, vol.278, pp.1168-1184, 2017.
URL : https://hal.archives-ouvertes.fr/hal-01942305

M. J. Orliac, J. Benoit, and M. A. O&apos;leary, The inner ear of Diacodexis, the oldest artiodactyl mammal, 2012.

, J. Anat, vol.221, pp.417-426

M. J. Orliac and M. A. O&apos;leary, Comparative Anatomy of the Petrosal Bone of Dichobunoids, Early Members of Artiodactylamorpha (Mammalia), J. Mamm. Evol, vol.21, pp.299-320, 2014.
URL : https://hal.archives-ouvertes.fr/hal-01897864

M. J. Orliac and M. A. O&apos;leary, The inner ear of Protungulatum (Pan-Euungulata, Mammalia), J. Mamm. Evol, vol.23, 2016.
URL : https://hal.archives-ouvertes.fr/hal-01925070

R. Presley, Alisphenoid equivalents in placentals, marsupials, monotremes and fossils.pdf, Nature, vol.294, pp.668-670, 1981.

A. Ravel and M. J. Orliac, The inner ear morphology of the 'condylarthran' Hyopsodus lepidus, Hist. Biol, vol.27, pp.957-969, 2015.
URL : https://hal.archives-ouvertes.fr/hal-01904989

S. Sisson, A Text-Book of Veterinary Anatomy, 1910.

J. R. Wible, On the cranial osteology of the short-tailed opossum Monodelphis brevicaudata (Didelphidae, Marsupialia), Ann. Carnegie Museum, vol.72, pp.137-202, 2003.

, BEGIN CHARACTERS

, DIMENSIONS, vol.80

. Format-datatype=standard-missing=?-gap=-symbols=, 012345

, Inflation of the mastoid/tegmen tympani at the external auditory meatus

, Ventrolateral tuberosity'

, Ventrolateral tuberosity -Size'

, Ventrolateral tuberosity -Shape

, Lateral process of epitympanic wing -Width' [11] 'Lateral process of epitympanic wing -Length

, Caudal tympanic process -Wing'

, Caudal tympanic process -Wing size' [18] 'Fenestra cochleae -Orientation of opening

, Stapedial muscle fossa -Size' [28] 'Tegmen tympani -Size' [29] 'Fossa for tensor tympani m. -Excavating the tegmen tympani' [30] 'Fossa for tensor tympani m. -Size' [31] 'Fossa for the head of malleus

, Subarcuate fossa -Anterior semicircular canal contribution

, Internal auditory meatus -Tube' [41] 'Internal auditory meatus -Depth' [42] 'Internal auditory meatus -Size' [43] 'Internal auditory meatus -Size of foramen acusticum superius relative to foramen acusticum inferius

, Mastoid plate'

, Prefacial commissure fossa

, Prefacial commissure fossa -Size'

, Prefacial commissure notch

, Prefacial commissure notch -Size

, Anteromedial tuberosity'

, Anteromedial tuberosity -Size

, Sulcus for capsuloparietal emissary vein

, Sulcus for capsuloparietal emissary vein -Depth, Basicapsular groove

, Basicapsular groove -Location'

, Epitympanic wing''s tip -Orientation

H. Fallopii--size,

, Hiatus Fallopii -Position'

, Crista Petrosa -Size' [66] 'Tegmen tympani fossa

, Lateral exposure of mastoid region

, Dorsal exposure of mastoid region

, Longer semicircular canal

, Shorter semicircular canal

, Secondary common crus

, Secondary common crus -Contribution'

, Branching of the lateral semicircular canal, Bretzel" contact

, Cochleovestibular ratio'

, Secondary bony lamina -Extension'

, Secondary bony lamina -Basis at basal turn'

, Number of turn of cochlear canal

, Basal ratio of cochlea

, Interturn distance ratio'

, Dichobune leporina' 1001001111 00-0---1-1 0300101111 1111411110 00200-0-01 01101010-1 ??11110001 10200?002? 'Homacodon vagans' 1001011010 111110-110 3320101211 1111111010 00000-1201 11111020-1 1111110101 10201?0121 'Helohyus plicodon' 1101011011 210100-100 031011?211 1110001030 1100101001 11101010-1 1110-00101 1020100132 'Gobiohyus, 1100120000 Pleuraspidotherium 1001101??? ?0-????1?1 3310101301 0?11311210 10210-0-01 011111?0-1 0010-0??21 1100120010 'Choeropotamus affinis' 100???1111 110110-1?0 3200102211 ?01111?030 1000??1001 2???11110? 11????2021 0-0101?112 'Haplobunodon, p.1100201123, 2010.

A. Assemat and N. Brualla, The petrosal bone of Prodremotherium sp, 2015.

. Morphomuseum,

M. Churchill, M. Martínez-cáceres, C. Muizon, . De, J. Mnieckowski et al., The Origin of High-Frequency Hearing in Whales, Curr. Biol, vol.26, pp.2144-2149, 2016.
URL : https://hal.archives-ouvertes.fr/hal-02612270

E. G. Ekdale and R. A. Racicot, Anatomical evidence for low frequency sensitivity in an archaeocete whale: Comparison of the inner ear of Zygorhiza kochii with that of crown Mysticeti, J. Anat, vol.226, pp.22-39, 2015.

J. H. Geisler, A. E. Sanders, and Z. Luo, A New Protocetid Whale (Cetacea: Archaeoceti) from the Late Middle Eocene of South Carolina, Am. Museum Novit, vol.3480, pp.1-65, 2005.

D. R. Ketten and D. Wartzok, Three-dimensional reconstructions of the dolphin ear, Sensory Abilities of Cetaceans: Laboratory and Field Evidence, pp.81-105, 1990.

S. Ladevèze, P. Missiaen, and T. Smith, Condylarthra") from the late Paleocene of Berru (France) and phylogenetic affinities of the enigmatic European family Pleuraspidotheriidae, J. Vertebr. Paleontol, vol.30, pp.1559-1578, 2010.

M. J. Mourlam and M. J. Orliac, Infrasonic and Ultrasonic Hearing Evolved after the Emergence of Modern Whales, Curr. Biol, vol.27, pp.1776-1781, 2017.
URL : https://hal.archives-ouvertes.fr/hal-01894431

M. J. Mourlam and M. J. Orliac, Protocetid (Cetacea, Artiodactyla) bullae and petrosals from the middle Eocene locality of Kpogamé, Togo: new insights into the early history of cetacean hearing, J. Syst. Palaeontol, vol.16, pp.621-644, 2018.

C. Muizon, . De, G. Billet, C. Argot, S. Ladevèze et al., Alcidedorbignya inopinata, a basal pantodont (Placentalia, Mammalia) from the early Palaeocene of Bolivia: Anatomy, phylogeny and palaeobiology, Geodiversitas, vol.37, pp.397-631, 2015.
URL : https://hal.archives-ouvertes.fr/hal-01455642

M. A. O&apos;leary, An anatomical and phylogenetic study of the osteology of the petrosal of extant and extinct artiodactylans (Mammalia) and relatives, Bull. Am. Museum Nat. Hist, vol.335, pp.1-206, 2010.

M. J. Orliac, The petrosal bone of extinct Suoidea (Mammalia, Artiodactyla), J. Syst. Palaeontol, vol.11, pp.925-945, 2013.

M. J. Orliac, R. Araújo, and F. Lihoreau, The petrosal and bony labyrinth of Diplobune minor, an enigmatic Artiodactyla from the Oligocene of Western Europe, J. Morphol, vol.278, pp.1168-1184, 2017.
URL : https://hal.archives-ouvertes.fr/hal-01942305

M. J. Orliac and M. A. O&apos;leary, Comparative Anatomy of the Petrosal Bone of Dichobunoids, Early Members of Artiodactylamorpha (Mammalia), J. Mamm. Evol, vol.21, pp.299-320, 2014.
URL : https://hal.archives-ouvertes.fr/hal-01897864

M. J. Orliac and M. A. O&apos;leary, The inner ear of Protungulatum (Pan-Euungulata, Mammalia), J. Mamm. Evol, vol.23, 2016.
URL : https://hal.archives-ouvertes.fr/hal-01925070

W. Segall, Morphological parallelisms of the bulla and auditory ossicles in some insectivores and marsupials, Fieldiana Zool, vol.51, pp.169-205, 1970.

J. G. Thewissen, L. N. Cooper, M. T. Clementz, S. Bajpai, and B. N. Tiwari, Whales originated from aquatic artiodactyls in the Eocene epoch of India, Nature, vol.450, pp.1190-1194, 2007.

, Petrosal and bony labyrinth morphology of Choeropotamidae: phylogenetic insights

M. J. Mourlam and M. J. Orliac, Petrosal and bony labyrinth morphology of Choeropotamidae: phylogenetic insights, 2018.