F. Achard, R. Beuchle, P. Mayaux, H. Stibig, C. Bodart et al., Determination of tropical deforestation rates and related carbon losses from, Global Change Biology, vol.20, pp.2540-2554, 1990.

D. D. Ackerly and F. A. Bazzaz, Seedling Crown Orientation and Interception of Diffuse Radiation in Tropical Forest Gaps, Ecology, vol.76, pp.1134-1146, 1995.

A. Agrawal, D. Nepstad, and A. Chhatre, Reducing emissions from deforestation and forest degradation, Annual Review of Environment and Resources, vol.36, pp.373-396, 2011.

A. H. Aldred and G. M. Bonner, Application of airborne laser to forest surveys, 1985.

A. P. Allen, J. H. Brown, and J. F. Gillooly, Global biodiversity, biochemical kinetics, and the energetic-equivalence rule, Science, vol.297, pp.1545-1548, 2002.

A. P. Allen, J. F. Gillooly, V. M. Savage, and J. H. Brown, Kinetic effects of temperature on rates of genetic divergence and speciation, Proceedings of the National Academy of Sciences, vol.103, pp.9130-9135, 2006.

P. Álvarez-loayza, J. F. Jr, M. S. Torres, H. Balslev, T. Kristiansen et al., Light Converts Endosymbiotic Fungus to Pathogen, Influencing Seedling Survival and Niche-Space Filling of a Common Tropical Tree, Iriartea deltoidea, PLOS ONE, vol.6, p.16386, 2011.

J. D. Anadón, O. E. Sala, and F. T. Maestre, Climate change will increase savannas at the expense of forests and treeless vegetation in tropical and subtropical Americas, Journal of Ecology, vol.102, pp.1363-1373, 2014.

H. Andersen, S. E. Reutebuch, R. J. Mcgaughey, M. V. Oliveira, and M. Keller, Monitoring selective logging in western Amazonia with repeat lidar flights, Remote Sensing of Environment, vol.151, pp.157-165, 2014.

A. B. Anderson, White-sand vegetation of Brazilian Amazonia, Biotropica, vol.13, pp.199-210, 1981.

K. J. Anderson-teixeira, S. J. Davies, A. C. Bennett, E. B. Gonzalez-akre, H. C. Muller-landau et al., CTFS-ForestGEO: a worldwide network monitoring forests in an era of global change, Global change biology, vol.21, pp.528-549, 2015.

L. Aragão, Y. Malhi, D. B. Metcalfe, J. E. Silva-espejo, E. Jiménez et al., Above-and below-ground net primary productivity across ten Amazonian forests on contrasting soils, Biogeosciences, vol.6, pp.2759-2778, 2009.

L. E. Aragão, Y. Malhi, R. M. Roman-cuesta, S. Saatchi, L. O. Anderson et al., Spatial patterns and fire response of recent Amazonian droughts, Research Letters, vol.34, issue.07, 2007.

H. Arp and C. A. Tranarg, Mapping in tropical forests: a new approach using the laser APR, 1982.

, Photogrammetric Engineering and Remote Sensing, p.48

G. P. Asner, E. N. Broadbent, P. J. Oliveira, M. Keller, D. E. Knapp et al., Condition and fate of logged forests in the Brazilian Amazon, Proceedings of the National Academy of Sciences, vol.103, pp.12947-12950, 2006.

G. P. Asner, J. K. Clark, J. Mascaro, R. Vaudry, K. D. Chadwick et al., Maatoug, L. & others. (2012a) Human and environmental controls over aboveground carbon storage in Madagascar, p.7

G. P. Asner, R. F. Hughes, T. A. Varga, D. E. Knapp, and T. Kennedy-bowdoin, Environmental and biotic controls over aboveground biomass throughout a tropical rain forest, Ecosystems, vol.12, pp.261-278, 2008.

G. P. Asner, M. Keller, and J. N. Silva, Spatial and temporal dynamics of forest canopy gaps following selective logging in the eastern Amazon, Global Change Biology, vol.10, pp.765-783, 2004.

G. P. Asner, J. R. Kellner, T. Kennedy-bowdoin, D. E. Knapp, C. Anderson et al., Forest canopy gap distributions in the southern Peruvian Amazon, PLoS ONE, vol.8, p.60875, 2013.

G. P. Asner, D. E. Knapp, E. N. Broadbent, P. J. Oliveira, M. Keller et al., Selective logging in the brazilian Amazon, Science, vol.310, pp.480-482, 2005.

G. P. Asner and J. Mascaro, Mapping tropical forest carbon: Calibrating plot estimates to a simple LiDAR metric. Remote Sensing of Environment, vol.140, pp.614-624, 2014.

G. P. Asner, J. Mascaro, C. Anderson, D. E. Knapp, R. E. Martin et al., High-fidelity national carbon mapping for resource management and REDD+, Carbon Balance and Management, vol.8, pp.1-14, 2013.

G. Asner, J. Mascaro, H. Muller-landau, G. Vieilledent, R. Vaudry et al., A universal airborne LiDAR approach for tropical forest carbon mapping, Oecologia, vol.168, pp.1147-1160, 2012.

G. P. Asner, G. V. Powell, J. Mascaro, D. E. Knapp, J. K. Clark et al., High-resolution forest carbon stocks and emissions in the Amazon, Proceedings of the National Academy of Sciences, vol.107, pp.16738-16742, 2010.

I. Axmanová, M. Chytrý, D. Zelený, C. Li, M. Vymazalová et al., The species richnessproductivity relationship in the herb layer of European deciduous forests, Global Ecology and Biogeography, vol.21, pp.657-667, 2012.

T. R. Baker, O. L. Phillips, Y. Malhi, S. Almeida, L. Arroyo et al., Increasing biomass in Amazonian forest plots. Philosophical Transactions of the Royal Society of London B: Biological Sciences, vol.359, pp.353-365, 2004.

E. P. Baltsavias, Airborne laser scanning: basic relations and formulas, ISPRS Journal of Photogrammetry and Remote Sensing, vol.54, pp.199-214, 1999.

L. Banin, T. R. Feldpausch, O. L. Phillips, T. R. Baker, J. Lloyd et al., What controls tropical forest architecture? Testing environmental, structural and floristic drivers, Global Ecology and Biogeography, vol.21, pp.1179-1190, 2012.

C. Baraloto and D. E. Goldberg, Microhabitat associations and seedling bank dynamics in a neotropical forest, Oecologia, vol.141, pp.701-712, 2004.
URL : https://hal.archives-ouvertes.fr/hal-01031928

C. Baraloto, Q. Molto, S. Rabaud, B. Hérault, R. Valencia et al., Rapid simultaneous estimation of aboveground biomass and tree diversity across neotropical forests: a comparison of field inventory methods, Biotropica, vol.45, pp.288-298, 2013.
URL : https://hal.archives-ouvertes.fr/hal-01032408

G. L. Baskerville, Use of logarithmic regression in the estimation of plant biomass, Canadian Journal of Forest Research, vol.2, pp.49-53, 1972.

C. C. Baskin and J. M. Baskin, Seeds: Ecology, Biogeography, and Evolution of Dormancy and Germination, 2001.

J. S. Beard, The classification of tropical American vegetation-types, Ecology, vol.36, pp.89-100, 1955.

F. Béchamel, Journal du Voyage que les Pères Jean Grillet et François Béchamel ont Fait dans la Guyane en 1674

C. Beer, M. Reichstein, E. Tomelleri, P. Ciais, M. Jung et al., Terrestrial gross carbon dioxide uptake: global distribution and covariation with climate, Science, vol.329, pp.834-838, 2010.
URL : https://hal.archives-ouvertes.fr/cea-00819125

J. Belmaker and W. Jetz, Relative roles of ecological and energetic constraints, diversification rates and region history on global species richness gradients, Ecology Letters, vol.18, pp.563-571, 2015.

R. A. Betts, P. M. Cox, M. Collins, P. P. Harris, C. Huntingford et al., The role of ecosystem-atmosphere interactions in simulated Amazonian precipitation 156 decrease and forest dieback under global climate warming, Theoretical and applied climatology, vol.78, pp.157-175, 2004.

C. A. Bode, M. P. Limm, M. E. Power, and J. C. Finlay, Subcanopy Solar Radiation model: Predicting solar radiation across a heavily vegetated landscape using LiDAR and GIS solar radiation models, Remote Sensing of Environment, vol.154, pp.387-397, 2014.

O. M. Bollandsås, T. G. Gregoire, E. Naesset, and B. Øyen, Detection of biomass change in a Norwegian mountain forest area using small footprint airborne laser scanner data, Statistical Methods & Applications, vol.22, pp.113-129, 2013.

R. A. Bone, D. W. Lee, and J. M. Norman, Epidermal cells functioning as lenses in leaves of tropical rain-forest shade plants, Applied Optics, vol.24, p.1408, 1985.

F. Bongers, P. Charles-dominique, P. Forget, and M. Théry, Nouragues: Dynamics and Plant-Animal Interactions in a Neotropical Rainforest, 2001.

M. T. Bonner, S. Schmidt, and L. P. Shoo, A meta-analytical global comparison of biomass accumulation between tropical secondary forests and monoculture plantations, Forest Ecology and Management, vol.291, pp.73-86, 2013.

A. Brenning, RSAGA: SAGA geoprocessing and terrain analysis in R, 2007.

R. J. Brienen, O. L. Phillips, T. R. Feldpausch, E. Gloor, T. R. Baker et al., Nature, vol.519, pp.344-348, 2015.

N. V. Brokaw, The Definition of Treefall Gap and Its Effect on Measures of Forest Dynamics, Biotropica, vol.14, pp.158-160, 1982.

J. H. Brown, Why are there so many species in the tropics, Journal of Biogeography, vol.41, pp.8-22, 2014.

S. Brown and A. E. Lugo, Effects of forest clearing and succession on the carbon and nitrogen content of soils in Puerto Rico and US Virgin Islands, Plant and Soil, vol.124, pp.53-64, 1990.

H. Bugmann and C. Bigler, Will the CO2 fertilization effect in forests be offset by reduced tree longevity?, Oecologia, vol.165, pp.533-544, 2010.

G. Caballé, Essai sur la géographie forestière du Gabon, Adansonia, vol.17, pp.425-440, 1978.

G. S. Campbell, Extinction coefficients for radiation in plant canopies calculated using an ellipsoidal inclination angle distribution, Agricultural and Forest Meteorology, vol.36, pp.317-321, 1986.

A. Campos-arceiz and S. Blake, Megagardeners of the forest-the role of elephants in seed dispersal, Acta Oecologica, vol.37, pp.542-553, 2011.

C. D. Canham, A. C. Finzi, S. W. Pacala, and D. H. Burbank, Causes and consequences of resource heterogeneity in forests: interspecific variation in light transmission by canopy trees, Canadian Journal of Forest Research, vol.24, pp.337-349, 1994.

R. S. Capers and R. L. Chazdon, Rapid assessment of understory light availability in a wet tropical forest, Agricultural and Forest Meteorology, vol.123, pp.177-185, 2004.

N. Carvalhais, M. Forkel, M. Khomik, J. Bellarby, M. Jung et al., Global covariation of carbon turnover times with climate in terrestrial ecosystems, Nature, vol.514, pp.213-217, 2014.

L. A. Cernusak, K. Winter, J. W. Dalling, J. A. Holtum, C. Jaramillo et al., Tropical forest responses to increasing atmospheric CO2: current knowledge and opportunities for future research, Functional Plant Biology, vol.40, pp.531-551, 2013.

M. T. Chahine, The hydrological cycle and its influence on climate, Nature, vol.359, pp.373-380, 1992.

J. Q. Chambers, R. I. Negrón-juárez, G. C. Hurtt, D. M. Marra, and N. Higuchi, Lack of intermediate-scale disturbance data prevents robust extrapolation of plot-level tree mortality rates for old-growth tropical forests, Ecology Letters, vol.12, pp.22-25, 2009.

J. Chave, C. Andalo, S. Brown, M. Cairns, J. Chambers et al., Tree allometry and improved estimation of carbon stocks and balance in tropical forests, Oecologia, vol.145, pp.87-99, 2005.

J. Chave, D. Coomes, S. Jansen, S. L. Lewis, N. G. Swenson et al., Towards a worldwide wood economics spectrum, Ecology Letters, vol.12, pp.351-366, 2009.

J. Chave, H. C. Muller-landau, T. R. Baker, T. A. Easdale, H. Ter-steege et al., Regional and phylogenetic variation of wood density across 2456 neotropical tree species, Ecological Applications, vol.16, pp.2356-2367, 2006.
URL : https://hal.archives-ouvertes.fr/hal-00132042

J. Chave, D. Navarrete, S. Almeida, E. Alvarez, L. E. Aragão et al., Regional and seasonal patterns of litterfall in tropical South America, Biogeosciences, vol.7, pp.43-55, 2010.
URL : https://hal.archives-ouvertes.fr/hal-00594514

J. Chave, J. Olivier, F. Bongers, P. Châtelet, P. Forget et al., Above-ground biomass and productivity in a rain forest of eastern South America, Journal of Tropical Ecology, vol.24, pp.355-366, 2008.
URL : https://hal.archives-ouvertes.fr/mnhn-02184858

J. Chave, M. Réjou-méchain, A. Búrquez, E. Chidumayo, M. S. Colgan et al., Improved allometric models to estimate the aboveground biomass of tropical trees, Global Change Biology, vol.20, pp.3177-3190, 2014.
URL : https://hal.archives-ouvertes.fr/hal-02063299

J. Chave, B. Riéra, and M. Dubois, Estimation of biomass in a Neotropical forest of French Guiana: Spatial and temporal variability, Journal of Tropical Ecology, vol.17, pp.79-96, 2001.

R. L. Chazdon and N. Fetcher, Photosynthetic Light Environments in a Lowland Tropical Rain Forest in Costa Rica, Journal of Ecology, vol.72, pp.553-564, 1984.

R. L. Chazdon and R. W. Pearcy, The Importance of Sunflecks for Forest Understory Plants, BioScience, vol.41, pp.760-766, 1991.

S. Chen and M. Ravallion, The developing world is poorer than we thought, but no less successful in the fight against poverty, The Quarterly Journal of Economics, vol.125, pp.1577-1625, 2010.

D. A. Clark, S. Brown, D. W. Kicklighter, J. Q. Chambers, J. R. Thomlinson et al., Net primary production in tropical forests: an evaluation and synthesis of existing field data, Ecological applications, vol.11, pp.371-384, 2001.

M. L. Clark, D. B. Clark, and D. A. Roberts, Small-footprint lidar estimation of subcanopy elevation and tree height in a tropical rain forest landscape, Remote Sensing of Environment, vol.91, pp.68-89, 2004.

D. B. Clark and J. R. Kellner, Tropical forest biomass estimation and the fallacy of misplaced concreteness, Journal of Vegetation Science, vol.23, pp.1191-1196, 2012.

D. B. Clark, P. C. Olivas, S. F. Oberbauer, D. A. Clark, and M. G. Ryan, First direct landscape-scale measurement of tropical rain forest Leaf Area Index, a key driver of global primary productivity, Ecology Letters, vol.11, pp.163-172, 2008.

R. Condit, N. Pitman, E. G. Leigh, J. Chave, J. Terborgh et al., Beta-diversity in tropical forest trees. Science, vol.295, pp.666-669, 2002.

J. H. Connell, On the role of natural enemies in preventing competitive exclusion in some marine animals and in rain forest trees. Dynamics of populations, vol.298, p.159, 1971.

A. C. Da-costa, D. Galbraith, S. Almeida, B. T. Portela, M. Da-costa et al., Effect of 7 yr of experimental drought on vegetation dynamics and biomass storage of an eastern Amazonian rainforest, New Phytologist, vol.187, pp.579-591, 2010.

H. Coudreau, Chez nos Indiens : quatre années dans la Guyane française, pp.1887-1891, 1893.

P. Hachette,

P. M. Cox, D. Pearson, B. B. Booth, P. Friedlingstein, C. Huntingford et al., Sensitivity of tropical carbon to climate change constrained by carbon dioxide variability, Nature, vol.494, pp.341-344, 2013.

E. P. Crist and R. C. Cicone, A physically-based transformation of thematic mapper data-The TM tasseled cap. Geoscience and Remote Sensing, IEEE Transactions On, pp.256-263, 1984.

M. Cucumo, D. Kaliakatsos, and V. Marinelli, A calculation method for the estimation of the Linke turbidity factor, Renewable Energy, vol.19, pp.249-258, 2000.

K. C. Cushman, H. C. Muller-landau, R. S. Condit, and S. P. Hubbell, Improving estimates of biomass change in buttressed trees using tree taper models, Methods in Ecology and Evolution, vol.5, pp.573-582, 2014.

I. C. Cuthill, J. C. Partridge, A. T. Bennett, S. C. Church, N. S. Hart et al., Ultraviolet vision in birds, Advances in the Study of Behavior, vol.29, pp.159-214, 2000.

J. W. Dalling and S. P. Hubbell, Seed size, growth rate and gap microsite conditions as determinants of recruitment success for pioneer species, Journal of Ecology, vol.90, pp.557-568, 2002.

J. W. Dalling, S. A. Schnitzer, C. Baldeck, K. E. Harms, R. John et al., Resource-based habitat associations in a neotropical liana community, Journal of Ecology, vol.100, pp.1174-1182, 2012.

E. A. Davidson, A. C. De-araújo, P. Artaxo, J. K. Balch, I. F. Brown et al., The Amazon basin in transition, Nature, vol.481, pp.321-328, 2012.

A. B. Davies and G. P. Asner, Advances in animal ecology from 3D-LiDAR ecosystem mapping, Trends in Ecology & Evolution, vol.29, pp.681-691, 2014.

J. Dauzat and M. N. Eroy, Simulating light regime and intercrop yields in coconut based farming systems, European Journal of Agronomy, vol.7, pp.63-74, 1997.

R. S. Defries, R. A. Houghton, M. C. Hansen, C. B. Field, D. Skole et al., Carbon emissions from tropical deforestation and regrowth based on satellite observations for the 1980s and 1990s, Proceedings of the National Academy of Sciences, vol.99, pp.14256-14261, 2002.

M. Detto, G. P. Asner, H. C. Muller-landau, and O. Sonnentag, Spatial variability in tropical forest leaf area density from multireturn LiDAR and modeling, Journal of Geophysical Research, 2015.

S. J. Dewalt and J. Chave, Structure and biomass of four lowland Neotropical Forests, Biotropica, vol.36, pp.7-19, 2004.

S. J. Dewalt, S. K. Maliakal, and J. S. Denslow, Changes in vegetation structure and composition along a tropical forest chronosequence: implications for wildlife, Forest Ecology and Management, vol.182, pp.139-151, 2003.

R. Dirzo, C. C. Horvitz, H. Quevedo, and M. A. Lopez, The Effects of Gap Size and Age on the Understorey Herb Community of a Tropical Mexican Rain Forest, Journal of Ecology, vol.80, pp.809-822, 1992.

J. B. Drake, R. O. Dubayah, D. B. Clark, R. G. Knox, J. B. Blair et al., Estimation of tropical forest structural characteristics using large-footprint lidar, Remote Sensing of Environment, vol.79, pp.305-319, 2002.

J. B. Drake, R. G. Knox, R. O. Dubayah, D. B. Clark, R. Condit et al., Above-ground biomass estimation in closed canopy Neotropical forests using lidar remote sensing: factors affecting the generality of relationships, Global Ecology and Biogeography, vol.12, pp.147-159, 2003.

R. O. Dubayah, S. L. Sheldon, D. B. Clark, M. A. Hofton, J. B. Blair et al., Estimation of tropical forest height and biomass dynamics using lidar remote sensing at La Selva, Costa Rica, Journal of Geophysical Research, vol.115, 2010.

J. A. Dulk, The Interpretation of Remote Sensing: A Feasibility Study, 1989.

A. K. Duraiappah, Poverty and environmental degradation: a review and analysis of the nexus, World development, vol.26, pp.2169-2179, 1998.

E. A. Eltahir and R. L. Bras, Precipitation recycling in the Amazon basin, Quarterly Journal of the Royal Meteorological Society, vol.120, pp.861-880, 1994.

L. H. Emmons and A. H. Gentry, Tropical forest structure and the distribution of gliding and prehensile-tailed vertebrates, American Naturalist, vol.121, pp.513-524, 1983.

J. A. Endler, The Color of Light in Forests and Its Implications, Ecological Monographs, vol.63, pp.2-27, 1993.

J. A. Endler and M. Thery, Interacting Effects of Lek Placement, Display Behavior, Ambient Light, and Color Patterns in Three Neotropical Forest-Dwelling Birds, The American Naturalist, vol.148, pp.421-452, 1996.

B. M. Engelbrecht and H. M. Herz, Evaluation of Different Methods to Estimate Understorey Light Conditions in Tropical Forests, Journal of Tropical Ecology, vol.17, pp.207-224, 2001.

S. Englhart, V. Keuck, and F. Siegert, Aboveground biomass retrieval in tropical forests -The potential of combined X-and L-band SAR data use, Remote Sensing of Environment, vol.115, pp.1260-1271, 2011.

S. Englhart, J. Jubanski, and F. Siegert, Quantifying dynamics in tropical peat swamp forest biomass with multi-temporal lidar datasets, Remote Sensing, vol.5, pp.2368-2388, 2013.

H. D. Eva, A. S. Belward, E. E. De-miranda, C. M. Di-bella, V. Gond et al., ) A land cover map of South America, Global Change Biology, vol.10, pp.731-744, 2004.

G. D. Farquhar, S. Von-caemmerer, and J. A. Berry, A biochemical model of photosynthetic CO2 assimilation in leaves of C3 species, Planta, vol.149, pp.78-90, 1980.

S. Fauset, M. O. Johnson, M. Gloor, T. R. Baker, M. et al., Nature Communications, p.6

P. M. Fearnside, Environment: Deforestation soars in the Amazon, Nature, vol.521, pp.423-423, 2015.

T. R. Feldpausch, L. Banin, O. L. Phillips, T. R. Baker, S. L. Lewis et al., Biogeosciences, vol.8, pp.1081-1106, 2011.

T. R. Feldpausch, J. Lloyd, S. L. Lewis, R. J. Brienen, M. Gloor et al., , p.162

E. J. Arroyo, L. Aymard, C. , G. A. Baker, T. R. Bánki et al., Biogeosciences, vol.9, pp.3381-3403

T. R. Feldpausch, M. A. Rondon, E. C. Fernandes, S. J. Riha, and E. Wandelli, Carbon and nutrient accumulation in secondary forests regenerating on pastures in central amazonia, Ecological Applications, vol.14, pp.164-176, 2004.

C. B. Field, M. J. Behrenfeld, J. T. Randerson, and P. Falkowski, Primary production of the biosphere: integrating terrestrial and oceanic components, Science, vol.281, pp.237-240, 1998.

A. G. Fischer, Latitudinal variations in organic diversity, Evolution, vol.14, pp.64-81, 1960.

T. H. Fleming, R. Breitwisch, and G. H. Whitesides, Patterns of tropical vertebrate frugivore diversity, Annual Review of Ecology and Systematics, vol.18, pp.91-109, 1987.

, Food Agriculture Organization of the United Nations, 2010.

J. R. Foster, P. A. Townsend, and C. E. Zganjar, Spatial and temporal patterns of gap dominance by low-canopy lianas detected using EO-1 Hyperion and Landsat Thematic Mapper. Remote Sensing of Environment, vol.112, pp.2104-2117, 2008.

D. Galbraith, P. E. Levy, S. Sitch, C. Huntingford, P. Cox et al., Multiple mechanisms of Amazonian forest biomass losses in three dynamic global vegetation models under climate change, New Phytologist, vol.187, pp.647-665, 2010.

D. Galbraith, Y. Malhi, K. Affum-baffoe, A. D. Castanho, C. E. Doughty et al., Residence times of woody biomass in tropical forests, Plant Ecology & Diversity, vol.6, pp.139-157, 2013.

C. Gehrig-downie, A. Obregón, J. Bendix, and S. R. Gradstein, Epiphyte biomass and canopy microclimate in the tropical lowland cloud forest of French Guiana, Biotropica, vol.43, pp.591-596, 2011.

J. J. Gerwing, S. A. Schnitzer, R. J. Burnham, F. Bongers, J. Chave et al., A standard protocol for liana censuses, Biotropica, vol.38, pp.256-261, 2006.

H. K. Gibbs, L. Rausch, J. Munger, I. Schelly, D. C. Morton et al., Brazil's soy moratorium, Science, vol.347, pp.377-378, 2015.

P. Glaister, 13 Least Squares Revisited. The Mathematical Gazette, vol.85, pp.104-107, 2001.

S. J. Goetz, D. Steinberg, M. G. Betts, R. T. Holmes, P. J. Doran et al., Lidar remote sensing variables predict breeding habitat of a Neotropical migrant bird, Ecology, vol.91, pp.1569-1576, 2010.

P. Good, C. Jones, J. Lowe, R. Betts, B. Booth et al., Quantifying environmental drivers of future tropical forest extent, Journal of Climate, vol.24, pp.1337-1349, 2011.

J. Granados and C. Körner, In deep shade, elevated CO2 increases the vigor of tropical climbing plants, Global Change Biology, vol.8, pp.1109-1117, 2002.

A. Granier, N. Bréda, B. Longdoz, P. Gross, and J. Ngao, Ten years of fluxes and stand growth in a young beech forest at Hesse, Annals of Forest Science, vol.65, p.1, 2008.
URL : https://hal.archives-ouvertes.fr/hal-00883444

J. D. Granville, Phytogeographical Characteristics of the Guianan Forests, Taxon, vol.37, pp.578-594, 1988.

, Geographic Resources Analysis Support System (GRASS) Software. Open Source Geospatial Foundation Project, GRASS Development Team, 2012.

J. Grillet and F. Béchamel, Journal du voyage que les pères jean grillet et françois béchamel ont fait dans la goyane, l'an 1674. C.Barbin, 1682.

S. M. Guzman-grajales and L. R. Walker, Differential seedling responses to litter after hurricane Hugo in the Luquillo experimental forest, Biotropica, vol.23, pp.407-413, 1991.

J. C. Hagar, B. N. Eskelson, P. K. Haggerty, S. K. Nelson, and D. G. Vesely, Modeling marbled murrelet (Brachyramphus marmoratus) habitat using LiDAR-derived canopy data, vol.38, pp.237-249, 2014.

M. C. Hansen, P. V. Potapov, R. Moore, M. Hancher, S. A. Turubanova et al., High-resolution global maps of 21st-century forest cover change, Science, vol.342, pp.850-853, 2013.

D. J. Harding, M. A. Lefsky, G. G. Parker, and J. B. Blair, Laser altimeter canopy height profiles: methods and validation for closed-canopy, broadleaf forests. Remote Sensing of Environment, vol.76, pp.283-297, 2001.

K. E. Harms, R. Condit, S. P. Hubbell, and R. B. Foster, Habitat associations of trees and shrubs in a 50-ha neotropical forest plot, Journal of Ecology, vol.89, pp.947-959, 2001.

N. L. Harris, S. Brown, S. C. Hagen, S. S. Saatchi, S. Petrova et al., Baseline map of carbon emissions from deforestation in tropical regions, Science, vol.336, pp.1573-1576, 2012.

G. M. Van-der-heijden, T. R. Feldpausch, A. Herrero, F. De-la, N. K. Van-der-velden et al., Calibrating the liana crown occupancy index in Amazonian forests, Forest Ecology and Management, vol.260, pp.549-555, 2010.

G. M. Van-der-heijden, O. L. Phillips, and S. A. Schnitzer, Impacts of lianas on forestlevel carbon storage and sequestration, Ecology of Lianas, pp.164-174, 2015.

G. M. Van-der-heijden and O. L. Phillips, Liana infestation impacts tree growth in a lowland tropical moist forest, Biogeosciences, vol.6, pp.2217-2226, 2009.

G. M. Van-der-heijden, S. A. Schnitzer, J. S. Powers, and O. L. Phillips, Liana impacts on carbon cycling, storage and sequestration in tropical forests, Biotropica, vol.45, pp.682-692, 2013.

M. Heimann and M. Reichstein, Terrestrial ecosystem carbon dynamics and climate feedbacks, Nature, vol.451, pp.289-292, 2008.

J. Heiskanen, L. Korhonen, J. Hietanen, and P. K. Pellikka, Use of airborne lidar for estimating canopy gap fraction and leaf area index of tropical montane forests, International Journal of Remote Sensing, vol.36, pp.2569-2583, 2015.

W. H. Hendershot, H. Lalande, and M. Duquette, Ion exchange and exchangeable cations, Soil Sampling and Methods of Analysis, vol.19, pp.167-176, 1993.

R. A. Hill, Image segmentation for humid tropical forest classification in Landsat TM data, International Journal of Remote Sensing, vol.20, pp.1039-1044, 1999.

R. A. Hill and S. A. Hinsley, Airborne LiDAR for woodland habitat quality monitoring: exploring the significance of LiDAR data characteristics when modelling organismhabitat relationships, Remote Sensing, vol.7, pp.3446-3466, 2015.

R. A. Hill, S. A. Hinsley, and R. K. Broughton, Assessing Habitats and Organism-Habitat Relationships by Airborne Laser Scanning. Forestry Applications of Airborne Laser Scanning, Managing Forest Ecosystems, pp.335-356, 2014.

J. Hofierka and M. Suri, The solar radiation model for open source GIS: implementation and applications, Proceedings of the Open source GIS-GRASS users conference, pp.1-19, 2002.

L. R. Holdridge, Life Zone Ecology, revised. Tropical Science Center, 1967.

J. Hortal, J. A. Diniz-filho, L. M. Bini, M. Á. Rodríguez, A. Baselga et al., Ice age climate, evolutionary constraints and diversity patterns of European dung beetles, Ecology Letters, vol.14, pp.741-748, 2011.

F. Hosoi and K. Omasa, Voxel-based 3-D modeling of individual trees for estimating leaf area density using high-resolution portable scanning lidar. Geoscience and Remote Sensing, IEEE Transactions on, vol.44, pp.3610-3618, 2006.

J. T. Houghton, The Global Climate, reprinted. CUP Archive, 1985.

H. F. Howe and J. Smallwood, Ecology of seed dispersal. Annual review of ecology and systematics, vol.13, pp.201-228, 1982.

S. P. Hubbell, The Unified Neutral Theory of Biodiversity and Biogeography (MPB-32), 2001.

S. P. Hubbell, R. B. Foster, S. T. O'brien, K. E. Harms, R. Condit et al., Light-gap disturbances, recruitment limitation, and tree diversity in a neotropical forest, Science, vol.283, pp.554-557, 1999.

A. T. Hudak, E. K. Strand, L. A. Vierling, J. C. Byrne, J. U. Eitel et al., Quantifying aboveground forest carbon pools and fluxes from repeat LiDAR surveys, Remote Sensing of Environment, vol.123, pp.25-40, 2012.

M. M. Humphries and K. S. Mccann, Metabolic ecology, Journal of Animal Ecology, vol.83, pp.7-19, 2014.

M. O. Hunter, M. Keller, D. Morton, B. Cook, M. Lefsky et al., Structural dynamics of tropical moist forest gaps, PLoS ONE, vol.10, p.132144, 2015.

C. Huntingford, P. Zelazowski, D. Galbraith, L. M. Mercado, S. Sitch et al., Simulated resilience of tropical rainforests to CO2-induced climate change, Nature Geoscience, vol.6, pp.268-273, 2013.

A. H. Hurlbert, Linking species-area and species-energy relationships in Drosophila microcosms, Ecology letters, vol.9, pp.287-294, 2006.

P. Hyde, R. Dubayah, B. Peterson, J. B. Blair, M. Hofton et al., Mapping forest structure for wildlife habitat analysis using waveform lidar: Validation of montane ecosystems, Remote Sensing of Environment, vol.96, pp.427-437, 2005.

P. Hyde, R. Dubayah, W. Walker, J. B. Blair, M. Hofton et al., Mapping forest structure for wildlife habitat analysis using multi-sensor (LiDAR, vol.102, pp.63-73, 2006.

L. L. Ingwell, J. Wright, S. Becklund, K. K. Hubbell, S. P. Schnitzer et al., The impact of lianas on 10 years of tree growth and mortality on, vol.98, pp.879-887, 2010.

M. Insenburg, LAStools -Efficient LiDAR Processing Software, 2014.

D. Jablonski, The tropics as a source of evolutionary novelty through geological time, Nature, vol.364, pp.142-144, 1993.

D. Jablonski, K. Roy, and J. W. Valentine, Out of the tropics: evolutionary dynamics of the latitudinal diversity gradient, Science, vol.314, pp.102-106, 2006.

D. H. Janzen, Herbivores and the number of tree species in tropical forests, The American Naturalist, vol.104, pp.501-528, 1970.

W. Jetz and P. V. Fine, Global gradients in vertebrate diversity predicted by historical area-productivity dynamics and contemporary environment, PLoS Biol, vol.10, p.1001292, 2012.

W. Jetz, H. Kreft, G. Ceballos, and J. Mutke, Global associations between terrestrial producer and vertebrate consumer diversity, Proceedings of the Royal Society of London B: Biological Sciences, vol.276, pp.269-278, 2009.

J. Jubanski, U. Ballhorn, K. Kronseder, and F. Siegert, Detection of large above-ground biomass variability in lowland forest ecosystems by airborne LiDAR, Biogeosciences, vol.10, pp.3917-3930, 2013.

J. R. Kellner and G. P. Asner, Convergent structural responses of tropical forests to diverse disturbance regimes, Ecology Letters, vol.12, pp.887-897, 2009.

J. R. Kellner, G. P. Asner, P. M. Vitousek, M. A. Tweiten, S. Hotchkiss et al., Dependence of forest structure and dynamics on substrate age and ecosystem development, Ecosystems, vol.14, pp.1156-1167, 2011.

D. Kim, J. O. Sexton, and J. R. Townshend, Accelerated deforestation in the humid tropics from the 1990s to the, Geophysical Research Letters, vol.42, 2000.

A. Kleidon and M. Heimann, Assessing the role of deep rooted vegetation in the climate system with model simulations: mechanism, comparison to observations and implications for Amazonian deforestation, Climate Dynamics, vol.16, pp.183-199, 2000.

P. H. Klopfer and R. H. Macarthur, On the causes of tropical species diversity: niche overlap, The American Naturalist, vol.95, pp.223-226, 1961.

H. Kobayashi, D. D. Baldocchi, Y. Ryu, Q. Chen, S. Ma et al., Modeling energy and carbon fluxes in a heterogeneous oak woodland: A threedimensional approach, Agricultural and Forest Meteorology, vol.152, pp.83-100, 2012.

C. Körner, Through enhanced tree dynamics carbon dioxide enrichment may cause tropical forests to lose carbon, Philosophical Transactions of the Royal Society of London B: Biological Sciences, vol.359, pp.493-498, 2004.

N. J. Kraft, L. S. Comita, J. M. Chase, N. J. Sanders, N. G. Swenson et al., Disentangling the drivers of ? diversity along latitudinal and elevational gradients, J. & others, vol.333, pp.1755-1758, 2011.

J. V. Lafrankie, P. S. Ashton, G. B. Chuyong, L. Co, R. Condit et al., Lagunzad, D. & others. (2006) Contrasting structure and 167 composition of the understory in species-rich tropical rain forests, Ecology, vol.87, pp.2298-2305

R. Lahtinen, Archaean-proterozoic transition: geochemistry, provenance and tectonic setting of metasedimentary rocks in central Fennoscandian shield, Precambrian Research, vol.104, pp.147-174, 2000.

W. F. Laurance, A. S. Andrade, A. Magrach, J. L. Camargo, J. J. Valsko et al., Long-term changes in liana abundance and forest dynamics in undisturbed Amazonian forests, Ecology, vol.95, pp.1604-1611, 2014.

W. F. Laurance, P. Delamônica, S. G. Laurance, H. L. Vasconcelos, and T. E. Lovejoy, Conservation: Rainforest fragmentation kills big trees, Nature, vol.404, pp.836-836, 2000.

W. F. Laurance, H. E. Nascimento, S. G. Laurance, A. C. Andrade, P. M. Fearnside et al., Rain forest fragmentation and the proliferation of successional trees, Ecology, vol.87, pp.469-482, 2006.

W. F. Laurance, D. Pérez-salicrup, P. Delamônica, P. M. Fearnside, S. Angelo et al., Rain forest fragmentation and the structure of Amazonian liana communities, Ecology, vol.82, pp.105-116, 2001.

A. Ledo and S. A. Schnitzer, Disturbance and clonal reproduction determine liana distribution and maintain liana diversity in a tropical forest, Ecology, vol.95, pp.2169-2178, 2014.

D. W. Lee, The Spectral Distribution of Radiation in Two Neotropical Rainforests, Biotropica, vol.19, pp.161-166, 1987.

H. Lee, K. C. Slatton, B. E. Roth, and W. P. Cropper, Prediction of forest canopy light interception using three-dimensional airborne LiDAR data, International Journal of Remote Sensing, vol.30, pp.189-207, 2009.

M. A. Lefsky, W. B. Cohen, G. G. Parker, and D. J. Harding, Lidar Remote Sensing for Ecosystem Studies Lidar, an emerging remote sensing technology that directly measures the three-dimensional distribution of plant canopies, can accurately estimate vegetation structural attributes and should be of particular interest to forest, landscape, and global ecologists, BioScience, vol.52, pp.19-30, 2002.

S. G. Letcher and R. L. Chazdon, Life history traits of lianas during tropical forest succession, Biotropica, vol.44, pp.720-727, 2012.

S. L. Lewis, P. M. Brando, O. L. Phillips, G. M. Van-der-heijden, and D. Nepstad, The 2010 Amazon Drought, Science, vol.331, pp.554-554, 2011.

S. L. Lewis, D. P. Edwards, and D. Galbraith, Increasing human dominance of tropical forests, Science, vol.349, pp.827-832, 2015.

S. L. Lewis, J. Lloyd, S. Sitch, E. T. Mitchard, and W. F. Laurance, Changing ecology of tropical forests: evidence and drivers, Evolution, and Systematics, vol.40, pp.529-549, 2009.

S. L. Lewis and M. A. Maslin, Defining the Anthropocene, Nature, vol.519, pp.171-180, 2015.

S. L. Lewis, O. L. Phillips, T. R. Baker, J. Lloyd, Y. Malhi et al., Concerted changes in tropical forest structure and dynamics: evidence from 50 South American long-term plots, Philosophical Transactions of the Royal Society of London. Series B: Biological Sciences, vol.359, pp.421-436, 2004.

S. L. Lewis, B. Sonké, T. Sunderland, S. K. Begne, G. Lopez-gonzalez et al., Above-ground biomass and structure of 260 African tropical forests. Philosophical Transactions of the Royal Society of London B: Biological Sciences, vol.368, 2013.

H. Lieth, Primary production of the major vegetation units of the world, Primary Productivity of the Biosphere, Ecological Studies, pp.203-215, 1975.

K. Lim, P. Treitz, M. Wulder, B. St-onge, and M. Flood, LiDAR remote sensing of forest structure, Progress in Physical Geography, vol.27, pp.88-106, 2003.

K. Linnet, Evaluation of regression procedures for methods comparison studies, Clinical Chemistry, vol.39, pp.424-432, 1993.

J. Lloyd and G. D. Farquhar, Effects of rising temperatures and [CO2] on the physiology of tropical forest trees, Philosophical Transactions of the Royal Society of London B: Biological Sciences, vol.363, pp.1811-1817, 2008.

M. V. Lomolino, B. R. Riddle, R. J. Whittaker, and J. H. Brown, , 2010.

M. H. Long, J. E. Rheuban, P. Berg, and J. C. Zieman, A comparison and correction of light intensity loggers to photosynthetically active radiation sensors, Limnology and Oceanography: Methods, vol.10, pp.416-424, 2012.

G. Lopez-gonzalez, S. L. Lewis, M. Burkitt, T. R. Baker, and O. L. Phillips, Forestplots. net database. University of Leeds, p.13, 2009.

G. Lopez-gonzalez, S. L. Lewis, M. Burkitt, and O. L. Phillips, ForestPlots. net: a web application and research tool to manage and analyse tropical forest plot data, Journal of Vegetation Science, vol.22, pp.610-613, 2011.

J. R. Lozada, C. Hernández, P. Soriano, and M. Costa, An assessment of the floristic composition, structure and possible origin of a liana forest in the Guayana Shield, Plant Biosystems -An International Journal Dealing with all Aspects of Plant Biology, vol.0, pp.1-10, 2015.

D. Lu, The potential and challenge of remote sensing-based biomass estimation, International Journal of Remote Sensing, vol.27, pp.1297-1328, 2006.

A. Luckman, J. Baker, M. Honzák, and R. Lucas, Tropical forest biomass density estimation using JERS-1 SAR: Seasonal variation, confidence limits, and application to image mosaics, Remote Sensing of Environment, vol.63, pp.126-139, 1998.

Y. Malhi, F. F. Amézquita, C. E. Doughty, J. E. Silva-espejo, C. A. Girardin et al., The productivity, metabolism and carbon cycle of two lowland tropical forest plots in south-western Amazonia, Peru. Plant Ecology & Diversity, vol.7, pp.85-105, 2014.

Y. Malhi, T. R. Baker, O. L. Phillips, S. Almeida, E. Alvarez et al., The above-ground coarse wood productivity of 104 Neotropical forest plots, Global Change Biology, vol.10, pp.563-591, 2004.

Y. Malhi, A. D. Nobre, J. Grace, B. Kruijt, M. G. Pereira et al., Carbon dioxide transfer over a Central Amazonian rain forest, Journal of Geophysical Research, vol.103, pp.31593-31612, 1984.

Y. Malhi, J. T. Roberts, R. A. Betts, T. J. Killeen, W. Li et al., Climate change, deforestation, and the fate of the Amazon, Science, vol.319, pp.169-172, 2008.

Y. Malhi, D. Wood, T. R. Baker, J. Wright, O. L. Phillips et al., The regional variation of aboveground live biomass in old-growth Amazonian forests, Global Change Biology, vol.12, pp.1107-1138, 2006.

E. Marín-spiotta, W. L. Silver, and R. Ostertag, Long-term patterns in tropical reforestation: plant community composition and aboveground biomass accumulation, Ecological Applications, vol.17, pp.828-839, 2007.

T. R. Marthews, D. F. Burslem, S. R. Paton, F. Yangüez, and C. E. Mullins, Soil drying in a tropical forest: Three distinct environments controlled by gap size, Ecological Modelling, vol.216, pp.369-384, 2008.

J. Mascaro, G. P. Asner, S. Davies, A. Dehgan, and S. Saatchi, These are the days of lasers in the jungle, Carbon Balance and Management, vol.9, p.7, 2014.

J. Mascaro, G. P. Asner, H. C. Muller-landau, M. Van-breugel, J. Hall et al., Controls over aboveground forest carbon density on, vol.8, pp.1615-1629, 2011.

J. Mascaro, M. Detto, G. P. Asner, and H. C. Muller-landau, Evaluating uncertainty in mapping forest carbon with airborne LiDAR. Remote Sensing of Environment, vol.115, pp.3770-3774, 2011.

R. J. Mcgaughey, FUSION/LDV: Software for LIDAR data analysis and visualization. US Department of Agriculture, p.123, 2012.

S. M. Mclennan, S. Hemming, D. K. Mcdaniel, and G. N. Hanson, Geochemical approaches to sedimentation, provenance, and tectonics, vol.284, pp.21-40, 1993.

P. J. Van-der-meer and F. Bongers, Patterns of tree-fall and branch-fall in a tropical rain forest in french guiana, Journal of Ecology, vol.84, pp.19-29, 1996.

S. Mermoz, M. Réjou-méchain, L. Villard, T. Le-toan, V. Rossi et al., Decrease of L-band SAR backscatter with biomass of dense forests, Remote Sensing of Environment, vol.159, pp.307-317, 2015.
URL : https://hal.archives-ouvertes.fr/hal-01902422

V. Meyer, S. S. Saatchi, J. Chave, J. W. Dalling, S. Bohlman et al., Detecting tropical forest biomass dynamics from repeated airborne lidar measurements, Biogeosciences, vol.10, pp.5421-5438, 2013.

D. L. Miranda, . De, C. R. Sanquetta, L. G. Costa, S. Da et al., Biomassa e carbono em Euterpe oleracea Mart. na ilha do Marajó -PA, vol.19, pp.336-343, 2012.

E. T. Mitchard, T. R. Feldpausch, R. J. Brienen, G. Lopez-gonzalez, A. Monteagudo et al., Global Ecology and Biogeography, vol.23, pp.935-946, 2014.

J. Monteith and M. Unsworth, Principles of Environmental Physics: Plants, Animals, and the Atmosphere, 2013.

R. A. Montgomery and R. L. Chazdon, Forest structure, canopy architecture, and light transmittance in tropical wet forests, Ecology, vol.82, pp.2707-2718, 2001.

R. Montgomery and R. Chazdon, Light gradient partitioning by tropical tree seedlings in the absence of canopy gaps, Oecologia, vol.131, pp.165-174, 2002.

P. R. Moorcroft, G. C. Hurtt, and S. W. Pacala, A method for scaling vegetation dynamics: the ecosystem demography model (ed), Ecological Monographs, vol.71, pp.557-586, 2001.

W. Mücke and M. Hollaus, Modelling light conditions in forests using airborne laser scanning data, Proceedings of the SilviLaser, 2011.

H. C. Muller-landau, Interspecific and inter-site variation in wood specific gravity of tropical trees, Biotropica, vol.36, pp.20-32, 2004.

H. C. Muller-landau, R. S. Condit, J. Chave, S. C. Thomas, S. A. Bohlman et al., Testing metabolic ecology theory for allometric scaling of tree size, Ecology Letters, vol.9, pp.575-588, 2006.
URL : https://hal.archives-ouvertes.fr/hal-00131882

E. Naesset, O. M. Bollandsås, T. Gobakken, T. G. Gregoire, and G. Ståhl, Modelassisted estimation of change in forest biomass over an 11year period in a sample survey supported by airborne LiDAR: A case study with post-stratification to provide "activity data, Remote Sensing of Environment, vol.128, pp.299-314, 2013.

R. Nelson, W. Krabill, and J. Tonelli, Estimating forest biomass and volume using airborne laser data. Remote sensing of environment, vol.24, pp.247-267, 1988.

D. Nepstad, P. Lefebvre, U. Lopes-da-silva, J. Tomasella, P. Schlesinger et al., Amazon drought and its implications for forest flammability and tree growth: A basin-wide analysis, Global Change Biology, vol.10, pp.704-717, 2004.

D. Nepstad, D. Mcgrath, C. Stickler, A. Alencar, A. Azevedo et al., Slowing Amazon deforestation through public policy and interventions in beef and soy supply chains, Science, vol.344, pp.1118-1123, 2014.

D. C. Nepstad, I. M. Tohver, D. Ray, P. Moutinho, and G. Cardinot, Mortality of large trees and lianas following experimental drought in an Amazon forest, Ecology, vol.88, pp.2259-2269, 2007.

A. B. Nicotra, R. L. Chazdon, and S. V. Iriarte, Spatial heterogeneity of light and woody seedling regeneration in tropical wet forests, Ecology, vol.80, pp.1908-1926, 1999.

W. Nijland, N. C. Coops, S. E. Nielsen, and G. Stenhouse, Integrating optical satellite data and airborne laser scanning in habitat classification for wildlife management, International Journal of Applied Earth Observation and Geoinformation, vol.38, pp.242-250, 2015.

N. Norden, J. Chave, P. Belbenoit, A. Caubère, P. Châtelet et al., Interspecific variation in seedling responses to seed limitation and habitat conditions for 14 Neotropical woody species, Journal of Ecology, vol.97, pp.186-197, 2009.
URL : https://hal.archives-ouvertes.fr/mnhn-02184845

N. Norden, J. Chave, A. Caubère, P. Châtelet, N. Ferroni et al., Is temporal variation of seedling communities determined by environment or by seed arrival? A test in a Neotropical forest, Journal of Ecology, vol.95, pp.507-516, 2007.
URL : https://hal.archives-ouvertes.fr/mnhn-02184893

A. Obregon, C. Gehrig-downie, S. R. Gradstein, R. Rollenbeck, and J. Bendix, Canopy level fog occurrence in a tropical lowland forest of French Guiana as a prerequisite for high epiphyte diversity, Agricultural and Forest Meteorology, vol.151, pp.290-300, 2011.

I. Olivares, J. Svenning, P. M. Van-bodegom, and H. Balslev, Effects of warming and drought on the vegetation and plant diversity in the Amazon basin, The Botanical Review, vol.81, pp.42-69, 2015.

M. V. D'-oliveira, S. E. Reutebuch, R. J. Mcgaughey, and H. Andersen, Estimating forest biomass and identifying low-intensity logging areas using airborne scanning lidar in Antimary State Forest, Acre State, Western Brazilian Amazon. Remote Sensing of Environment, vol.124, pp.479-491, 2012.

M. B. Palomaki, R. L. Chazdon, J. P. Arroyo, and S. G. Letcher, Juvenile tree growth in relation to light availability in second-growth tropical rain forests, Journal of Tropical Ecology, vol.22, pp.223-226, 2006.

Y. Pan, R. A. Birdsey, J. Fang, R. Houghton, P. E. Kauppi et al., & others. (2011) A large and persistent carbon sink in the world's forests, Science, vol.333, pp.988-993

G. G. Parker, M. A. Lefsky, and D. J. Harding, Light transmittance in forest canopies determined using airborne laser altimetry and in-canopy quantum measurements, Remote Sensing of Environment, vol.76, pp.298-309, 2001.

S. Peng, C. Zhao, and Z. Xu, Modeling spatiotemporal patterns of understory light intensity using airborne laser scanner (LiDAR), ISPRS Journal of Photogrammetry and Remote Sensing, vol.97, pp.195-203, 2014.

D. R. Pérez-salicrup, Effect of liana cutting on tree regeneration in a liana forest in Amazonian Bolivia, Ecology, vol.82, pp.389-396, 2001.

O. L. Phillips, L. E. Aragão, S. L. Lewis, J. B. Fisher, J. Lloyd et al., Drought sensitivity of the Amazon rainforest, A. & others, vol.323, pp.1344-1347, 2009.
URL : https://hal.archives-ouvertes.fr/hal-01032111

O. L. Phillips, T. R. Baker, L. Arroyo, N. Higuchi, T. J. Killeen et al., Pattern and process in Amazon tree turnover, Philosophical Transactions of the Royal Society of London Series B-Biological Sciences, vol.359, pp.381-407, 1976.

O. L. Phillips and A. H. Gentry, Increasing turnover through time in tropical forests, Science, vol.263, pp.954-958, 1994.

O. L. Phillips, P. N. Vargas, A. L. Monteagudo, A. P. Cruz, M. C. Zans et al., Habitat association among Amazonian tree species: a landscape-scale approach, Journal of Ecology, vol.91, pp.757-775, 2003.

O. L. Phillips, R. Vásquez-martínez, L. Arroyo, T. R. Baker, T. Killeen et al., Increasing dominance of large lianas in Amazonian forests, Nature, vol.418, pp.770-774, 2002.

E. R. Pianka, Latitudinal gradients in species diversity: a review of concepts, American Naturalist, vol.100, pp.33-46, 1966.

A. Porté and H. H. Bartelink, Modelling mixed forest growth: a review of models for forest management, Ecological Modelling, vol.150, pp.141-188, 2002.

C. A. Price, J. S. Weitz, V. M. Savage, J. Stegen, A. Clarke et al., Testing the metabolic theory of ecology, Ecology Letters, vol.15, pp.1465-1474, 2012.

F. E. Putz, Liana biomass and leaf area of a "tierra firme" forest in the Rio Negro basin, Biotropica, vol.15, pp.185-189, 1983.

F. E. Putz, The natural history of lianas on Barro Colorado Island, Panama. Ecology, vol.65, pp.1713-1724, 1984.

F. E. Putz, How trees avoid and shed lianas, Biotropica, vol.16, pp.19-23, 1984.

F. E. Putz and C. Romero, Futures of tropical forests (sensu lato), Biotropica, vol.46, pp.495-505, 2014.

F. Ramade, Eléments D'écologie: Ecologie Fondamentale-4e édition, 2009.

D. S. Reay, F. Dentener, P. Smith, J. Grace, and R. A. Feely, Global nitrogen deposition and carbon sinks, Nature Geoscience, vol.1, pp.430-437, 2008.

M. Reichstein, A. Rey, A. Freibauer, J. Tenhunen, R. Valentini et al., Modeling temporal and large-scale spatial variability of soil respiration from soil water availability, temperature and vegetation productivity indices, Global Biogeochemical Cycles, vol.17, p.1104, 2003.

D. T. Reindl, W. A. Beckman, and J. A. Duffie, Diffuse fraction correlations, Solar Energy, vol.45, pp.1-7, 1990.

M. Réjou-méchain, H. C. Muller-landau, M. Detto, S. C. Thomas, T. Le-toan et al., Local spatial structure of forest biomass and its consequences for remote sensing of carbon stocks, vol.11, pp.6827-6840, 2014.

M. Réjou-méchain, B. Tymen, L. Blanc, S. Fauset, T. R. Feldpausch et al., Using repeated small-footprint LiDAR acquisitions to infer spatial and temporal variations of a high-biomass Neotropical forest, Remote Sensing of Environment, vol.169, pp.93-101, 2015.

J. Remund, L. Wald, M. Lefevre, T. Ranchin, H. Page et al., Worldwide Linke turbidity information, International Solar Energy Society (ISES), 2003.
URL : https://hal.archives-ouvertes.fr/hal-00465791

J. Ross and M. Sulev, Sources of errors in measurements of PAR, Agricultural and Forest Meteorology, vol.100, pp.103-125, 2000.

. R-core-team, R: A Language and Environment for Statistical Computing. R Foundation for Statistical Computing, 2012.

S. S. Saatchi, N. L. Harris, S. Brown, M. Lefsky, E. T. Mitchard et al., Benchmark map of forest carbon stocks in tropical regions across three continents, vol.108, pp.9899-9904, 2011.

D. Sabatier and M. Prévost, Variations du peuplement forestier à l'échelle stationnelle: le cas de la station des Nouragues en Guyane Francaise, 1990.

D. Sabatier and M. F. Prévost, Quelques données sur la composition floristique et la diversité des peuplements forestiers de Guyane française, vol.219, pp.31-55, 1990.

N. Salinas, Y. Malhi, P. Meir, M. Silman, R. Roman-cuesta et al., The sensitivity of tropical leaf litter decomposition to temperature: results from a large-scale leaf translocation experiment along an elevation gradient in Peruvian forests, New Phytologist, vol.189, pp.967-977, 2011.

M. T. Van-der-sande, L. Poorter, S. A. Schnitzer, and L. Markesteijn, Are lianas more drought-tolerant than trees? A test for the role of hydraulic architecture and other stem and leaf traits, Oecologia, vol.172, pp.961-972, 2013.

S. E. Scanga, Population dynamics in canopy gaps: nonlinear response to variable light regimes by an understory plant, Plant Ecology, vol.215, pp.927-935, 2014.

S. A. Schnitzer, A mechanistic explanation for global patterns of liana abundance and distribution, American Naturalist, vol.166, pp.262-276, 2005.

S. A. Schnitzer and F. Bongers, The ecology of lianas and their role in forests, Trends in Ecology & Evolution, vol.17, pp.223-230, 2002.

S. A. Schnitzer and F. Bongers, Increasing liana abundance and biomass in tropical forests: emerging patterns and putative mechanisms, Ecology Letters, vol.14, pp.397-406, 2011.

S. A. Schnitzer and W. P. Carson, Lianas suppress tree regeneration and diversity in treefall gaps, Ecology Letters, vol.13, pp.849-857, 2010.

S. A. Schnitzer, J. W. Dalling, and W. P. Carson, The impact of lianas on tree regeneration in tropical forest canopy gaps: evidence for an alternative pathway of gap-phase regeneration, Journal of Ecology, vol.88, pp.655-666, 2000.

S. A. Schnitzer, S. J. Dewalt, and J. Chave, Censusing and measuring lianas: a quantitative comparison of the common methods, Biotropica, vol.38, pp.581-591, 2006.

S. A. Schnitzer, G. Van-der-heijden, J. Mascaro, and W. P. Carson, Lianas in gaps reduce carbon accumulation in a tropical forest, Ecology, vol.95, pp.3008-3017, 2014.

S. A. Schnitzer, S. A. Mangan, J. W. Dalling, C. A. Baldeck, S. P. Hubbell et al., Liana abundance, diversity, and distribution on, vol.7, p.52114, 2012.

D. M. Schwertfeger and W. H. Hendershot, Determination of effective cation exchange capacity and exchange acidity by a one-step BaCl method, Soil Science Society of America Journal, vol.73, pp.737-743, 2009.

D. Sheil and R. M. May, Mortality and recruitment rate evaluations in heterogeneous tropical forests, Journal of Ecology, vol.84, pp.91-100, 1996.

D. Skole and C. Tucker, Tropical deforestation and habitat fragmentation in the Amazon. Satellite data from 1978 to, Science, vol.260, pp.1905-1910, 1988.

N. S. Skowronski, K. L. Clark, M. Gallagher, R. A. Birdsey, and J. L. Hom, Airborne laser scanner-assisted estimation of aboveground biomass change in a temperate oakpine forest, Remote Sensing of Environment, vol.151, pp.166-174, 2014.

J. W. Slik, V. Arroyo-rodríguez, S. Aiba, P. Alvarez-loayza, L. F. Alves et al., , p.176

M. Santiago, E. L. Catharino, S. Chai, Y. Chen, R. K. Colwell et al., , vol.112, pp.7472-7477

A. P. Smith, K. P. Hogan, and J. R. Idol, Spatial and Temporal Patterns of Light and Canopy Structure in a Lowland Tropical Moist Forest, Biotropica, vol.24, pp.503-511, 1992.

B. Smith, I. C. Prentice, and M. T. Sykes, Representation of vegetation dynamics in the modelling of terrestrial ecosystems: comparing two contrasting approaches within European climate space, Global Ecology and Biogeography, vol.10, pp.621-637, 2001.

P. Sollins, Factors influencing species composition in tropical lowland rain forest: does soil matter?, Ecology, vol.79, pp.23-30, 1998.

T. A. Spies, Forest structure: a key to the ecosystem, Northwest science, vol.72, pp.34-36, 1998.

S. C. Stark, B. J. Enquist, S. R. Saleska, V. Leitold, J. Schietti et al., Linking canopy leaf area and light environments with tree size distributions to explain Amazon forest demography, Ecology Letters, vol.18, pp.636-645, 2015.

S. C. Stark, V. Leitold, J. L. Wu, M. O. Hunter, C. V. De-castilho et al., Amazon forest carbon dynamics predicted by profiles of canopy leaf area and light environment, Ecology Letters, vol.15, pp.1406-1414, 2012.

H. Ter-steege, N. C. Pitman, O. L. Phillips, J. Chave, D. Sabatier et al., Continental-scale patterns of canopy tree composition and function across Amazonia, Nature, vol.443, pp.444-447, 2006.
URL : https://hal.archives-ouvertes.fr/hal-00106584

H. Ter-steege, D. Sabatier, H. Castellanos, T. V. Andel, J. Duivenvoorden et al., An analysis of the floristic composition and diversity of Amazonian forests including those of the Guiana shield, Journal of Tropical Ecology, vol.16, pp.801-828, 2000.

J. C. Stegen, N. G. Swenson, B. J. Enquist, E. P. White, O. L. Phillips et al., Variation in above-ground forest biomass across broad climatic gradients, Global Ecology and Biogeography, vol.20, pp.744-754, 2011.

N. L. Stephenson, P. J. Van-mantgem, A. G. Bunn, H. Bruner, M. E. Harmon et al., Causes and implications of the correlation between forest productivity and tree mortality rates, Ecological Monographs, vol.81, pp.527-555, 2011.

G. C. Stevens, The latitudinal gradient in geographical range: how so many species coexist in the tropics, American naturalist, vol.133, pp.240-256, 1989.

J. Svenning, On the Role of Microenvironmental Heterogeneity in the Ecology and Diversification of Neotropical Rain-Forest Palms (Arecaceae), Botanical Review, vol.67, pp.1-53, 2001.

J. Svenning, D. A. Kinner, R. F. Stallard, B. M. Engelbrecht, and S. J. Wright, Ecological determinism in plant community structure across a tropical forest landscape, Ecology, vol.85, pp.2526-2538, 2004.

S. M. Swinton, G. Escobar, and T. Reardon, Poverty and environment in Latin America: concepts, evidence and policy implications, World Development, vol.31, pp.1865-1872, 2003.

H. Tang, R. Dubayah, A. Swatantran, M. Hofton, S. Sheldon et al., Retrieval of vertical LAI profiles over tropical rain forests using waveform lidar at La Selva, vol.124, pp.242-250, 2012.

P. Taylor, G. Asner, K. Dahlin, C. Anderson, D. Knapp et al., Landscape-scale controls on aboveground forest carbon stocks on the osa peninsula, costa rica, PLoS ONE, vol.10, 2015.

M. Thery and D. Larpin, Seed Dispersal and Vegetation Dynamics at a Cock-of-the-Rock's Lek in the Tropical Forest of French Guiana, Journal of Tropical Ecology, vol.9, pp.109-116, 1993.

J. Terborgh and E. Andresen, The composition of Amazonian forests: patterns at local and regional scales, Journal of Tropical Ecology, vol.14, pp.645-664, 1998.

S. C. Thomas and A. R. Martin, Carbon Content of Tree Tissues: A Synthesis. Forests, vol.3, pp.332-352, 2012.

C. Tinoco-ojanguren and R. W. Pearcy, A Comparison of Light Quality and Quantity Effects on the Growth and Steady-State and Dynamic Photosynthetic Characteristics of Three Tropical Tree Species, Functional Ecology, vol.9, pp.222-230, 1995.

T. Le-toan, S. Quegan, M. W. Davidson, H. Balzter, P. Paillou et al., The BIOMASS mission: Mapping global forest biomass to better understand the terrestrial carbon cycle, Remote Sensing of Environment, vol.115, pp.2850-2860, 2011.
URL : https://hal.archives-ouvertes.fr/hal-00992890

C. Tinoco-ojanguren and R. W. Pearcy, A Comparison of Light Quality and Quantity Effects on the Growth and Steady-State and Dynamic Photosynthetic Characteristics of Three Tropical Tree Species, Functional Ecology, vol.9, pp.222-230, 1995.

K. E. Trenberth, Atmospheric moisture recycling: Role of advection and local evaporation, Journal of Climate, vol.12, pp.1368-1381, 1999.

H. Tuomisto, A. D. Poulsen, K. Ruokolainen, R. C. Moran, C. Quintana et al., Linking floristic patterns with soil heterogeneity and satellite imagery in Ecuadorian Amazonia, Ecological Applications, vol.13, pp.352-371, 2003.

B. L. Turner and T. E. Romero, Short-term changes in extractable inorganic nutrients during storage of tropical rain forest soils, Soil Science Society of America Journal, vol.73, 1972.

G. Vaglio-laurin, Q. Chen, J. A. Lindsell, D. A. Coomes, F. D. Frate et al., Above ground biomass estimation in an African tropical forest with lidar and hyperspectral data, ISPRS Journal of Photogrammetry and Remote Sensing, vol.89, pp.49-58, 2014.

F. Valladares, Light heterogeneity and plants: from ecophysiology to species coexistence and biodiversity, pp.439-471, 2003.

O. Vanderhaeghe, P. Ledru, D. Thiéblemont, E. Egal, A. Cocherie et al., Contrasting mechanism of crustal growth: Geodynamic evolution of the Paleoproterozoic granite-greenstone belts of French Guiana, Precambrian Research, vol.92, pp.165-193, 1998.

P. J. Van-der-meer and F. Bongers, Patterns of tree-fall and branch-fall in a tropical rain forest in french guiana, Journal of Ecology, vol.84, pp.19-29, 1996.

K. T. Vierling, C. Bässler, R. Brandl, L. A. Vierling, I. Weiß et al., Spinning a laser web: predicting spider distributions using LiDAR, Ecological Applications, vol.21, pp.577-588, 2011.

P. Vihervaara, L. Mononen, A. Auvinen, R. Virkkala, Y. Lü et al., How to integrate remotely sensed data and biodiversity for ecosystem assessments at landscape scale -Springer, vol.30, pp.501-516, 2015.

G. Vincent, D. Sabatier, L. Blanc, J. Chave, E. Weissenbacher et al., Accuracy of small footprint airborne LiDAR in its predictions of tropical moist forest stand structure, Remote Sensing of Environment, vol.125, pp.23-33, 2012.
URL : https://hal.archives-ouvertes.fr/hal-02313581

F. Wagner, E. Rutishauser, L. Blanc, and B. Herault, Effects of plot size and census interval on descriptors of forest structure and dynamics, Biotropica, vol.42, pp.664-671, 2010.
URL : https://hal.archives-ouvertes.fr/hal-01093212

W. Wagner, A. Ullrich, V. Ducic, T. Melzer, and N. Studnicka, Gaussian decomposition and calibration of a novel small-footprint full-waveform digitising airborne laser scanner, ISPRS Journal of Photogrammetry and Remote Sensing, vol.60, pp.100-112, 2006.

G. L. Webster, S. P. Churchill, H. Balslev, E. Forero, and J. L. Luteyn, The panorama of Neotropical cloud forests. Biodiversity and conservation of Neotropical montane forests, Proceedings of a symposium, pp.53-77, 1993.

J. F. Weishampel, J. B. Blair, R. G. Knox, R. Dubayah, and D. B. Clark, Volumetric lidar return patterns from an old-growth tropical rainforest canopy, International Journal of Remote Sensing, vol.21, pp.409-415, 2000.

R. H. Whittaker, Communities and Ecosystems, 2nd revised, 1975.

C. G. Willis, C. C. Baskin, J. M. Baskin, J. R. Auld, D. L. Venable et al., The evolution of seed dormancy: environmental cues, evolutionary hubs, and diversification of the seed plants, New Phytologist, vol.203, pp.300-309, 2014.

A. Wolf, P. Ciais, V. Bellassen, N. Delbart, C. B. Field et al., Forest biomass allometry in global land surface models, Global Biogeochemical Cycles, vol.25, p.3015, 2011.
URL : https://hal.archives-ouvertes.fr/hal-01050528

S. J. Wright, O. Calderón, A. Hernandéz, and S. Paton, Are lianas increasing in importance in tropical forests? A 17-year record from Panama, Ecology, vol.85, pp.484-489, 2004.

M. A. Wulder, J. C. White, R. F. Nelson, E. N\a-esset, H. O. Ørka et al., Lidar sampling for large-area forest characterization: A review, Remote Sensing of Environment, vol.121, pp.196-209, 2012.

J. Xia, S. Niu, P. Ciais, I. A. Janssens, J. Chen et al., Joint control of terrestrial gross primary productivity by plant phenology and physiology, Proceedings of the National Academy of Sciences, vol.112, pp.2788-2793, 2015.
URL : https://hal.archives-ouvertes.fr/hal-01268995

A. E. Zanne, D. C. Tank, W. K. Cornwell, J. M. Eastman, S. A. Smith et al., , p.180

N. G. Warman, L. Beaulieu, and J. M. , Three keys to the radiation of angiosperms into freezing environments, Nature, vol.506, pp.89-92, 2014.

K. Zhang, A. D. De-almeida-castanho, D. R. Galbraith, S. Moghim, N. M. Levine et al., The fate of Amazonian ecosystems over the coming century arising from changes in climate, atmospheric CO2, and land use, Global Change Biology, vol.21, pp.2569-2587, 2015.

S. G. Zolkos, S. J. Goetz, and R. Dubayah, A meta-analysis of terrestrial aboveground biomass estimation using lidar remote sensing. Remote Sensing of Environment, vol.128, pp.289-298, 2013.

, Supplementary information

G. P. Asner and J. Mascaro, Mapping tropical forest carbon: Calibrating plot estimates to a simple LiDAR metric. Remote Sensing of Environment, vol.140, pp.614-624, 2014.

G. Asner, J. Mascaro, H. Muller-landau, G. Vieilledent, R. Vaudry et al., A universal airborne LiDAR approach for tropical forest carbon mapping, Oecologia, vol.168, pp.1147-1160, 2012.

G. L. Baskerville, Use of logarithmic regression in the estimation of plant biomass, Canadian Journal of Forest Research, vol.2, pp.49-53, 1972.

J. Böhner, R. Köthe, O. Conrad, J. Gross, A. Ringeler et al., Soil regionalisation by means of terrain analysis and process parameterisation. Soil classification, Research Report, issue.7, pp.213-222, 2001.

A. Brenning, RSAGA: SAGA geoprocessing and terrain analysis in R, 2007.

C. V. De-castilho, W. E. Magnusson, R. N. De-araújo, R. C. Luizão, F. J. Luizão et al., Variation in aboveground tree live biomass in a central Amazonian Forest: Effects of soil and topography, Forest Ecology and Management, vol.234, pp.85-96, 2006.

D. A. Clark, Are tropical forests an important carbon sink? reanalysis of the long-term plot data, Ecological Applications, vol.12, pp.3-7, 2002.

D. B. Clark and D. A. Clark, Landscape variation in forest structure and biomass in a tropical rain forest, Forest Ecology Management, vol.137, pp.185-198, 2000.

M. Detto, H. C. Muller-landau, J. Mascaro, and G. P. Asner, Hydrological Networks and Associated Topographic Variation as Templates for the Spatial Organization of Tropical Forest Vegetation, PLoS ONE, vol.8, p.76296, 2013.

T. Emilio, C. A. Quesada, F. R. Costa, W. E. Magnusson, J. Schietti et al., Soil physical conditions limit palm and tree basal area in Amazonian forests, Plant Ecology & Diversity, vol.0, pp.1-15, 2013.

B. M. Engelbrecht, L. S. Comita, R. Condit, T. A. Kursar, M. T. Tyree et al., Drought sensitivity shapes species distribution patterns in tropical forests, Nature, vol.447, pp.80-82, 2007.

B. Ferry, F. Morneau, J. Bontemps, L. Blanc, and V. Freycon, Higher treefall rates on slopes and waterlogged soils result in lower stand biomass and productivity in a tropical rain forest, Journal of Ecology, vol.98, pp.106-116, 2010.
URL : https://hal.archives-ouvertes.fr/hal-01032100

K. Harms, R. Condit, S. P. Hubbel, and R. B. Foster, Habitat associations of trees and shrubs in a 50-ha neotropical forest plot, Journal of Ecology, vol.89, pp.947-959, 2001.

W. F. Laurance, P. M. Fearnsideb, S. G. Laurancea, P. Delamonicaa, T. E. Lovejoyc et al., Relationship between soils and Amazon forest biomass: a landscape-scale study, Forest Ecology and Management, vol.118, pp.127-138, 1999.

G. Lopez-gonzalez, S. L. Lewis, M. Burkitt, and O. L. Phillips, ForestPlots. net: a web application and research tool to manage and analyse tropical forest plot data, Journal of Vegetation Science, vol.22, pp.610-613, 2011.

J. Mascaro, G. P. Asner, H. C. Muller-landau, M. Van-breugel, J. Hall et al., Controls over aboveground forest carbon density on, vol.8, pp.1615-1629, 2011.

R. W. Mcewan, Y. Lin, I. Sun, C. Hsieh, S. Su et al., Topographic and biotic regulation of aboveground carbon storage in subtropical broad-leaved forests of Taiwan, Forest Ecology and Management, vol.262, pp.1817-1825, 2011.

D. L. Miranda, . De, C. R. Sanquetta, L. G. Costa, S. Da et al., Biomassa e carbono em Euterpe oleracea Mart. na ilha do Marajó -PA, vol.19, pp.336-343, 2012.

L. Wang and H. Liu, An efficient method for identifying and filling surface depressions in digital elevation models for hydrologic analysis and modelling, International Journal of Geographical Information Science, vol.20, pp.193-213, 2006.

L. W. Zevenbergen and C. R. Thorne, Quantitative analysis of land surface topography, Earth Surface Processes and Landforms, vol.12, pp.47-56, 1987.

, the Supporting Information

D. B. Clark and D. A. Clark, Distribution and effects on tree growth of lianas and woody hemiepiphytes in a Costa Rican tropical wet forest, Journal of Tropical Ecology, vol.6, pp.321-331, 1990.

L. L. Ingwell, J. Wright, S. Becklund, K. K. Hubbell, S. P. Schnitzer et al., The impact of lianas on 10 years of tree growth and mortality on, vol.98, pp.879-887, 2010.

R. J. Mcgaughey, FUSION/LDV: Software for LIDAR data analysis and visualization. US Department of Agriculture, p.123, 2009.

O. L. Phillips, R. Vásquez-martínez, A. Monteagudo-mendoza, T. R. Baker, and P. Vargas, Large lianas as hyperdynamic elements of the tropical forest canopy, Ecology, vol.86, pp.1250-1258, 2005.

S. A. Schnitzer, M. E. Kuzee, and F. Bongers, Disentangling above-and below-ground competition between lianas and trees in a tropical forest, Journal of Ecology, vol.93, pp.1115-1125, 2005.

G. Vincent, D. Sabatier, L. Blanc, J. Chave, E. Weissenbacher et al., Accuracy of small footprint airborne LiDAR in its predictions of tropical moist forest stand structure, Remote Sensing of Environment, vol.125, pp.23-33, 2012.
URL : https://hal.archives-ouvertes.fr/hal-02313581

N. V. Brokaw, The Definition of Treefall Gap and Its Effect on Measures of Forest Dynamics, Biotropica, vol.14, pp.158-160, 1982.