M. Gaete and A. S. Tucker, Organized Emergence of Multiple-Generations of Teeth in Snakes Is Dysregulated by Activation of Wnt/Beta-Catenin Signalling, PLoS One, vol.8, p.74484, 2013.

G. R. Handrigan and J. M. Richman, A network of Wnt, hedgehog and BMP signaling pathways regulates tooth replacement in snakes, Dev Biol, vol.348, pp.130-141, 2010.

N. Itasaki, C. M. Jones, and S. Mercurio, Wise, a context-dependent activator and inhibitor of Wnt signalling, Development, vol.121, pp.3627-3636, 2003.

E. Järvinen, J. Shimomura-kuroki, and A. Balic, Mesenchymal Wnt/?-catenin signaling limits tooth number, Development, vol.145, 2018.

E. Järvinen, M. Tummers, and I. Thesleff, The role of the dental lamina in mammalian tooth replacement, J Exp Zool Part B Mol Dev Evol, vol.312, issue.4, pp.281-291, 2009.

E. Järvinen, K. Välimäki, and M. Pummila, The taming of the shrew milk teeth, Evol Dev, vol.10, pp.477-486, 2008.

M. Jussila, C. Yanez, X. Thesleff, and I. , Initiation of teeth from the dental lamina in the ferret, Differentiation, vol.87, pp.32-43, 2014.

E. Juuri and A. Balic, The Biology Underlying Abnormalities of Tooth Number in Humans, J Dent Res, vol.96, pp.1248-1256, 2017.

E. Juuri, M. Jussila, and K. Seidel, Sox2 marks epithelial competence to generate teeth in mammals and reptiles, Development, vol.140, pp.1424-1456, 2013.

E. Juuri, K. Saito, and L. Ahtiainen, Sox2+ Stem Cells Contribute to All Epithelial Lineages of the Tooth via Sfrp5+ Progenitors, Dev Cell, vol.23, pp.317-328, 2012.

J. Kim, . Ahn, D. Adasooriya, E. J. Woo, H. J. Kim et al., Shh Plays an Inhibitory Role in Cusp Patterning by Regulation of Sostdc1, J Dent Res, vol.98, pp.98-106, 2018.

J. D. Kormish, D. Sinner, and A. M. Zorn, Interactions between SOX factors and Wnt/betacatenin signaling in development and disease, Dev Dyn, vol.239, pp.56-68, 2010.

Y. Lan, S. Jia, and R. Jiang, Molecular patterning of the mammalian dentition, Semin Cell Dev Biol, pp.61-70, 2014.

M. Lee, E. Kim, K. Otsu, H. Harada, and H. Jung, Sox2 contributes to tooth development via Wnt signaling, Cell Tissue Res, vol.365, issue.1, pp.77-84, 2016.

L. Merametdjian, T. Prud'homme, L. Caignec, and C. , Oro-dental phenotype in patients with RUNX2 duplication, Eur J Med Genet, vol.62, pp.85-89, 2019.
URL : https://hal.archives-ouvertes.fr/inserm-01844233

P. M. Munne, M. Tummers, and E. Järvinen, Tinkering with the inductive mesenchyme: Sostdc1 uncovers the role of dental mesenchyme in limiting tooth induction, Development, vol.136, 2009.

L. Muzio, L. Tetè, S. Mastrangelo, and F. , A novel mutation of gene CBFA1/RUNX2 in cleidocranial dysplasia, Ann Clin Lab Sci, vol.37, pp.115-135, 2007.

E. M. Popa, M. Buchtova, and A. S. Tucker, Revitalising the rudimentary replacement dentition in the mouse, Development, vol.146, p.171363, 2019.

E. M. Popa, N. Anthwal, and A. S. Tucker, Complex patterns of tooth replacement revealed in the fruit bat ( Eidolon helvum ), J Anat, vol.229, pp.847-856, 2016.

L. J. Rasch, K. J. Martin, and R. L. Cooper, An ancient dental gene set governs development and continuous regeneration of teeth in sharks, 2016.

J. M. Richman and G. R. Handrigan, Reptilian tooth development, Genesis, vol.49, pp.247-260, 2011.

Y. Ahn, B. W. Sanderson, O. D. Klein, and R. Krumlauf, Inhibition of Wnt signaling by Wise (Sostdc1) and negative feedback from Shh controls tooth number and patterning, Development, vol.137, pp.3221-3252, 2010.

A. O. Averianov, Homology of the cusps in the molars of the lagomorpha (Mammalia) and certain general problems of homology in the morphological structures, Paleontol. J, vol.32, pp.73-77, 1998.

P. M. Butler, The ontogeny of molar pattern, Biol. Rev, vol.31, pp.30-69, 1956.

B. M. Davis, Evolution of the Tribosphenic Molar Pattern in Early Mammals, with Comments on the "Dual-Origin, Hypothesis. J. Mamm. Evol, vol.18, pp.227-244, 2011.

S. Glasstone, A Comparative Study of the Development in vivo and in vitro of Rat and Rabbit Molars, Proc. R. Soc. London. Ser. B-Biological Sci, vol.126, pp.315-330, 1938.

S. J. Gould, Ontogeny and phylogeny, 1977.

J. P. Hunter and J. Jernvall, The hypocone as a key innovation in mammalian evolution, Proc. Natl. Acad. Sci. U. S. A, vol.92, pp.10718-10740, 1995.

. Nesolagus-(leporidae, Lagomorpha) from Southeast Asia. Science China Earth Sciences, vol.53, issue.8, pp.1134-1140

J. Jernvall, Linking development with generation of novelty in mammalian teeth, Proc. Natl. Acad. Sci, vol.97, pp.2641-2645, 2002.

B. P. Kraatz, D. Badamgarav, and F. Bibi, Gomphos ellae, a new mimotonid from the middle Eocene of Mongolia and its implications for the origin of Lagomorpha, Journal of Vertebrate Paleontology, vol.29, issue.2, pp.576-583, 2009.

B. P. Kraatz, J. Meng, M. Weksler, and C. Li, Evolutionary patterns in the dentition of duplicidentata (Mammalia) and a novel trend in the molarization of premolars, PLoS One, vol.5, pp.1-15, 2010.

M. D. Laubichler, Homology in Development and the Development of the Homology Concept, Am. Zool, vol.40, pp.777-788, 2000.

W. P. Luckett, An Ontogenetic Assessment of Dental Homologies in Therian Mammals, pp.182-204, 1993.

B. D. Metscher, MicroCT for comparative morphology: simple staining methods allow high-contrast 3D imaging of diverse non-mineralized animal tissues, BMC Physiol, vol.9, p.11, 2009.

H. F. Osborn, Evolution of Mammalian Molar Teeth: To and from the Triangular Type Including Collected and Revised Researches Trituberculy and New Sections on the Forms and Homologies of the Molar Teeth in the Different Orders of Mammals, 1907.

. Macmillan,

S. Pantalacci, L. Guéguen, C. Petit, A. Lambert, R. Peterkovà et al., Transcriptomic signatures shaped by cell proportions shed light on comparative developmental biology, Genome Biol, vol.18, p.29, 2017.
URL : https://hal.archives-ouvertes.fr/hal-01943814

C. Petit, Évolution et Développement d'un organe sériel Transcriptomique comparée des bourgeons de molaire chez les rongeurs, Thèse Dr. en Sci. la Vie, 2017.

A. Sadier, M. Twarogowska, K. Steklikova, L. Hayden, A. Lambert et al., Modeling Edar expression reveals the hidden dynamics of tooth signaling center patterning, PLOS Biol, vol.17, p.3000064, 2019.
URL : https://hal.archives-ouvertes.fr/hal-02080524

L. M. Van-valen, Serial homology: the crests and cusps of mammalian teeth, Acta Palaeontol. Pol, vol.38, pp.145-158, 1994.

C. Angelone and L. Rook, Alilepus meini nov. sp. (Leporidae, Lagomorpha) du Messinien inférieur de Toscane (Italie centro-occidentale), Geobios, vol.44, pp.151-156, 2011.

C. Angelone, J. A. Schultz, and M. A. Erbajeva, Determining the ontogenetic variation of lower cheek teeth occlusal surface patterns in lagomorphs using micro-ct technology, Palaeontol. Electron, vol.17, pp.1-13, 2014.

R. J. Asher, J. Meng, J. R. Wible, M. C. Mckenna, G. W. Rougier et al., Stem Lagomorpha and the antiquity of Glires, Science, vol.307, pp.1091-1095, 2005.

A. O. Averianov, A. V. Abramov, and A. N. Tikhonov, A new species of. Nesolagus (Lagomorpha, Leporidae) from Vietnam with osteological description, Contrib. from Zool. Inst, vol.3, 2000.

A. O. Averianov and A. S. Tesakov, Evolutionary trends in Mio-Pliocene Leporinae, based on Trischizolagus, vol.71, pp.145-153, 1997.

. Springer-verlag,

M. Ben-amar and F. Jia, Anisotropic growth shapes intestinal tissues during embryogenesis, Proc. Natl. Acad. Sci. U. S. A, vol.110, pp.10525-10555, 2013.

T. J. Bertin, B. Thivichon-prince, A. R. Leblanc, M. W. Caldwell, and L. Viriot, Current Perspectives on Tooth Implantation, Attachment, and Replacement in Amniota, Front. Physiol, vol.9, p.1630, 2018.

Z. Bosze and L. M. Houdebine, Application of rabbits in biomedical research: a review, World Rabbit Sci, vol.14, pp.1-14, 2006.

M. Carneiro, C. J. Rubin, F. D. Palma, F. W. Albert, J. Alföldi et al.,

G. Barrell, S. Bolet, H. A. Boucher, R. Burbano, J. L. Campos et al.,

M. Rogel-gaillard, S. Ruffier, R. Searle, A. Villafuerte, S. Xiong et al., Rabbit genome analysis reveals a polygenic basis for phenotypic change during domestication, Science, vol.345, pp.1074-1079, 2014.

S. ?ermák, C. Angelone, and M. Sinitsa, New late miocene alilepus (Lagomorpha, Mammalia) from Eastern Europe -a new light on the evolution of the earliest old world leporinae, Bull. Geosci, vol.90, pp.431-451, 2015.

S. ?ermák, The Late Miocene species Ochotona kalfense (Mammalia, Lagomorpha) of Moldova: The oldest European record of the genus in the context of the earliest Ochotoninae, Comptes Rendus Palevol, vol.15, pp.927-940, 2016.

S. ?ermák and J. Wagner, The Pliocene record of Trischizolagus and Pliopentalagus (Leporidae, Lagomorpha, Mammalia) in Central Europe with comments on taxonomy and evolutionary history of Leporinae, Neues Jahrbuch fur Geologie und Paläontologie Abhandlungen, vol.268, pp.97-111, 2013.

J. Chaline, P. Brunet-lecomte, S. Montuire, L. Viriot, and F. Courant, Anatomy of the arvicoline radiation (Rodentia): palaeogeographical, palaeoecological history and evolutionary data, 1999.

L. Chuan-kuei, R. W. Wilson, M. R. Dawson, and L. Krishtalka, The Origin of Rodents and Lagomorphs. Pp. 97-108 in Current Mammalogy, 1987.

V. Conte, J. J. Muñoz, and M. Miodownik, A 3D finite element model of ventral furrow invagination in the Drosophila melanogaster embryo, J. Mech. Behav. Biomed. Mater, vol.1, pp.188-198, 2008.

J. Damuth and C. M. Janis, On the relationship between hypsodonty and feeding ecology in ungulate mammals, and its utility in palaeoecology, Biol. Rev, vol.86, pp.733-758, 2011.

A. Dare, R. Hachisu, A. Yamaguchi, S. Yokose, S. Yoshiki et al., Effects of Ionizing Radiation on Proliferation and Differentiation of Osteoblast-like Cells, J. Dent. Res, vol.76, pp.658-664, 1997.

L. A. Davidson, M. A. Koehl, R. Keller, and G. F. Oster, How do sea urchins invaginate? Using biomechanics to distinguish between mechanisms of primary invagination, Development, vol.121, pp.2005-2023, 1995.

M. R. Dawson, Later Tertiary Leporidae of North America, Univ. Kansas Paleontol. Contrib. Vertebrata, pp.1-75, 1958.

J. Dommergues, B. David, and D. Marchand, Les relations ontogenèsephylogenèse:Applications paléontologiques, Geobios, vol.19, pp.335-356, 1986.

M. Erbajeva, B. Baatarjav, G. Daxner-höck, and L. J. Flynn, Occurrences of Sinolagomys (Lagomorpha) from the Valley of Lakes (Mongolia), Palaeobiodiversity and palaeoenvironments, vol.97, pp.11-24, 2017.

M. A. Erbajeva and S. Zheng, New data on Late Miocene -Pleistocene ochotonids (Ochotonidae, Lagomorpha) from North China, Acta Zool. Cracoviensia, vol.48, pp.93-117, 2005.

A. Forsten, Latest Hipparion Christol, 1832 in Europe. A review of the Pliocene Hipparion crassum Gervais group and other finds, 2002.

L. Fostowicz-frelik, Species distribution and differentiation of Eurasian Hypolagus, vol.10, pp.197-211, 2003.

?. Fostowicz-frelik, Reassessment of Chadrolagus and Litolagus (Lagomorpha ) and a New Genus of North American Eocene Lagomorph from Wyoming, Am. Museum Novit, vol.3773, pp.1-76, 2013.

?. Fostowicz-frelik and A. R. Tabrum, Leporids (Mammalia, Lagomorpha) from the Diamond O Ranch Local Fauna, Latest Middle Eocene of Southwestern Montana, Ann. Carnegie Museum, vol.78, pp.253-271, 2009.

S. J. Gould, Ontogeny and phylogeny, 1977.

E. Hannezo, J. Prost, and J. F. Joanny, Instabilities of monolayered epithelia: Shape and structure of villi and crypts, Phys. Rev. Lett, vol.107, p.78104, 2011.

C. W. Hibbard, The origin of the P3 pattern of Sylvilagus, Caprolagus, Oryctolagus and Lepus, J. Mammal, vol.14, pp.1-15, 1963.

R. S. Hoffmann, A. Smith-;-wilson, D. E. Reeder, and D. M. , Order Lagomorpha, Mammal Species World, pp.185-211, 2005.

D. Huchon, O. Madsen, M. J. Sibbald, K. Ament, M. J. Stanhope et al., Rodent Phylogeny and a Timescale for the Evolution of Glires: Evidence\rfrom an Extensive Taxon Sampling Using Three Nuclear Genes, 2002.

, Mol Biol Evol, vol.19, pp.1053-1065

C. M. Janis and M. Fortelius, On the means whereby mammals achieve increased functional durability of their dentitions, with special reference to limiting factors, Biol. Rev, vol.63, pp.197-230, 1988.

C. Z. Jin, Y. Tomida, Y. Wang, and Y. Q. Zhang, Lagomorpha) from Southeast Asia. Sci. China Earth Sci, vol.53, pp.1134-1140, 2010.

W. J. Kennedy, Thoughts on the evolution and extinction of Cretaceous ammonites, Proc. Geol. Assoc, vol.100, pp.251-279, 1989.

W. Koenigswald and . Von, Diversity of hypsodont teeth in mammalian dentitionsconstruction and classification, Palaeontographica, vol.294, pp.63-94, 2011.

W. Koenigswald, U. Von, S. Anders, J. A. Engels, I. Schultz et al., Tooth Morphology in Fossil and Extant Lagomorpha (Mammalia) Reflects Different Mastication Patterns, 2010.

, J. Mamm. Evol, vol.17, pp.275-299

W. W. Korth and J. Hageman, Lagomorphs (Mammalia) from the Oligocene (Orellan and Whitneyan) Brule Formation, Nebraska. Trans. Nebraska Acad. Sci. Affil. Soc. Pap. Trans. Nebraska Acad. Sci, vol.182, pp.141-152, 1988.

B. P. Kraatz, J. Meng, M. Weksler, and C. Li, Evolutionary patterns in the dentition of duplicidentata (Mammalia) and a novel trend in the molarization of premolars, PLoS One, vol.5, pp.1-15, 2010.

J. O. Kriegs, G. Churakov, J. Jurka, J. Brosius, and J. Schmitz, Evolutionary history of 7SL RNA-derived SINEs in Supraprimates, 2007.

M. Kücken and A. C. Newell, A model for fingerprint formation, Europhys. Lett, vol.68, pp.141-146, 2004.

O. Kullmer, Evolution of African Plio-Pleistocene suids (Artiodactyla: Suidae) based on tooth pattern analysis, Kaupia Darmstädter Beiträg zur Naturgeschichte, vol.9, pp.1-34, 1999.

L. Scala, G. C. , D. A. O'donovan, I. Yeung, J. Darko et al., Radiation-Induced Craniofacial Bone Growth Inhibition: Efficacy of Cytoprotection following a Fractionated Dose Regimen, Plast. Reconstr. Surg, vol.115, pp.1973-1985, 2005.

C. Li, Y. Wang, Z. Zhang, F. Mao, and J. Meng, A new mimotonidan Mina hui (Mammalia, Glires) from the Middle Paleocene of Qianshan, Vertebr. Palasiat, vol.54, pp.121-136, 2016.

C. Li, J. Meng, and Y. Wang, Dawsonolagus antiquus, A Primitive Lagomorph from the Eocene Arshanto Formation, Bull. Carnegie Museum Nat. Hist, vol.39, pp.97-110, 2007.

A. V. Lopatin and A. O. Averianov, Eocene Lagomorpha (Mammalia) of Asia: 2. Strenulagus and Gobiolagus (Strenulagidae), Paleontol. J, vol.40, pp.198-206, 2006.

N. Lopez-martinez, The lagomorph fossil record and the origin of the European rabbit, Lagomorph biology, pp.27-46, 2008.

R. H. Madden, Hypsodonty in Mammals: Evolution, Geomorphology and the Role of Earth Surface Processes, 2015.

, On Fossil and Recent Lagomorpha, Transactions of the Linnean Society of London, vol.7, pp.433-520

C. A. Matthee, B. J. Van-vuuren, D. Bell, and T. J. Robinson, A molecular supermatrix of the rabbits and hares (Leporidae) allows for the identification of five intercontinental exchanges during the Miocene, Syst. Biol, vol.53, pp.433-447, 2004.

H. Medak, M. Weinreb, H. Sicher, J. P. Weinmann, and I. Schour, The Effect of Single Doses of Irradiation Upon the Tissues of the Upper Rat Incisor, J. Dent. Res, vol.31, pp.559-574, 1952.

J. Meng, Chapter 7: Phylogeny and Divergence of Basal Glires, Bull. Am. Museum Nat. Hist, vol.285, pp.93-109, 2004.

A. A. Mieloch and W. M. Suchorska, The concept of radiation-enhanced stem cell differentiation, Radiol Oncol, vol.49, pp.209-216, 2015.

J. Müller, M. Clauss, D. Codron, E. Schulz, J. Hummel et al.,

. Hatt, Growth and wear of incisor and cheek teeth in domestic rabbits ( Oryctolagus cuniculus ) fed diets of different abrasiveness, J. Exp. Zool. Part A Ecol, 2014.

, Genet. Physiol, vol.321, pp.283-298

R. M. Nowak, Walker's mammals of the world, 1999.

M. A. O'leary, J. I. Bloch, J. J. Flynn, T. J. Gaudin, A. Giallombardo et al., Z. S

G. W. Randall, E. J. Rougier, M. T. Sargis, N. B. Silcox, M. Simmons et al.,

M. Velazco, M. Weksler, J. R. Wible, and A. L. Cirranello, The Placental Mammal Ancestor and the Post-K-Pg Radiation of Placentals, Science, vol.339, pp.662-667, 2013.

G. M. Odell, G. Oster, P. Alberch, and B. Burnside, The mechanical basis of morphogenesis: I. Epithelial folding and invagination, Dev. Biol, vol.85, pp.446-462, 1981.

. Macmillan,

O. A. Reig, A proposed unified nomenclature for the enamelled components of the molar teeth of the Cricetidae (Rodentia), J. Zool, vol.181, pp.227-241, 1977.

J. M. Rensberger, Function in the Cheek Tooth Evolution of Some Hypsodont Geomyoid Rodents, J. Paleontol, vol.49, pp.10-22, 1975.

E. Renvoisé and F. Michon, An Evo-Devo perspective on ever-growing teeth in mammals and dental stem cell maintenance, Front. Physiol, vol.5, pp.1-12, 2014.

D. P. Richman, R. M. Stewart, J. W. Hutchinson, and V. S. Caviness, Mechanical model of brain convolutional development, Science, vol.189, pp.18-21, 1975.

K. D. Rose, The beginning of the age of mammals, 2006.

J. M. Sawyer, J. R. Harrell, G. Shemer, J. Sullivan-brown, M. Roh-johnson et al., Apical constriction: a cell shape change that can drive morphogenesis, Developmental biology, vol.341, issue.1, pp.5-19, 2010.

N. Schmidt-kittler, Feeding specializations in rodents, Senckenbergiana lethaea, vol.82, pp.141-152, 2002.

L. Sych, Lagomorpha from the Oligocene of Mongolia, Palaeontogica Polonica, vol.33, pp.183-200, 1975.

M. Tummers and I. Thesleff, Root or crown: a developmental choice orchestrated by the differential regulation of the epithelial stem cell niche in the tooth of two rodent species, Development, vol.130, pp.1049-1057, 2003.

L. Viriot, The use of serial sections to estimate age-related occlusal variability in arvicolid molars, Acta Zoologica Cracoviensia, issue.1, p.39, 1996.

L. Viriot, J. Chaline, A. Schaaf, and E. Le-boulengé, Ontogenetic change of Ondatra zibethicus (Arvicolidae, Rodentia) cheek teeth analyzed by digital image processing. Morphological change in Quaternary mammals of North America, pp.373-391, 1993.

Y. Wang, C. Li, Q. Li, and D. Li, A synopsis of Paleocene stratigraphy and vertebrate paleontology in the Qianshan Basin, p.54, 2016.

Y. Wang, J. Meng, N. Xijun, and L. Chuankui, Major events of Paleogene mammal radiation in China, Geol. J, vol.42, pp.415-430, 2007.

J. A. White, North American Leporinae (Mammalia: Lagomorpha) from late Miocene (Clarendonian) to latest Pliocene (Blancan), Journal of Vertebrate Paleontology, vol.11, issue.1, pp.67-89, 1991.

A. E. Wood, The Mammalian Fauna of the White River Oligocene: Part III, Lagomorpha. Trans. Am. Philos. Soc, vol.28, p.271, 1940.

S. Abtahi, M. Poosti, and N. Saghravanian, Histological evaluation of orthodontic tooth movement following low level laser irradiation in rabbits, Med. Oral Patol. Oral y Cir. Bucal, 2018.

A. K. Al-hamdany, A. R. Al-khatib, and H. I. Sadi, Influence of olive oil on alveolar bone response during orthodontic retention period: rabbit model study, Acta Odontol. Scand, vol.75, pp.413-422, 2017.

Z. H. Ali and R. Mubarak, Histomorphological study of dentine pulp complex of continuously growing teeth in the rabbits, Life Sci. J, vol.9, pp.1554-1564, 2012.

C. Angelone and L. Rook, Alilepus meini nov. sp. (Leporidae, Lagomorpha) du Messinien inférieur de Toscane (Italie centro-occidentale), Geobios, vol.44, pp.151-156, 2011.

R. J. Asher, J. Meng, J. R. Wible, M. C. Mckenna, G. W. Rougier et al., Stem Lagomorpha and the antiquity of Glires, Science, vol.307, pp.1091-1095, 2005.

A. O. Averianov, Homology of the cusps in the molars of the lagomorpha (Mammalia) and certain general problems of homology in the morphological structures, Paleontol. J, vol.32, pp.73-77, 1998.

A. O. Averianov, A. V. Abramov, and A. N. Tikhonov, A new species of. Nesolagus (Lagomorpha, Leporidae) from Vietnam with osteological description, Contrib. from Zool. Inst, vol.3, 2000.

A. O. Averianov and A. S. Tesakov, Evolutionary trends in Mio-Pliocene Leporinae, based on Trischizolagus (Mammalia, Lagomorpha), Palaeontol. Zeitschrift, vol.71, pp.145-153, 1997.

A. Balic and I. Thesleff, Tissue Interactions Regulating Tooth Development and Renewal, Curr. Top. Dev. Biol, vol.115, pp.157-186, 2015.

R. S. Ball, Issues to Consider for Preparing Ferrets as Research Subjects in the Laboratory, ILAR J, vol.47, pp.348-357, 2006.

M. Ben-amar and F. Jia, Anisotropic growth shapes intestinal tissues during embryogenesis, Proc. Natl. Acad. Sci. U. S. A, vol.110, 2013.

L. Bertonnier-brouty, L. Viriot, T. Joly, C. , and C. , 3D reconstructions of dental epithelium during Oryctolagus cuniculus embryonic development related to the publication "Morphological features of tooth development and replacement in the rabbit Oryctolagus cuniculus, MorphoMuseuM, vol.5, p.90, 2019.

B. Biehs, J. K. Hu, .. Strauli, N. B. Sangiorgi, E. Jung et al., BMI1 represses Ink4a/Arf and Hox genes to regulate stem cells in the rodent incisor, Nat. Cell Biol, vol.15, pp.846-852, 2013.

Z. Bosze and L. M. Houdebine, Application of rabbits in biomedical research: a review, World Rabbit Sci, vol.14, pp.1-14, 2006.

P. M. Butler, The ontogeny of molar pattern, Biol. Rev, vol.31, pp.30-69, 1956.

M. Carneiro, C. J. Rubin, F. Palma, A. Di, F. W. Alföldi et al., Rabbit genome analysis reveals a polygenic basis for phenotypic change during domestication, vol.345, pp.1074-1079, 2014.
URL : https://hal.archives-ouvertes.fr/hal-01193854

S. ?ermák, C. Angelone, and M. V. Sinitsa, New late miocene alilepus (Lagomorpha, Mammalia) from Eastern Europe -a new light on the evolution of the earliest old world leporinae, Bull. Geosci, vol.90, pp.431-451, 2015.

J. Chaline, P. Brunet-lecomte, S. Montuire, L. Viriot, and F. Courant, Anatomy of the arvicoline radiation (Rodentia): palaeogeographical, palaeoecological history and evolutionary data, Ann. Zool. Fennici, vol.36, pp.239-267, 1999.

M. G. Chavez, W. Yu, B. Biehs, H. Harada, M. L. Snead et al., Characterization of Dental Epithelial Stem Cells from the Mouse Incisor with Two-Dimensional and Three-Dimensional Platforms, Tissue Eng. Part C Methods, vol.19, p.120816080322006, 2012.

L. Chuan-kuei, R. W. Wilson, M. R. Dawson, and L. Krishtalka, The Origin of Rodents and Lagomorphs, Current Mammalogy, pp.97-108, 1987.

M. T. Cobourne and P. T. Sharpe, Making up the numbers: The molecular control of mammalian dental formula, Semin. Cell Dev. Biol, vol.21, pp.314-324, 2010.

V. Conte, J. J. Muñoz, and M. Miodownik, A 3D finite element model of ventral furrow invagination in the Drosophila melanogaster embryo, J. Mech. Behav. Biomed, 2008.

. Mater, , vol.1, pp.188-198

J. Damuth, J. , and C. M. , On the relationship between hypsodonty and feeding ecology in ungulate mammals, and its utility in palaeoecology, Biol. Rev, vol.86, pp.733-758, 2011.

A. Dare, R. Hachisu, A. Yamaguchi, S. Yokose, S. Yoshiki et al., Effects of Ionizing Radiation on Proliferation and Differentiation of Osteoblast-like Cells, J. Dent. Res, vol.76, pp.658-664, 1997.

B. M. Davis, Evolution of the Tribosphenic Molar Pattern in Early Mammals, with Comments on the "Dual-Origin, Hypothesis. J. Mamm. Evol, vol.18, pp.227-244, 2011.

J. Dommergues, B. David, and D. Marchand, Les relations ontogenèsephylogenèse:Applications paléontologiques, Geobios, vol.19, issue.86, pp.80022-80027, 1986.

H. Dosed?lová, J. Dumková, H. Lesot, K. Glocová, M. Kunová et al., , 2015.

, Fate of the molar dental lamina in the monophyodont mouse, PLoS One, vol.10

Q. Eastman and R. Grosschedl, Regulation of LEF-1/TCF transcription factors by Wnt and other signals, Curr. Opin. Cell Biol, vol.11, pp.233-273, 1999.

M. Erbajeva, B. Baatarjav, G. Daxner-höck, and L. J. Flynn, Occurrences of Sinolagomys (Lagomorpha) from the Valley of Lakes (Mongolia). Palaeobiodiversity and palaeoenvironments 97, pp.11-24, 2017.

A. Forsten, Latest Hipparion Christol, 1832 in Europe. A review of the Pliocene Hipparion crassum Gervais group and other finds, Mammalia, Equidae). Geodiversitas, vol.24, pp.465-486, 2002.

?. Fostowicz-frelik, Reassessment of Chadrolagus and Litolagus (Lagomorpha ) and a New Genus of North American Eocene Lagomorph from Wyoming, Am. Museum Novit, vol.3773, pp.1-76, 2013.

?. Fostowicz-frelik and A. R. Tabrum, Leporids (Mammalia, Lagomorpha) from the Diamond O Ranch Local Fauna, 2009.

, Carnegie Museum, vol.78, pp.253-271

M. Gaete and A. S. Tucker, Organized Emergence of Multiple-Generations of Teeth in Snakes Is Dysregulated by Activation of Wnt/Beta-Catenin Signalling, PLoS One, vol.8, 2013.

S. Glasstone, A Comparative Study of the Development in vivo and in vitro of Rat and Rabbit Molars, Proc. R. Soc. London. Ser. B-Biological Sci, vol.126, pp.315-330, 1938.

S. J. Gould, Ontogeny and phylogeny, 1977.

G. R. Handrigan, K. J. Leung, J. M. Richman, N. Barker, J. H. Es et al., Identification of putative dental epithelial stem cells in a lizard with life-long tooth replacement, Development, vol.137, pp.3545-3554, 2010.

G. R. Handrigan and J. M. Richman, A network of Wnt, hedgehog and BMP signaling pathways regulates tooth replacement in snakes, Dev. Biol, vol.348, pp.130-141, 2010.

G. R. Handrigan and J. M. Richman, Autocrine and paracrine Shh signaling are necessary for tooth morphogenesis, but not tooth replacement in snakes and lizards (Squamata), Dev. Biol, vol.337, pp.171-186, 2010.

E. Hannezo, J. Prost, and J. F. Joanny, Instabilities of monolayered epithelia: Shape and structure of villi and crypts, Phys. Rev. Lett, vol.107, p.78104, 2011.

C. W. Hibbard, The origin of the P3 pattern of Sylvilagus, Caprolagus, Oryctolagus and Lepus, J. Mammal, vol.14, pp.1-15, 1963.

Z. Hirschfeld, M. M. Weinreb, and Y. Michaeli, Incisors of the Rabbit: Morphology, Histology, and Development, J. Dent. Res, vol.52, pp.377-384, 1973.

R. S. Hoffmann, A. Smith, D. E. Wilson, and D. M. Reeder, Order Lagomorpha, Mammal Species World, pp.185-211, 2005.

S. L. Horowitz, S. H. Weisbroth, and S. Scher, Deciduous dentition in the rabbit (Oryctolagus cuniculus). A roentgenographic study, Arch. Oral Biol, vol.18, pp.517-540, 1973.

J. P. Hunter, J. , and J. , The hypocone as a key innovation in mammalian evolution, Proc. Natl. Acad. Sci. U. S. A, vol.92, pp.10718-10740, 1995.

N. Itasaki, C. M. Jones, S. Mercurio, A. Rowe, P. M. Domingos et al., , 2003.

, Wise, a context-dependent activator and inhibitor of Wnt signalling, Development, vol.121, pp.3627-3636

V. Jaks, N. Barker, M. Kasper, J. H. Van-es, H. J. Snippert et al., Lgr5 marks cycling, yet long-lived, hair follicle stem cells, Nat. Genet, vol.40, pp.1291-1299, 2008.

E. Jarvinen, I. Salazar-ciudad, W. Birchmeier, M. M. Taketo, J. Jernvall et al., Continuous tooth generation in mouse is, 2006.

, Wnt/beta-catenin signaling, Proc. Natl. Acad. Sci, vol.103, pp.18627-18632

E. Järvinen, J. Shimomura-kuroki, A. Balic, M. Jussila, and I. Thesleff, , 2018.

, Mesenchymal Wnt/?-catenin signaling limits tooth number, Development, vol.145, p.158048

E. Järvinen, M. Tummers, and I. Thesleff, The role of the dental lamina in mammalian tooth replacement, J. Exp. Zool. Part B Mol. Dev. Evol, vol.312, pp.281-291, 2009.

E. Järvinen, K. Välimäki, M. Pummila, I. Thesleff, J. et al., The taming of the shrew milk teeth, Evol. Dev, vol.10, pp.477-486, 2008.

J. Jernvall, Linking development with generation of novelty in mammalian teeth, Proc. Natl. Acad. Sci. 97, pp.2641-2645, 2002.

J. Jernvall and I. Thesleff, Tooth shape formation and tooth renewal: evolving with the same signals, Development, vol.139, pp.3487-3497, 2012.

C. Z. Jin, Y. Tomida, Y. Wang, and Y. Q. Zhang, First discovery of fossil Nesolagus (Leporidae, Lagomorpha) from Southeast Asia, Sci. China Earth Sci, vol.53, pp.1134-1140, 2010.

M. Jussila, X. Crespo-yanez, and I. Thesleff, Initiation of teeth from the dental lamina in the ferret, Differentiation, vol.87, pp.32-43, 2014.

E. Juuri and A. Balic, The Biology Underlying Abnormalities of Tooth Number in Humans, J. Dent. Res, vol.96, pp.1248-1256, 2017.

E. Juuri, M. Jussila, K. Seidel, S. Holmes, P. Wu et al., Sox2 marks epithelial competence to generate teeth in mammals and reptiles, Development, vol.140, pp.1424-1456, 2013.

E. Juuri, K. Saito, L. Ahtiainen, K. Seidel, M. Tummers et al., , 2012.

, Sox2+ Stem Cells Contribute to All Epithelial Lineages of the Tooth via Sfrp5+ Progenitors, Dev. Cell, vol.23, pp.317-328

W. J. Kennedy, Thoughts on the evolution and extinction of Cretaceous ammonites, Proc. Geol. Assoc, vol.100, pp.80047-80048, 1989.

J. Kim, . Ahn, D. Adasooriya, E. J. Woo, H. J. Kim et al., Shh Plays an Inhibitory Role in Cusp Patterning by Regulation of Sostdc1, J. Dent. Res, vol.98, pp.98-106, 2018.

E. King, Oral sequelae and rehabilitation considerations for survivors of childhood cancer, Br. Dent. J, vol.226, pp.323-329, 2019.

O. D. Klein, G. Minowada, R. Peterkova, A. Kangas, B. D. Yu et al., , 2006.

, Sprouty Genes Control Diastema Tooth Development via Bidirectional Antagonism of Epithelial-Mesenchymal FGF Signaling. Dev. Cell, vol.11, pp.181-190

W. Koenigswald and . Von, Diversity of hypsodont teeth in mammalian dentitionsconstruction and classification, Palaeontographica, vol.294, pp.63-94, 2011.

W. Koenigswald, U. Anders, S. Engels, J. A. Schultz, and I. Ruf, Tooth Morphology in Fossil and Extant Lagomorpha (Mammalia) Reflects Different Mastication Patterns, J. Mamm. Evol, vol.17, pp.275-299, 2010.

J. D. Kormish, D. Sinner, and A. M. Zorn, Interactions between SOX factors and Wnt/beta-catenin signaling in development and disease, Dev. Dyn, vol.239, pp.56-68, 2010.

B. P. Kraatz, J. Meng, M. Weksler, L. , and C. , Evolutionary patterns in the dentition of duplicidentata (Mammalia) and a novel trend in the molarization of premolars, PLoS One, vol.5, pp.1-15, 2010.

J. O. Kriegs, G. Churakov, J. Jurka, J. Brosius, and J. Schmitz, Evolutionary history of 7SL RNA-derived SINEs in Supraprimates, Trends Genet, vol.23, pp.158-161, 2007.

M. Kücken and A. C. Newell, A model for fingerprint formation, Europhys. Lett, vol.68, pp.141-146, 2004.

G. S. Kumar, Orban's Oral Histology &amp, 2014.

. Embryology,

L. Scala, G. C. Odonovan, D. A. Yeung, I. Darko, J. Addison et al., Radiation-Induced Craniofacial Bone Growth Inhibition: Efficacy of Cytoprotection following a Fractionated Dose Regimen, Plast. Reconstr. Surg, vol.115, 1973.

Y. Lan, S. Jia, and R. Jiang, Molecular patterning of the mammalian dentition, Semin. Cell Dev. Biol, pp.61-70, 2014.

M. D. Laubichler, Homology in Development and the Development of the Homology Concept, Am. Zool, vol.40, pp.777-788, 2000.

C. Li, Y. Wang, Z. Zhang, F. Mao, and J. Meng, A new mimotonidan Mina hui (Mammalia, Glires) from the Middle Paleocene of Qianshan, Vertebr. Palasiat, vol.54, pp.121-136, 2016.

C. Li, J. Meng, W. , and Y. , Dawsonolagus antiquus, A Primitive Lagomorph from the Eocene Arshanto Formation, Bull. Carnegie Museum Nat. Hist, vol.39, pp.97-110, 2007.

L. Lo-muzio, S. Tetè, F. Mastrangelo, A. P. Cazzolla, M. G. Lacaita et al., A novel mutation of gene CBFA1/RUNX2 in cleidocranial dysplasia, Ann. Clin. Lab. Sci, vol.37, pp.115-135, 2007.

W. P. Luckett, An Ontogenetic Assessment of Dental Homologies in Therian Mammals, Mammal Phylogeny, pp.182-204, 1993.

Z. Luo, Z. Kielan-jaworowska, and R. L. Cifelli, Evolution of dental replacement in mammals, Bull. Carnegie Museum Nat. Hist, vol.36, p.36, 2004.

C. I. Major, IX. On Fossil and Recent Lagomorpha, 1899.

C. A. Matthee, B. J. Van-vuuren, D. Bell, R. , and T. J. , A molecular supermatrix of the rabbits and hares (Leporidae) allows for the identification of five intercontinental exchanges during the Miocene, Syst. Biol, vol.53, pp.433-447, 2004.

H. Medak, M. Weinreb, H. Sicher, J. P. Weinmann, and I. Schour, The Effect of Single Doses of Irradiation Upon the Tissues of the Upper Rat Incisor, J. Dent. Res, vol.31, pp.559-574, 1952.

J. Meng, Chapter 7: Phylogeny and Divergence of Basal Glires, Bull. Am. Museum Nat. Hist, vol.285, pp.93-109, 2004.

L. Merametdjian, T. Prud'homme, C. Le-caignec, B. Isidor, and S. Lopez-cazaux, , 2019.

, Oro-dental phenotype in patients with RUNX2 duplication, Eur. J. Med. Genet, vol.62, pp.85-89

A. Meredith, Rabbit dentistry, European Journal of Companion Animal Practice, pp.55-62, 2007.

B. D. Metscher, MicroCT for comparative morphology: simple staining methods allow high-contrast 3D imaging of diverse non-mineralized animal tissues, BMC Physiol, vol.9, p.11, 2009.

Y. Michaeli, Z. Hirschfeld, and M. M. Weinreb, The cheek teeth of the rabbit: morphology, histology and development, Cells Tissues Organs, vol.106, pp.223-239, 1980.

A. A. Mieloch and W. M. Suchorska, The concept of radiation-enhanced stem cell differentiation, Radiol Oncol, vol.49, pp.209-216, 2015.

K. D. Moses, W. T. Butler, and C. Qin, Immunohistochemical study of small integrinbinding ligand, N-linked glycoproteins in reactionary dentin of rat molars at different ages, Eur. J. Oral Sci, vol.114, pp.216-238, 2006.

M. L. Mucchielli and T. A. Mitsiadis, Correlation of asymmetric Notch2 expression and mouse incisor rotation, Mech. Dev, vol.91, issue.99, pp.293-295, 2000.

J. Müller, M. Clauss, D. Codron, E. Schulz, J. Hummel et al., Growth and wear of incisor and cheek teeth in domestic rabbits ( Oryctolagus cuniculus ) fed diets of different abrasiveness, J. Exp. Zool. Part A Ecol. Genet. Physiol, vol.321, pp.283-298, 2014.

P. M. Munne, M. Tummers, E. Järvinen, I. Thesleff, J. et al., Tinkering with the inductive mesenchyme: Sostdc1 uncovers the role of dental mesenchyme in limiting tooth induction, Development, vol.136, pp.393-402, 2009.

J. A. Navarro, D. Sottovia-filho, M. C. Leite-ribeiro, and R. Taga, Histological Study on the Postnatal Development and Sequence of Eruption of the Maxillary Cheek-Teeth of Rabbits (Oryctolagus cuniculus), Arch. Histol. Jpn, vol.38, pp.17-30, 1975.

J. A. Navarro, D. Sottovia-filho, M. C. Leite-ribeiro, and R. Taga, Histological study on the postnatal development and sequence of eruption of the mandibular cheekteeth of rabbits (Oryctolagus cuniculus), Arch. Histol. Jpn. Nippon soshikigaku kiroku, vol.39, pp.23-32, 1976.

R. M. Nowak, M. A. Leary, J. I. Bloch, J. J. Flynn, T. J. Gaudin et al., Walker's mammals of the world, The Placental Mammal Ancestor and the Post-K-Pg Radiation of Placentals, 1999.

, , vol.339, pp.662-667

G. M. Odell, G. Oster, P. Alberch, and B. Burnside, The mechanical basis of morphogenesis: I. Epithelial folding and invagination, Dev. Biol, vol.85, pp.90276-90277, 1981.

T. Ooë, Développement embryonnaire des incisives chez le lapin (Oryctolagus cuniculus L.). Interpretation de la formule dentaire, Mammalia, vol.44, pp.259-270, 1980.

. Macmillan,

S. Pantalacci, L. Guéguen, C. Petit, A. Lambert, R. Peterkovà et al., , 2017.

, Transcriptomic signatures shaped by cell proportions shed light on comparative developmental biology, Genome Biol, vol.18, p.29

C. Petit, Évolution et Développement d'un organe sériel Transcriptomique comparée des bourgeons de molaire chez les rongeurs, Thèse Dr. en Sci. la Vie. Available at, 2017.

S. Pillet, N. Svitek, and V. Messling, Ferrets as a Model for Morbillivirus Pathogenesis, Complications, and Vaccines, pp.73-87, 2009.
URL : https://hal.archives-ouvertes.fr/pasteur-00819655

J. Pispa and I. Thesleff, Mechanisms of ectodermal organogenesis, Dev. Biol, vol.262, pp.195-205, 2003.

E. M. Popa, N. Anthwal, and A. S. Tucker, Complex patterns of tooth replacement revealed in the fruit bat ( Eidolon helvum ), J. Anat, vol.229, pp.847-856, 2016.

E. M. Popa, M. Buchtova, and A. S. Tucker, Revitalising the rudimentary replacement dentition in the mouse, Development, vol.146, p.171363, 2019.

L. J. Rasch, K. J. Martin, R. L. Cooper, B. D. Metscher, C. J. Underwood et al., An ancient dental gene set governs development and continuous regeneration of teeth in sharks, 2016.

O. A. Reig, A proposed unified nomenclature for the enamelled components of the molar teeth of the Cricetidae (Rodentia), J. Zool, vol.181, pp.227-241, 1977.

J. M. Rensberger, Function in the Cheek Tooth Evolution of Some Hypsodont Geomyoid Rodents, J. Paleontol, vol.49, pp.10-22, 1975.

E. Renvoisé and F. Michon, An Evo-Devo perspective on ever-growing teeth in mammals and dental stem cell maintenance, Front. Physiol, vol.5, pp.1-12, 2014.

D. P. Richman, R. M. Stewart, J. W. Hutchinson, and V. S. Caviness, Mechanical model of brain convolutional development, Science, vol.189, pp.18-21, 1975.

J. M. Richman and G. R. Handrigan, Reptilian tooth development, Genesis, vol.49, pp.247-260, 2011.

K. D. Rose, M. Twarogowska, K. Steklikova, L. Hayden, A. Lambert et al., Modeling Edar expression reveals the hidden dynamics of tooth signaling center patterning, PLOS Biol, vol.17, p.3000064, 2006.

M. Sanz-navarro, K. Seidel, Z. Sun, L. Bertonnier-brouty, B. A. Amendt et al., Plasticity within the niche ensures the maintenance of a Sox2+ stem cell population in the mouse incisor, Development, vol.145, p.155929, 2018.

N. Schmidt-kittler, Feeding specializations in rodents, Senckenbergiana lethaea, vol.82, pp.141-152, 2002.

M. Seppala, G. J. Fraser, A. A. Birjandi, G. M. Xavier, M. T. Cobourne et al., Sonic Hedgehog Signaling and Development of the Dentition, J. Dev. Biol, vol.5, 2017.

A. Sharir and O. D. Klein, Watching a deep dive: Live imaging provides lessons about tooth invagination, J. Cell Biol, vol.214, pp.645-652, 2016.

Z. Sun, W. Yu, M. S. Navarro, M. Sweat, S. Eliason et al., Sox2 and Lef-1 interact with Pitx2 to regulate incisor development and stem cell renewal, 2016.

, Development

L. S. Sych and P. C. Reade, Heterochrony in the development of vestigial and functional deciduous incisors in rabbits (Oryctolagus cuniculus L.), J. Craniofac. Genet. Dev. Biol, vol.7, pp.81-94, 1987.

I. Thesleff, How to build a tooth?, Developmental biology is revealing the instructions. Nor Tann. Tid, p.327, 2004.

Y. Togo, K. Takahashi, K. Saito, H. Kiso, H. Tsukamoto et al., , 2016.

, Antagonistic Functions of USAG-1 and RUNX2 during Tooth Development, PLoS One, vol.11

A. S. Tucker and G. J. Fraser, Evolution and developmental diversity of tooth regeneration, Semin. Cell Dev. Biol, pp.25-26, 2014.

T. Tumbar, G. Guasch, V. Greco, C. Blanpain, W. E. Lowry et al., Defining the Epithelial Stem Cell Niche in Skin, Science, vol.303, pp.359-363, 2004.

A. Vaahtokari, T. Aberg, J. Jernvall, S. Keränen, and I. Thesleff, The enamel knot as a signaling center in the developing mouse tooth, Mech. Dev, vol.54, pp.39-43, 1996.

L. M. Van-valen, Serial homology: the crests and cusps of mammalian teeth, Acta Palaeontol. Pol, vol.38, pp.145-158, 1994.

L. Viriot, R. Peterková, M. Peterka, and H. Lesot, Evolutionary Implications of the Occurrence of Two Vestigial Tooth Germs During Early Odontogenesis in the Mouse Lower Jaw, Connect. Tissue Res, vol.43, pp.129-133, 2002.

F. Wang, G. Li, Z. Wu, Z. Fan, M. Yang et al., Tracking diphyodont development in miniature pig in vitro and in vivo, Biol. Open, 2019.

F. Wang, J. Xiao, W. Cong, A. Li, T. Song et al., Morphology and chronology of diphyodont dentition in miniature pigs, Sus Scrofa, Oral Dis, vol.20, pp.367-379, 2014.

F. Wang, J. Xiao, W. Cong, A. Li, F. Wei et al., Stage-specific differential gene expression profiling and functional network analysis during morphogenesis of diphyodont dentition in miniature pigs, Sus Scrofa, BMC Genomics, vol.15, p.103, 2014.

X. Wang, T. Aberg, M. J. James, D. Levanon, Y. Groner et al., Runx2 (Cbfa1) Inhibits Shh Signaling in the Lower but not Upper Molars of Mouse Embryos and Prevents the Budding of Putative Successional Teeth, J. Dent. Res, vol.84, pp.138-143, 2005.

Y. Wang, C. Li, Q. Li, and D. Li, A synopsis of Paleocene stratigraphy and vertebrate paleontology in the Qianshan Basin, p.54, 2016.

Y. Wang, J. Meng, N. Xijun, and L. Chuankui, Major events of Paleogene mammal radiation in China, Geol. J, vol.42, pp.415-430, 2007.

J. A. Whitlock and J. M. Richman, Biology of tooth replacement in amniotes, Int. J. Oral Sci, vol.5, pp.66-70, 2013.

A. E. Wood, The Mammalian Fauna of the White River Oligocene: Part III, 1940.

, Lagomorpha. Trans. Am. Philos. Soc, vol.28, p.271

L. K. Xie, K. Wangrangsimakul, S. Suttapreyasri, L. K. Cheung, and T. Nuntanaranont, A preliminary study of the effect of low intensity pulsed ultrasound on new bone formation during mandibular distraction osteogenesis in rabbits, Int. J. Oral Maxillofac, 2011.

. Surg, , vol.40, pp.730-736

Q. Yan, Q. Zhang, H. Yang, Q. Zou, C. Tang et al., Generation of multigene knockout rabbits using the Cas9/gRNA system, Cell Regen, vol.3, p.12, 2014.

M. Yardin, Evolution des dents du jeune lapin, Mammalia, vol.32, pp.677-689, 1968.

M. Zeichner-david, K. Oishi, Z. Su, V. Zakartchenko, L. Chen et al., Role of Hertwig's epithelial root sheath cells in tooth root development, Dev. Dyn, vol.228, pp.651-663, 2003.