I. Afán, J. Navarro, L. Cardador, F. Ramírez, A. Kato et al., Foraging movements and habitat niche of two closely-related seabirds breeding in sympatry, Marine Biology, vol.161, pp.657-668, 2014.

L. Amo, J. M. Avilés, D. Parejo, A. Peña, J. Rodríguez et al., Sex recognition by odour and variation in the uropygial gland secretion in starlings, Journal of Animal Ecology, vol.81, pp.605-613, 2012.

E. C. Anderson and E. A. Thompson, A model-based method for identifying species hybrids using multilocus genetic data, Genetics, vol.160, pp.1217-1229, 2002.

M. J. Anderson, A new method for non-parametric multivariate analysis of variance, Austral Ecology, vol.26, pp.32-46, 2001.

M. J. Anderson and T. J. Willis, Canonical analysis of principal coordinates: a useful method of constrained ordination for ecology, Ecology, vol.84, pp.511-525, 2003.

M. J. Anderson, R. N. Gorley, and K. R. Clarke, PERMANOVA+ for PRIMER: guide to software and statistical methods, 2008.

M. J. Anderson and R. J. , Generalized discriminant analysis based on distances, Australian & New Zealand Journal of Statistics, vol.45, pp.301-318, 2003.

M. C. Baker and A. Baker, Reproductive behavior of female buntings : isolating mechanisms in a hybridizing pair of species, Evolution, vol.44, pp.332-338, 1990.

K. Belkhir, P. Borsa, L. Chikhi, N. Raufaste, and F. Bonhomme, GENETIX 4.05, logiciel sous Windows TM pour la génétique des populations. Laboratoire génome, populations, interactions, vol.5000, pp.1996-2004, 1996.

, The BirdLife checklist of the birds of the world: Version 8, 2015.

F. Bonadonna and G. A. Nevitt, Partner-specific odor recognition in an Antarctic seabird, Science, vol.306, p.835, 2004.

F. Bonadonna, M. E. Grosbois, V. Jouventin, P. Bessiere, and J. M. , Individual odor recognition in birds: an endogenous olfactory signature on petrels' feathers?, Journal of Chemical Ecology, vol.33, pp.1819-1829, 2007.

F. Bonadonna and J. Mardon, One house two families: petrel squatters get a sniff of low-cost breeding opportunities, Ethology, vol.116, pp.176-182, 2010.

F. Bonadonna and A. Sanz-aguilar, Kin recognition and inbreeding avoidance in wild birds: the first evidence for individual kin-related odour recognition, Animal Behaviour, vol.84, pp.509-513, 2012.

F. Bonadonna and J. Mardon, Besides colours and songs, odour is the new black of avian communication, Chemical Signals in Vertebrates, vol.12, pp.325-339, 2013.

V. Bretagnolle and B. Lequette, Structural variation in the call of the Cory's shearwater (Calonectris diomedea, Aves, Procellariidae), Ethology, vol.85, pp.313-323, 1990.

J. Bried, M. P. Dubois, P. Jouventin, and R. S. Santos, Eleven polymorphic microsatellite markers in Cory's shearwater, Calonectris diomedea, and cross-species amplification on threatened Procellariiformes, Molecular Ecology Resources, vol.8, pp.602-604, 2008.

J. Bried, M. P. Dubois, P. Jarne, P. Jouventin, and R. S. Santos, Does competition for nests affect genetic monogamy in Cory's shearwater Calonectris diomedea, Journal of Avian Biology, vol.41, pp.407-418, 2010.

B. Mde and L. , Determination of the absolute visual threshold of a nocturnal seabird, the Common Diving Petrel Pelecanoides urinatrix, Ibis, vol.131, pp.290-300, 1989.

U. Candolin, The use of multiple cues in mate choice, Biological Reviews of the Cambridge Philosophical Society, vol.78, pp.575-595, 2003.

, CALONECTRIS SHEARWATERS 725

, make-up in a seabird species, Proceedings of the Royal Society B: Biological Sciences, vol.181, pp.1185-1193, 2017.

J. Mardon and F. Bonadonna, Atypical homing or self-odour avoidance? Blue petrels (Halobaena caerulea) are attracted to their mate's odour but avoid their own, Behavioral Ecology and Sociobiology, vol.63, pp.537-542, 2009.

J. Mardon, S. M. Saunders, M. J. Anderson, C. Couchoux, and F. Bonadonna, Species, gender, and identity: cracking petrels' sociochemical code, Chemical Senses, vol.35, pp.309-321, 2010.

J. Mardon, S. M. Saunders, and F. Bonadonna, From preen secretions to plumage: the chemical trajectory of blue petrels' Halobaena caerulea social scent, Journal of Avian Biology, vol.42, pp.29-38, 2011.

G. R. Martin, B. Mde, and L. , The eye of a procellariiform seabird, the Manx shearwater, Puffinus puffinus: visual fields and optical structure, vol.37, pp.65-78, 1991.

J. Martín and P. López, Pheromones and chemical communication in lizards, The reproductive biology and phylogeny of Lizards and Tuatara, pp.43-77, 2014.

A. Martínez-abraín, A. Sanchez, and D. Oro, Atlantic Cory's Shearwaters breeding in a colony of Mediterranean Cory's Shearwaters, Waterbirds, vol.25, p.221, 2002.

B. H. Mcardle and M. J. Anderson, Fitting multivariate models to community data: a comment on distance-based redundancy analysis, Ecology, vol.82, pp.290-297, 2001.

F. Mougeot and V. V. Bretagnolle, Predation as a cost of sexual communication in nocturnal seabirds: an experimental approach using acoustic signals, Animal Behaviour, vol.60, pp.647-656, 2000.
URL : https://hal.archives-ouvertes.fr/hal-00193498

J. L. Mougin, Pairing in the Cory's Shearwater (Calonectris diomedea) of Selvagem Grande, Journal für Ornithologie, vol.141, pp.319-326, 2000.

I. Munilla, M. Genovart, V. H. Paiva, and A. Velando, Colony foundation in an Oceanic Seabird, PLoS One, vol.11, p.147222, 2016.

C. P. Nava, S. Y. Kim, M. C. Magalhães, and V. Neves, Do Cory's Shearwaters Calonectris borealis choose mates based on size, Journal of Ornithology, vol.155, pp.869-875, 2014.

J. Navarro, M. G. Forero, J. González-solís, J. M. Igual, J. Bécares et al., Foraging segregation between two closely related shearwaters breeding in sympatry, Biology Letters, vol.5, pp.545-548, 2009.

E. E. Nielsen, L. A. Bach, and P. Kotlicki, HYBRIDLAB (version 1.0): a program for generating simulated hybrids from population samples, Molecular Ecology Notes, vol.6, pp.971-973, 2006.

T. M. Panhuis, R. Butlin, M. Zuk, and T. Tregenza, Sexual selection and speciation, Trends in Ecology & Evolution, vol.16, pp.364-371, 2001.

T. D. Price, Speciation in birds, 2008.

A. C. Alberts, Constraints on the design of chemical communication systems in terrestrial vertebrates, The American Naturalist, vol.139, pp.62-69, 1992.

D. Allainé, Sociality, mating system and reproductive skew in marmots: evidence and hypotheses, Behavioural Processes, vol.51, pp.21-34, 2000.

D. Allainé, F. Brondex, L. Graziani, J. Coulon, and I. Till-bottraud, Male-biased sex ratio in litters of Alpine marmots supports the helper repayment hypothesis, Behavioral Ecology, vol.11, pp.507-514, 2000.

M. J. Anderson, A new method for non-parametric multivariate analysis of variance, Austral Ecology, vol.26, pp.32-46, 2001.

M. J. Anderson, Permutational multivariate analysis of variance (PERMANOVA), Wiley statsref: statistics reference online, pp.1-15, 2017.

K. B. Armitage, Marmot biology: sociality, individual fitness, and population dynamics, 2014.

W. Arnold, The evolution of marmot sociality: I. Why disperse late?, Behavioral ecology and Sociobiology, vol.27, pp.229-237, 1990.

W. Arnold and J. Dittami, Reproductive suppression in male Alpine marmots, Animal Behaviour, vol.53, pp.53-66, 1997.

M. Bell, H. J. Nichols, J. S. Gilchrist, M. A. Cant, and S. J. Hodge, The cost of dominance: suppressing subordinate reproduction affects the reproductive success of dominant female banded mongooses, Proceedings of the Royal Society B: Biological Sciences, vol.279, pp.619-624, 2012.

M. C. Bel, J. Coulon, L. Sreng, D. Allainé, A. G. Bagnères et al., Social signals involved in scent-marking behavior by cheek-rubbing in Alpine marmots (Marmota marmota), Journal of Chemical Ecology, vol.25, pp.2267-2293, 1999.
URL : https://hal.archives-ouvertes.fr/halsde-00295519

M. C. Bel, C. Porteret, and J. Coulon, Scent deposition by cheek rubbing in the alpine marmot (Marmota marmota) in the French Alps, Canadian Journal of Zoology, vol.73, pp.2065-2071, 1995.

A. Bretman, J. D. Westmancoat, M. Gage, and T. Chapan, Males use multiple, redundant cues to detect mating rivals, Current Biology, vol.21, pp.617-622, 2011.

R. E. Brown, Mammalian social odors: a critical review, Advances in Study of Behavior, vol.10, pp.103-162, 1979.

C. D. Buesching, J. S. Waterhouse, and D. W. Macdonald, Gaschromatographic analyses of the subcaudal gland secretion of the European badger (Meles meles) part II: time-related variation in the individual-specific composition, Journal of Chemical Ecology, vol.28, pp.57-69, 2002.

N. Burgener, M. Dehnhard, H. Hofer, and M. L. East, Does anal gland scent signal identity in the spotted hyaena?, Animal Behaviour, vol.77, pp.707-715, 2009.

U. Candolin, The use of multiple cues in mate choice, Biological Reviews of the Cambridge Philosophical Society, vol.78, pp.575-595, 2003.

M. A. Cant, S. J. Hodge, M. Bell, J. S. Gilchrist, and H. J. Nicols, Reproductive control via eviction (but not the threat of eviction) in banded mongooses, Proceedings of the Royal Society B: Biological Sciences, vol.277, pp.2219-2226, 2010.

M. A. Cant and R. A. Johnstone, How threats influence the evolutionary resolution of within-group conflict, The American Naturalist, vol.173, pp.759-771, 2009.

B. A. Casper, F. C. Schroeder, S. Franke, and C. C. Voigt, Scents of adolescence: the maturation of the olfactory, 2011.

T. Zidat and . Al, The Linnean Society of London, phenotype in a free-ranging mammal (T Burne, vol.125, p.21162, 2018.

K. R. Clarke and R. M. Warwick, Change in marine communities: an approach to statistical analysis and interpretation, 2001.

T. Clutton-brock, Cooperation between non-kin in animal societies, Nature, vol.462, pp.51-57, 2009.

T. H. Clutton-brock, P. Brotherton, R. Smith, G. M. Mcilrath, R. Kansky et al., Infanticide and expulsion of females in a cooperative mammal, Proceedings of the Royal Society B: Biological Sciences, vol.265, pp.2291-2295, 1998.

T. H. Clutton-brock, S. J. Hodge, G. Spong, A. F. Russell, N. R. Jordan et al., Intrasexual competition and sexual selection in cooperative mammals, Nature, vol.444, pp.21-28, 2006.

T. H. Clutton-brock and E. Huchard, Social competition and selection in males and females, Philosophical Transactions of the Royal Society of London. Series B, Biological Sciences, vol.368, 2013.

A. Cohas, N. G. Yoccoz, D. Silva, A. Goossens, B. Allainé et al., Night and day: the comparative study of strepsirrhine primates reveals socioecological and phylogenetic patterns in olfactory signals, Philosophical Transactions of the Royal Society of London. Series B, Biological Sciences, vol.59, pp.1909-1922, 2006.

F. S. Dobson, W. F. Van-dongen, J. White, H. B. Brandl, Y. Moodley et al., Competition for mates and predominant juvenile male dispersal in mammals, Animal Behaviour, vol.30, p.11, 1982.

S. Dray and A. B. Dufour, The ade4 package: implementing the duality diagram for ecologists, Journal of Statistical Software, vol.22, pp.1-20, 2007.
URL : https://hal.archives-ouvertes.fr/hal-00434575

C. M. Drea, M. Boulet, J. Delbarco-trillo, L. K. Greene, C. R. Sacha et al., The "secret" in secretions: methodological considerations in deciphering primate olfactory communication, American Journal of Primatology, vol.75, pp.621-642, 2013.

C. M. Drea, S. N. Vignieri, H. S. Kim, M. L. Weldele, and S. E. Glickman, Responses to olfactory stimuli in spotted hyenas (Crocuta crocuta): II. Discrimination of conspecific scent, Journal of Comparative Psychology, vol.116, pp.342-349, 1983.

I. Garin, A. Aldezabal, J. Herrero, A. García-serrano, and J. L. Remón, Diet selection of the Alpine marmot (Marmota m. marmota L.) in the Pyrenees, vol.63, pp.383-390, 2008.

B. Goossens, L. Graziani, L. P. Waits, F. E. Magnolon, S. Coulon et al., Extrapair paternity in the monogamous Alpine marmot revealed by nuclear DNA microsatellite analysis, Behavioral Ecology and Sociobiology, vol.43, pp.281-288, 1998.
URL : https://hal.archives-ouvertes.fr/halsde-00281632

L. M. Gosling and S. C. Roberts, Scent-marking by male mammals: cheat-proof signals to competitors and mates. Advances in the Study of Behavior, vol.30, pp.169-217, 2001.

K. Hackländer, E. Möstl, and W. Arnold, Reproductive suppression in female Alpine marmots, Marmota marmota, Animal Behaviour, vol.65, pp.1133-1140, 2003.

L. R. Hagey, M. A. Gavrilkina, and A. F. Hofmann, Agerelated changes in the biliary bile acid composition of bovids, Canadian Journal of Zoology, vol.75, pp.1193-1201, 1997.

R. L. Harris, N. W. Davies, and S. C. Nicol, Chemical composition of odorous secretions in the Tasmanian short-beaked echidna (Tachyglossus aculeatus setosus), Chemical Senses, vol.37, pp.819-836, 2012.

R. L. Harris, B. R. Holland, E. Z. Cameron, N. W. Davies, and S. C. Nicol, Chemical signals in the echidna: differences between seasons, sexes, individuals and gland types, Journal of Zoology, vol.293, pp.171-180, 2014.

M. E. Hauber and E. A. Lacey, Bateman's principle in cooperatively breeding vertebrates: the effects of non-breeding alloparents on variability in female and male reproductive success, Integrative and Comparative Biology, vol.45, pp.903-914, 2005.

R. A. Hayes, T. L. Morelli, and P. C. Wright, Anogenital gland secretions of Lemur catta and Propithecus verreauxi coquereli: a preliminary chemical examination, American Journal of Primatology, vol.63, pp.49-62, 2004.

R. A. Hayes, B. J. Richardson, and S. G. Wyllie, Semiochemicals and social signaling in the wild European rabbit in Australia: I. Scent profiles of chin gland secretion from the field, Journal of Chemical Ecology, vol.28, pp.363-384, 2002.

E. W. Haymann, Scent-marking strategies of New World primates, American Journal of Primatology, vol.68, pp.650-661, 2006.

B. Jemiolo, J. Alberts, S. Sochinski-wiggins, S. Harvey, and M. Novotny, Behavioural and endocrine responses of female mice to synthetic analogues of volatile compounds in male urine, Animal Behaviour, vol.33, pp.1114-1118, 1985.

B. G. Johansson and T. M. Jones, The role of chemical communication in mate choice, Biological Reviews of the Cambridge Philosophical Society, vol.82, pp.265-289, 2007.

R. A. Johnstone and M. A. Cant, Reproductive skew and the threat of eviction: a new perspective, Proceedings of the Royal Society B: Biological Sciences, vol.266, pp.275-279, 1999.

E. F. Kean, C. T. Müller, and E. A. Chadwick, Otter scent signals age, sex, and reproductive status, Chemical Senses, vol.36, pp.555-564, 2011.

J. Kwak, A. Willse, K. Matsumura, M. Curran-opiekun, Y. W. Preti et al., Geneticallybased olfactory signatures persist despite dietary variation, PLoS ONE, vol.3, p.3591, 2008.

S. Leclaire, J. F. Nielsen, and C. M. Drea, Bacterial communities in meerkat anal scent secretions vary with host sex, age, and group membership, Behavioral Ecology, vol.25, pp.996-1004, 2014.

W. L. Linklater, K. Mayer, and R. R. Swaisgood, Chemical signals of age, sex and identity in black rhinoceros, Animal Behaviour, vol.85, pp.671-677, 2013.

, The Linnean Society of London, Biological Journal of the Linnean Society, vol.125, pp.229-239, 2018.

E. A. Macdonald, E. Fernandez-duque, S. Evans, and L. R. Hagey, Sex, age, and family differences in the chemical composition of owl monkey (Aotus nancymaae) subcaudal scent secretions, American Journal of Primatology, vol.70, pp.12-18, 2008.

W. D. Magrath, R. A. Johnstone, and R. G. Heinsohn, Reproductive skew, Ecology and evolution of cooperative breeding in birds, pp.157-176, 2004.

J. Martín, J. Carranza, P. López, S. Alarcos, and J. Pérez-gonzález, A new sexual signal in rutting male red deer: age related chemical scent constituents in the belly black spot, Mammalian Biology, vol.79, pp.362-368, 2014.

S. Massemin, C. Gibault, R. Ramousse, and A. Butet, Premières données sur le régime alimentaire de la marmotte alpine (Marmota marmota) en France, Mammalia, vol.60, pp.351-361, 1996.

M. Novotny, S. Harvey, B. Jemiolo, and J. Alberts, Synthetic pheromones that promote inter-male aggression in mice, Proceedings of the National Academy of Sciences of the United States of America, vol.82, pp.2059-2061, 1985.

J. Oksanen, F. G. Blanchet, M. Friendly, R. Kindt, P. Legendre et al., , p.6, 2018.

K. Osada, M. Kashiwayanagi, and H. Izumi, Profiles of volatiles in male rat urine: the effect of puberty on the female attraction, Chemical Senses, vol.34, pp.713-721, 2009.

H. Psenner, Neues vom Murmeltier, Marmota m. marmota (Linné 1758), Säugetierkundliche Mitteilungen, vol.5, pp.4-10, 1957.

H. Psenner, Das verhalten der Murmeltiere (Marmota m. marmota), ins besondere von Mutter und Jungen, Säugetierkundliche Mitteilungen, vol.8, pp.144-148, 1960.

. R-core-team, R: a language and environment for statistical computing, p.6, 2017.

K. Ralls, Mammalian scent marking, Science, vol.171, pp.443-449, 1971.

L. Rasmussen, H. S. Riddle, and V. Krishnamurthy, Mellifluous matures to malodorous in musth, Nature, vol.415, pp.975-976, 2002.

F. Rosell, S. M. Jojola, K. Ingdal, B. A. Lassen, J. E. Swenson et al., Brown bears possess anal sacs and secretions may code for sex, Journal of Zoology, vol.283, pp.143-152, 2011.

W. Saltzman, K. J. Liedl, O. J. Salper, R. R. Pick, and D. H. Abbott, Post-conception reproductive competition in cooperatively breeding common marmosets, Hormones and Behavior, vol.53, pp.274-286, 2008.

W. Saltzman, S. D. Tardif, and J. N. Rutherford, Hormones and reproductive cycles in primates, Hormones and reproduction of vertebrates, pp.291-327, 2011.

M. L. Schaefer, K. Wongravee, M. E. Holmboe, N. M. Heinrich, S. J. Dixon et al., Mouse urinary biomarkers provide signatures of maturation, diet, stress level, and diurnal rhythm, Chemical Senses, vol.35, pp.459-471, 2010.

E. S. Scordato, G. Dubay, and C. M. Drea, Chemical composition of scent marks in the ringtailed lemur (Lemur catta): glandular differences, seasonal variation, and individual signatures, Chemical Senses, vol.32, pp.493-504, 2007.

J. M. Setchell, S. Vaglio, J. Moggi-cecchi, F. Boscaro, L. Calamai et al., Chemical composition of scent-gland secretions in an old world monkey (Mandrillus sphinx): influence of sex, male status, and individual identity, Chemical Senses, vol.35, pp.205-220, 2010.

A. Siberchicot, A. Julien-laferrière, A. B. Dufour, and J. Thioulouse, adegraphics: an S4 lattice-based package for the representation of multivariate data. R package version 1.0-9, 2017.
URL : https://hal.archives-ouvertes.fr/hal-01887857

J. B. Silk, The adaptive value of sociality in mammalian groups, Philosophical Transactions of the Royal Society of London. Series B, Biological Sciences, vol.362, pp.539-559, 2007.

A. Spence-aizenberg, B. A. Kimball, L. E. Williams, and E. Fernandez-duque, Chemical composition of glandular secretions from a pair-living monogamous primate: sex, age, and gland differences in captive and wild owl monkeys (Aotus spp, American Journal of Primatology, vol.80, p.22730, 2018.

H. V. Tinnesand, S. Jojola, A. Zedrosser, and F. Rosell, The smell of desperadoes? Beavers distinguish between dominant and subordinate intruders, Behavioral Ecology and Sociobiology, vol.67, pp.895-904, 2013.

S. Vaglio, P. Minicozzi, R. Romoli, F. Boscaro, G. Pieraccini et al., Sternal gland scentmarking signals sex, age, rank, and group identity in captive mandrills, Chemical Senses, vol.41, pp.177-186, 2016.

W. N. Venables and B. D. Ripley, Modern applied statistics with S, 2002.

T. E. Wolf, F. S. Schaebs, N. C. Bennett, R. Burroughs, and A. Ganswindt, Age and socially related changes in fecal androgen metabolite concentrations in free-ranging male giraffes, General and Comparative Endocrinology, vol.255, pp.19-25, 2018.

T. D. Wyatt, Pheromones and animal behavior: chemical signals and signatures, 2014.

A. J. Young, The causes of physiological suppression in vertebrate societies: a synthesis, Reproductive skew in vertebrates: proximate and ultimate causes, pp.397-436, 2009.

H. Yuan, D. Liu, L. Sun, R. Wei, G. Zhang et al., Anogenital gland secretions code for sex and age in the giant panda, Ailuropoda melanoleuca, Canadian Journal of Zoology, vol.82, pp.1596-1604, 2004.

R. Zabaras, B. J. Richardson, and S. G. Wyllie, Evolution in the suite of semiochemicals secreted by the sternal gland of Australian marsupials, Australian Journal of Zoology, vol.53, pp.257-263, 2005.

T. E. Ziegler, Social effects via olfactory sensory stimuli on reproductive function and dysfunction in cooperative breeding marmosets and tamarins, American Journal of Primatology, vol.75, pp.202-211, 2013.