, ) en respectant les abréviations des titres des journaux selon les instructions mentionnées dans le site Index Medicus, vol.4094

, Références Bibliographiques

A. M. Abdel-mawgoud, M. M. Aboulwafa, and N. A. Hassouna, Characterization of Surfactin Produced by Bacillus subtilis Isolate BS5, Applied Biochemistry and Biotechnology, vol.150, issue.3, pp.289-303, 2008.

A. M. Abdel-mawgoud, F. Lépine, and E. Déziel, Rhamnolipids: diversity of structures, microbial origins and roles, Applied Microbiology and Biotechnology, vol.86, issue.5, pp.1323-1336, 2010.
URL : https://hal.archives-ouvertes.fr/pasteur-00819624

A. S. Abouhend and K. M. El-moselhy, Spatial and seasonal variations of heavy metals in water and sediments at the Northern Red Sea Coast, American Journal of Water Resources, vol.3, issue.3, pp.73-85, 2015.

M. Abouseoud, A. Yataghene, A. Amrane, and R. Maachi, Biosurfactant production by free and alginate entrapped cells of Pseudomonas fluorescens, Journal of industrial microbiology & biotechnology, vol.35, issue.11, pp.1303-1308, 2008.

A. Acosta-gonzález and S. Marqués, Bacterial diversity in oil-polluted marine coastal sediments, Current Opinion in Biotechnology, vol.38, pp.24-32, 2016.

T. A. Adedosu, H. O. Adedosu, O. S. Sojinu, and A. A. Olajire, N-Alkanes and polycyclic aromatic hydrocarbons (PAHs) profile of soil from some polluted sites in Niger Delta, Nigeria. Environmental Earth Sciences, vol.68, issue.8, pp.2139-2144, 2013.

A. O. Adeniji, O. O. Okoh, and A. I. Okoh, Petroleum Hydrocarbon Profiles of Water and Sediment of Algoa Bay, International Journal of Environmental Research and Public Health, vol.14, issue.10, p.1263, 2017.

J. Aislabie, M. Mcleod, and R. Fraser, Potential for biodegradation of hydrocarbons in soil from the Ross Dependency, Antarctica. Applied Microbiology and Biotechnology, vol.49, issue.2, pp.210-214, 1998.

F. Al-abdali, M. Massoud, and A. Al-ghadban, Bottom sediments of the Arabian Gulf-III. Trace metal contents as indicators of pollution and implications for the effect and fate of the Kuwait oil slick, Environmental Pollution, vol.93, issue.3, pp.285-301, 1996.

D. M. Al-mailem, N. A. Sorkhoh, H. Al-awadhi, M. Eliyas, and S. S. Radwan, Biodegradation of crude oil and pure hydrocarbons by extreme halophilic archaea from hypersaline coasts of the Arabian Gulf, Extremophiles, vol.14, issue.3, pp.321-328, 2010.

T. Al-said, S. Naqvi, F. Al-yamani, A. Goncharov, and L. Fernandes, High total organic carbon in surface waters of the northern Arabian Gulf: Implications for the oxygen minimum zone of the Arabian Sea, Marine pollution bulletin, vol.129, issue.1, pp.35-42, 2018.

H. R. Ali, N. S. El-gendy, Y. M. Moustafa, M. I. Roushdy, and A. I. Hashem, Degradation of Asphaltenic Fraction by Locally Isolated Halotolerant Bacterial Strains, ISRN Soil Science, p.11, 2012.

A. Khan, A. H. Tanveer, S. Alia, S. Anees, M. Sultan et al., Role of nutrients in bacterial biosurfactant production and effect of biosurfactant production on petroleum hydrocarbon biodegradation, Ecological Engineering, vol.104, pp.158-164, 2017.

F. Aloulou, M. Kallel, M. Dammak, B. Elleuch, and A. Saliot, Even-numbered n-alkanes/nalkenes predominance in surface sediments of Gabes Gulf in Tunisia, Environmental Earth Sciences, vol.61, issue.1, pp.1-10, 2010.

R. I. Amann, W. Ludwig, and K. Schleifer, Phylogenetic identification and in situ detection of individual microbial cells without cultivation, Microbiological reviews, vol.59, issue.1, pp.143-169, 1995.

C. Amiard-triquet, T. Burgeot, and D. Claisse, La surveillance de la qualité du milieu marin: l'expérience du Réseau national d'observation (RNO) et le développement de biomarqueurs, Océanis, vol.25, issue.4, pp.651-684, 1999.

J. W. Anderson and R. F. Lee, Use of biomarkers in oil spill risk assessment in the marine environment, Human and Ecological Risk Assessment, vol.12, issue.6, pp.1192-1222, 2006.

F. Anjum, G. Gautam, G. Edgard, and S. Negi, Biosurfactant production through Bacillus sp. MTCC 5877 and its multifarious applications in food industry, Bioresource Technology, vol.213, pp.262-269, 2016.

, Agence nationale de protection de l'environnement 2012. Manuel de procédures du pniu pour lutter contre les évènements de pollution marine, ANPE, pp.30-59

A. Aparna, G. Srinikethan, and H. Smitha, Production and characterization of biosurfactant produced by a novel Pseudomonas sp. 2B, Colloids and Surfaces B: Biointerfaces, vol.95, pp.23-29, 2012.

T. Austad and K. Taugbøl, Chemical flooding of oil reservoirs 2. Dissociative surfactant-polymer interaction with a negative effect on oil recovery, Colloids and Surfaces A: Physicochemical and Engineering Aspects, vol.103, issue.1-2, pp.73-81, 1995.

J. Avoine, L'estuaire de la Seine : sédiments et dynamique sédimentaire, p.236, 1981.

C. C. Azubuike, C. B. Chikere, and G. C. Okpokwasili, Bioremediation techniques-classification based on site of application: principles, advantages, limitations and prospects, World Journal of Microbiology and Biotechnology, vol.32, issue.11, p.180, 2016.

A. Bahuguna, M. K. Lily, K. Dangwal, A. Munjal, and R. N. Singh, Degradation of Naphthalene by a novel strain Bacillus licheniformis BMIT5ii (MTCC 9446), Journal of Pharmacy Research, vol.5, issue.3, pp.1600-1604, 2012.

B. R. Baldwin, C. H. Nakatsu, and L. Nies, Detection and enumeration of aromatic oxygenase genes by multiplex and real-time PCR, Applied and Environmental Microbiology, vol.69, issue.6, pp.3350-3358, 2003.

I. M. Banat, M. A. De-rienzo, and G. A. Quinn, Microbial biofilms: biosurfactants as antibiofilm agents, Applied microbiology and biotechnology, vol.98, issue.24, pp.9915-9929, 2014.

I. M. Banat, R. S. Makkar, and S. S. Cameotra, Potential commercial applications of microbial surfactants, Applied microbiology and biotechnology, vol.53, issue.5, pp.495-508, 2000.

I. M. Banat, S. K. Satpute, S. S. Cameotra, R. Patil, and N. V. Nyayanit, Cost effective technologies and renewable substrates for biosurfactants' production, Frontiers in microbiology, vol.5, p.697, 2014.

F. F. Barros, A. N. Ponezi, and G. M. Pastore, Production of biosurfactant by Bacillus subtilis LB5a on a pilot scale using cassava wastewater as substrate, Journal of Industrial Microbiology & Biotechnology, vol.35, issue.9, pp.1071-1078, 2008.

S. Bayen, Occurrence, bioavailability and toxic effects of trace metals and organic contaminants in mangrove ecosystems: A review, Environmental International, vol.48, issue.1, pp.84-101, 2012.

H. Ben-ayed, S. Bardaa, D. Moalla, M. Jridi, H. Maalej et al., Wound healing and in vitro antioxidant activities of lipopeptides mixture produced by Bacillus mojavensis A21, Process Biochemistry, vol.50, issue.6, pp.1023-1030, 2015.

H. Ben-ayed and M. Nasri, Les lipopeptides de Bacillus: des molécules bioactives à structure particulière et à forte potentialité d'application, Nutrition et Santé, vol.5, issue.2, pp.115-125, 2016.

J. Bertrand, M. Bianchi, A. Mallah, M. Acquaviva, M. Mille et al., Hydrocarbon biodegradation and hydrocarbonoclastic bacterial communities composition grown in seawater as a function of sodium chloride concentration, Journal of experimental marine biology and ecology, vol.168, issue.1, pp.125-138, 1993.

J. C. Bertrand, P. Caumette, G. Mille, M. Gilewicz, and M. Denis, Anaerobic biodegradation of hydrocarbons, Science Progress, vol.73, issue.3, pp.333-350, 1933.

J. Bertrand and G. Mille, Devenir de la matière organique exogène, Un modèle: les hydrocarbures. Microorganismes dans les écosystèmes océaniques. Masson (Paris), vol.13, pp.343-385, 1989.

F. A. Bezza and E. M. Nkhalambayausi-chirwa, Biosurfactant-enhanced bioremediation of aged polycyclic aromatic hydrocarbons (PAHs) in creosote contaminated soil, Chemosphere, vol.144, pp.635-644, 2016.

J. Boateng and O. Catanzano, Advanced therapeutic dressings for effective wound healing-a review, Journal of pharmaceutical sciences, vol.104, issue.11, pp.3653-3680, 2015.

J. S. Boateng, K. H. Matthews, H. N. Stevens, and G. M. Eccleston, Wound healing dressings and drug delivery systems: a review, Journal of pharmaceutical sciences, vol.97, issue.8, pp.2892-2923, 2008.

A. A. Bodour, K. P. Drees, and R. M. Maier, Distribution of biosurfactant-producing bacteria in undisturbed and contaminated arid southwestern soils, Applied and environmental microbiology, vol.69, issue.6, pp.3280-3287, 2003.

D. Borah and R. Yadav, Biodegradation of diesel, crude oil, kerosene and used engine oil by a newly isolated Bacillus cereus strain DRDU1 from an automobile engine in liquid culture, Arabian Journal for Science and Engineering, vol.39, issue.7, pp.5337-5345, 2014.

M. Bouchez-naïtali, H. Rakatozafy, R. Marchal, J. Y. Leveau, and J. P. Vandecasteele, Diversity of bacterial strains degrading hexadecane in relation to the mode of substrate uptake, Journal of Applied Microbiology, vol.86, issue.3, pp.421-428, 1999.

M. Bouchez, D. Blanchet, and J. Vandecasteele, Degradation of polycyclic aromatic hydrocarbons by pure strains and by defined strain associations: inhibition phenomena and cometabolism, Applied Microbiology and Biotechnology, vol.43, issue.1, pp.156-164, 1995.

F. R. Branquinho, Unveiling population diversity, biosurfactant and antibacterial agents production of Bacillus pumilis group species. Thesis, Faculty of Pharmacy of the University of Porto, Chapter, pp.351-389, 2014.

F. Brian-jaisson, Identification et caractérisation des exopolymères de biofilms de bactéries marines, 2014.

A. Bridier, P. Sanchez-vizuete, M. Guilbaud, J. Piard, M. Naïtali et al., Biofilmassociated persistence of food-borne pathogens, Food microbiology, vol.45, pp.167-178, 2015.
URL : https://hal.archives-ouvertes.fr/hal-01204463

B. Callaghan, H. Lydon, S. L. Roelants, I. N. Van-bogaert, R. Marchant et al., Lactonic Sophorolipids increase tumor burden in Apcmin+/-mice, PloS one, vol.11, issue.6, p.156845, 2016.

S. S. Cameotra and R. S. Makkar, Recent applications of biosurfactants as biological and immunological molecules, Current Opinion in Microbiology, vol.7, issue.3, pp.262-266, 2004.

S. S. Cameotra and R. S. Makkar, Synthesis of biosurfactants in extreme conditions, Applied Microbiology and Biotechnology, vol.50, issue.5, pp.520-529, 1998.

J. G. Caporaso, J. Kuczynski, J. Stombaugh, K. Bittinger, F. D. Bushman et al., QIIME allows analysis of high-throughput community sequencing data, Nature methods, vol.7, issue.5, p.335, 2010.

M. A. Castillo, I. S. Trujillo, E. V. Alonso, A. G. De-torres, and J. C. Pavón, Bioavailability of heavy metals in water and sediments from a typical Mediterranean Bay, Marine pollution bulletin, vol.76, pp.427-434, 2013.

M. L. Cazalet, Caractérisation physico-chimique d'un sédiment marin traité aux liants hydrauliques: Évaluation de la mobilité potentielle des polluants inorganiques, 2012.

A. Cébron, M. P. Norini, T. Beguiristain, and C. Leyval, Real-Time PCR quantification of PAHring hydroxylating dioxygenase (PAH-RHD?) genes from Gram positive and Gram negative Bacteria in soil and sediment samples, Journal of Microbiological Methods, vol.73, pp.148-159, 2008.

C. E. Cerniglia, Biodegradation of polycyclic aromatic hydrocarbons, Current Opinion in Biotechnology, vol.4, issue.3, pp.331-338, 1993.

C. E. Cerniglia, J. B. Sutherland, and . Timmis, Degradation of Polycyclic Aromatic Hydrocarbons by Fungi, Handbook of Hydrocarbon and Lipid Microbiology, pp.2079-2110, 2010.

M. Chachkhiani, P. Dabert, T. Abzianidze, G. Partskhaladze, L. Tsiklauri et al., 16S rDNA characterisation of bacterial and archaeal communities during start-up of anaerobic thermophilic digestion of cattle manure, Bioresource Technology, vol.93, issue.3, pp.227-232, 2004.

S. K. Chaerun, K. Tazaki, R. Asada, and K. Kogure, Bioremediation of coastal areas 5 years after the Nakhodka oil spill in the Sea of Japan: isolation and characterization of hydrocarbondegrading bacteria, Environment International, vol.30, issue.7, pp.911-922, 2004.

S. Chakraborty and G. Owens, Metal distributions in seawater, sediment and marine benthic macroalgae from the South Australian coastline, International Journal of Environmental Science and Technology, vol.11, issue.5, pp.1259-1270, 2014.

H. Chalghmi, Etude de la pollution marine par les hydrocarbures et caractérisation de leurs effets biochimiques et moléculaires sur la palourde de Ruditapes sp, 2015.

M. Chamkha, S. Sayadi, V. Bru, and J. Godon, Microbial diversity in Tunisian olive fermentation brine as evaluated by small subunit rRNA-Single strand conformation polymorphism analysis, International journal of food microbiology, vol.122, issue.1-2, pp.211-215, 2008.

A. Chaouay, R. Okhrib, M. Hilali, L. Bazzi, A. Chahid et al., Contribution à l'étude de l'analyse physico-chimique et de la contamination métallique de l'eau de mer du littoral d'Agadir (Sud du Maroc).(Contribution to the study of physico-chemical analysis and metal contamination of coastal Agadir seawater

A. Charpiot, La méthanisation dans la filière laitière. Compte rendu-n° 001233011. Département Techniques d'élevage et, Qualité. Service Bâtiment-Environnement, 2012.

A. Chebbi, M. Elshikh, F. Haque, S. Ahmed, S. Dobbin et al., Rhamnolipids from Pseudomonas aeruginosa strain W10; as antibiofilm/antibiofouling products for metal protection, Journal of basic microbiology, vol.57, issue.5, pp.364-375, 2017.

, Références Bibliographiques

A. Chebbi, D. Hentati, M. Cheffi, R. Bouabdallah, C. Choura et al., Promising abilities of mercapto-degrading Staphylococcus capitis strain SH6 in both crude oil and waste motor oil as sole carbon and energy sources: its biosurfactant production and preliminary characterization, Journal of Chemical Technology & Biotechnology, vol.93, issue.5, pp.1401-1412, 2018.

A. Chebbi, D. Hentati, H. Zaghden, N. Baccar, F. Rezgui et al., Polycyclic aromatic hydrocarbon degradation and biosurfactant production by a newly isolated Pseudomonas sp. strain from used motor oil-contaminated soil, International Biodeterioration & Biodegradation, vol.122, pp.128-140, 2017.

J. Chen, P. Huang, K. Zhang, and F. Ding, Isolation of biosurfactant producers, optimization and properties of biosurfactant produced by Acinetobacter sp. from petroleum-contaminated soil, Journal of applied microbiology, vol.112, issue.4, pp.660-671, 2012.

Q. Chen, J. Li, M. Liu, H. Sun, and M. Bao, Study on the biodegradation of crude oil by free and immobilized bacterial consortium in marine environment, PloS one, vol.12, issue.3, p.174445, 2017.

Q. Chen, J. Liu, F. Liu, B. Wang, and Z. Cao, Biologic risk and source diagnose of 16 PAHs from Haihe River Basin, China. Frontiers of Environmental Science & Engineering, vol.10, issue.1, pp.46-52, 2016.

C. Chiellini, R. Iannelli, F. Verni, and G. Etroni, Bacterial communities in polluted seabed sediments: a molecular biology assay in Leghorn Harbor, Scientific World Journal, p.165706, 2013.

H. Chong and Q. Li, Microbial production of rhamnolipids: opportunities, challenges and strategies. Microbial cell factories, vol.16, p.137, 2017.
DOI : 10.1186/s12934-017-0753-2

URL : https://doi.org/10.1186/s12934-017-0753-2

K. Chong and A. Alexander, Early detection and management of ganoderma basal stem rot disease: a special report from Sabah, Transactions on Science and Technology, vol.3, issue.3, pp.517-523, 2016.

C. S. Chooklin, S. Maneerat, and A. Saimmai, Utilization of Banana Peel as a Novel Substrate for Biosurfactant Production by Halobacteriaceae archaeon AS65, Applied Biochemistry and Biotechnology, vol.173, issue.2, pp.624-645, 2014.

R. Christen, Global sequencing: a review of current molecular data and new methods available to assess microbial diversity, Microbes and Environments, vol.23, issue.4, pp.253-268, 2008.

M. Christopher, Logistics & supply chain management, 2016.

N. Christova, B. Tuleva, Z. Lalchev, A. Jordanova, and B. Jordanov, Rhamnolipid biosurfactants produced by Renibacterium salmoninarum 27BN during growth on n-hexadecane, Zeitschrift für Naturforschung C, vol.59, issue.1-2, pp.70-74, 2004.
DOI : 10.1515/znc-2004-1-215

URL : http://www.degruyter.com/downloadpdf/j/znc.2004.59.issue-1-2/znc-2004-1-215/znc-2004-1-215.xml

O. Chuan, Y. Chuen, and F. Mei, Determination of total organic carbon concentration in surficial sediments of Sungai Pinang, Penang, Malaysia [Penentuan jumlah karbon organik dalam sedimen permukaan di Sungai Pinang, 2016.

C. Ciacci, C. Barmo, G. Gallo, M. Maisano, T. Cappello et al., Effects of sublethal, environmentally relevant concentrations of hexavalent chromium in the gills of Mytilus galloprovincialis, Aquatic Toxicology, vol.120, pp.109-118, 2012.

M. C. Cirigliano and G. M. Carma, Isolation of a Bioemulsifier from Candida lipolyticat, Applied and environmental microbiology, vol.48, issue.4, pp.747-750, 1984.

L. L. Clark, J. J. Dajcs, C. H. Mclean, J. G. Bartell, and D. W. Stroman, Pseudomonas otitidis sp. nov., isolated from patients with otic infections, International Journal of Systematic and Evolutionary Microbiology, vol.56, issue.4, pp.709-714, 2006.
DOI : 10.1099/ijs.0.63753-0

URL : http://ijs.microbiologyresearch.org/deliver/fulltext/ijsem/56/4/709.pdf?itemId=/content/journal/ijsem/10.1099/ijs.0.63753-0&mimeType=pdf&isFastTrackArticle=

S. A. Cochrane and J. C. Vederas, Lipopeptides from Bacillus and Paenibacillus spp.: a gold mine of antibiotic candidates, Medicinal Research Reviews, vol.36, issue.1, pp.4-31, 2016.
DOI : 10.1002/med.21321

K. A. Colati, G. P. Dalmaschio, E. V. De-castro, A. O. Gomes, B. G. Vaz et al., Monitoring the liquid/liquid extraction of naphthenic acids in brazilian crude oil using electrospray ionization FT-ICR mass spectrometry (ESI FT-ICR MS), Fuel, vol.108, pp.647-655, 2013.

J. C. Colombo, M. Cabello, and A. M. Arambarri, Biodegradation of aliphatic and aromatic hydrocarbons by natural soil microflora and pure cultures of imperfect and lignolitic fungi, Environmental pollution, vol.94, issue.3, pp.355-362, 1996.

J. Coronel-león, G. De-grau, A. Grau-campistany, M. Farfan, F. Rabanal et al., Biosurfactant production by AL 1.1, a Bacillus licheniformis strain isolated from Antarctica: production, chemical characterization and properties, Annals of Microbiology, vol.65, issue.4, pp.2065-2078, 2015.

J. Coronel-león, A. Marqués, J. Bastida, and A. Manresa, Optimizing the production of the biosurfactant lichenysin and its application in biofilm control, Journal of applied microbiology, vol.120, issue.1, pp.99-111, 2016.

J. Cortés-sánchez-ade, H. Hernández-sánchez, and M. E. Jaramillo-flores, Biological activity of glycolipids produced by microorganisms: New trends and possible therapeutic alternatives, Microbiological Research, vol.168, issue.1, pp.22-32, 2013.

S. Costa, E. Déziel, and F. Lépine, Characterization of rhamnolipid production by Burkholderia glumae, Letters in applied microbiology, vol.53, issue.6, pp.620-627, 2011.
URL : https://hal.archives-ouvertes.fr/pasteur-00722219

C. M. Couillard, K. Lee, B. Légaré, and T. L. King, Effect of dispersant on the composition of the water-accommodated fraction of crude oil and its toxicity to larval marine fish, Environmental Toxicology and Chemestry, vol.24, issue.6, pp.1496-504, 2005.

C. D. Da-cunha, A. S. Rosado, G. V. Sebastián, and L. Seldin, Von der Weid, I. 2006. Oil biodegradation by Bacillus strains isolated from the rock of an oil reservoir located in a deep-water production basin in Brazil, Applied microbiology and biotechnology, vol.73, issue.4, pp.949-959

G. Dalmaso, D. Ferreira, and A. Vermelho, Marine extremophiles: a source of hydrolases for biotechnological applications, Marine drugs, vol.13, issue.4, pp.1925-1965, 2015.

N. Das and P. Chandran, Microbial Degradation of Petroleum Hydrocarbon Contaminants: An Overview, Biotechnology Research International, 2011.

P. Das, X. Yang, and L. Z. Ma, Analysis of biosurfactants from industrially viable Pseudomonas strain isolated from crude oil suggests how rhamnolipids congeners affect emulsification property and antimicrobial activity, Frontiers in Microbiology, vol.5, issue.696, 2014.

D. G. De-almeida, . Soares-da, R. D. Silva, J. M. Luna, R. D. Rufino et al., Biosurfactants: Promising Molecules for Petroleum Biotechnology Advances, Frontiers in Microbiology, vol.7, 1718.

C. J. De-andrade, F. F. Barros, L. M. De-andrade, S. A. Rocco, L. Sforça et al., Ultrafiltration based purification strategies for surfactin produced by Bacillus subtilis LB5A using cassava wastewater as substrate, Journal of Chemical Technology & Biotechnology, vol.91, issue.12, pp.3018-3027, 2016.

Í. W. De-frança, A. P. Lima, J. A. Lemos, C. G. Lemos, V. M. Melo et al., Production of a biosurfactant by Bacillus subtilis ICA56 aiming bioremediation of impacted soils, Catalysis Today, vol.255, pp.10-15, 2015.

M. R. De-oliveira, A. Magri, C. Baldo, D. Camilios-neto, T. Minucelli et al., Sophorolipids A promising biosurfactant and it's applications, Int J Adv Biotechnol Res, vol.6, pp.161-174, 2015.

E. F. Delong, Archaea in coastal marine environments, Proceedings of the National Academy of Sciences, vol.89, issue.12, pp.5685-5689, 1992.

G. R. Denton, B. Bearden, L. Concepcion, H. Siegrist, D. Vann et al., Contaminant assessment of surface sediments from Tanapag Lagoon, Saipan, 2001.

J. D. Desai and I. M. Banat, Microbial production of surfactants and their commercial potential, Microbiology and Molecular Biology Reviews, vol.61, issue.1, pp.47-64, 1997.

M. A. Díaz-de-rienzo, I. M. Banat, B. Dolman, J. Winterburn, and P. J. Martin, Sophorolipid biosurfactants: Possible uses as antibacterial and antibiofilm agent, New Biotechnology, vol.32, issue.6, pp.720-726, 2015.

N. Dijkstra, J. Junttila, K. Skirbekk, J. Carroll, K. Husumd et al., Benthic foraminifera as bio-indicators of chemical and physical stressors in Hammerfest harbor, 2017.

, Marine Pollution Bulletin, vol.114, issue.1, pp.384-396

G. Ding, H. Heuer, S. Zühlke, M. Spiteller, G. J. Pronk et al., Soil type-dependent responses to phenanthrene as revealed by determining the diversity and abundance of polycyclic aromatic hydrocarbon ring-hydroxylating dioxygenase genes by using a novel PCR detection system, Applied and environmental microbiology, vol.76, issue.14, pp.4765-4771, 2010.

S. Dooley, J. Heyne, S. H. Won, P. Dievart, Y. Ju et al., Importance of a Cycloalkane Functionality in the Oxidation of a Real Fuel, Energy & Fuels, vol.28, issue.12, pp.7649-7661, 2014.

Z. Drira, S. Kmiha-megdiche, H. Sahnoun, A. Hammami, N. Allouche et al., Assessment of anthropogenic inputs in the surface waters of the southern coastal area of Sfax during spring, Marine pollution bulletin, vol.104, pp.355-363, 2016.
URL : https://hal.archives-ouvertes.fr/hal-01435961

W. A. Ducker and L. M. Grant, Effect of Substrate Hydrophobicity on Surfactant Surface?Aggregate Geometry, The Journal of Physical Chemistry, vol.100, issue.28, pp.11507-11511, 1996.

K. Durairaj, P. Velmurugan, J. Park, W. Chang, Y. Park et al., Characterization and assessment of two biocontrol bacteria against Pseudomonas syringae wilt in Solanum lycopersicum and its genetic responses, Microbiological Research, vol.206, pp.43-49, 2018.

R. Duran and C. Cravo-laureau, Role of environmental factors and microorganisms in determining the fate of polycyclic aromatic hydrocarbons in the marine environment, FEMS Microbiology Reviews, vol.40, issue.6, pp.814-830, 2016.
URL : https://hal.archives-ouvertes.fr/hal-01495414

K. Dutta, S. Shityakov, P. P. Das, and C. Ghosh, Enhanced biodegradation of mixed PAHs by mutated naphthalene 1, 2-dioxygenase encoded by Pseudomonas putida strain KD6 isolated from petroleum refinery waste, Biotech, vol.7, issue.3, p.365, 2017.

K. Eddouaouda, S. Mnif, A. Badis, S. B. Younes, S. Cherif et al., Characterization of a novel biosurfactant produced by Staphylococcus sp. strain 1E with potential application on hydrocarbon bioremediation, Journal of basic microbiology, vol.52, issue.4, pp.408-418, 2012.

R. Eganhouse and P. Sherblom, Anthropogenic organic contaminants in the effluent of a combined sewer overflow: impact on Boston Harbor, Marine Environmental Research, vol.51, issue.1, pp.51-74, 2001.

M. Elshikh, S. Ahmed, S. Funston, P. Dunlop, M. Mcgaw et al., Resazurin-based 96-well plate microdilution method for the determination of minimum inhibitory concentration of biosurfactants, Biotechnology Letters, vol.38, issue.6, pp.1015-1019, 2016.

H. Er-raioui, S. Bouzid, M. Marhraoui, and A. Saliot, Hydrocarbon pollution of the Mediterranean coastline of Morocco, Ocean & Coastal Management, vol.52, issue.2, pp.124-129, 2009.

S. Eskandari, M. Hoodaji, A. Tahmourespour, A. Abdollahi, T. Mohammadian-baghi et al., Bioremediation of Polycyclic Aromatic Hydrocarbons by Bacillus Licheniformis ATHE9 and Bacillus Mojavensis ATHE13 as Newly Strains Isolated from Oil-Contaminated Soil, Environment and Earth Science International, vol.11, issue.2, pp.1-11, 2017.

X. Fang, C. Thornton, B. E. Scheffler, and K. L. Willett, Benzo [a] pyrene decreases global and gene specific DNA methylation during zebrafish development, Environmental toxicology and pharmacology, vol.36, issue.1, pp.40-50, 2013.

H. Faragallah, A. Askar, M. Okbah, and H. Moustafa, Physico-chemical characteristics of the open Mediterranean sea water far about 60 Km from Damietta harbor, Egypt. Journal of Ecology and the Natural Environment, vol.1, issue.5, pp.106-119, 2009.

B. Z. Fathepure, Recent studies in microbial degradation of petroleum hydrocarbons in hypersaline environments, Frontiers in Microbiology, vol.5, issue.173, 2014.

B. W. Feng, X. R. Li, J. H. Wang, Z. Y. Hu, H. Meng et al., Bacterial diversity of water and sediment in the Changjiang estuary and coastal area of the East China Sea, FEMS Microbiology Ecology, vol.70, issue.2, pp.236-248, 2009.

A. Filibeli, R. Yilmaz, and I. Alyanak, Dredged material of Izmir Harbor: its behavior and pollution potential, Water Science and Technology, vol.32, issue.2, pp.105-113, 1995.

H. Flemming, Microbial biofouling: unsolved problems, insufficient approaches, and possible solutions, Biofilm Highlights, pp.81-109, 2011.

M. Fondi, V. Orlandini, G. Emiliani, M. C. Papaleo, I. Maida et al., Draft Genome Sequence of the Hydrocarbon-Degrading and Emulsan-Producing Strain Acinetobacter venetianus RAG-1T, Journal of Bacteriology, vol.194, issue.17, pp.4771-4772, 2012.

G. E. Fox, K. R. Pechman, and C. R. Woese, Comparative cataloging of 16S ribosomal ribonucleic acid: molecular approach to procaryotic systematics, International Journal of Systematic and Evolutionary Microbiology, vol.27, issue.1, pp.44-57, 1977.

J. Gans, M. Wolinsky, and J. Dunbar, Computational improvements reveal great bacterial diversity and high metal toxicity in soil, Science, vol.309, issue.5739, pp.1387-1390, 2005.

D. Gargouri, C. Azri, M. M. Serbaji, Y. Jedoui, and M. Montacer, Heavy metal concentrations in the surface marine sediments of Sfax Coast, Tunisia. Environmental Monitoring and Assessment, vol.175, issue.1-4, pp.519-530, 2011.

F. Gassara, N. Suri, and G. Voordouw, Nitrate-mediated microbially enhanced oil recovery (NMEOR) from model upflow bioreactors, Journal of hazardous materials, vol.324, pp.94-99, 2017.
DOI : 10.1016/j.jhazmat.2015.12.039

M. J. Gauthier, B. Lafay, R. Christen, L. Fernandez, M. Acquaviva et al., Marinobacter hydrocarbonoclasticus gen. nov., sp. nov., a New, Extremely Halotolerant, Hydrocarbon-Degrading Marine Bacterium, International Journal of Systematic and Evolutionary Microbiology, vol.42, issue.4, pp.568-576, 1992.
URL : https://hal.archives-ouvertes.fr/hal-00412938

S. Geetha, I. M. Banat, and S. J. Joshi, Biosurfactants: Production and potential applications in microbial enhanced oil recovery (MEOR), Biocatalysis and Agricultural Biotechnology, 2018.

S. George and K. Jayachandran, Production and characterization of rhamnolipid biosurfactant from waste frying coconut oil using a novel Pseudomonas aeruginosa D, Journal of Applied Microbiology, vol.114, issue.2, pp.373-383, 2013.

Y. Gerchman, R. Patichov, and T. Zeltzer, Lipolytic, Proteolytic, and Cholesterol-Degrading Bacteria from the Human Cerumen, Current Microbiology, vol.64, issue.6, pp.588-591, 2012.
DOI : 10.1007/s00284-012-0113-8

, United nation group of expert on the scientific aspect of marine pollution. Article premier, GESAMP, vol.1, 1989.

I. M. Gillespie and J. C. Philp, Bioremediation, an environmental remediation technology for the bioeconomy, Trends in Biotechnology, vol.31, issue.6, pp.329-332, 2013.
DOI : 10.1016/j.tibtech.2013.01.015

, Global Market Insights, 2018.

J. Godon, F. Duthoit, C. Delbes, L. Millet, and M. Montel, Use of molecular fingerprint for the study of complex microbial ecosystem. Application to" AOC Salers" cheese, 2001.

. Lait-(france,

J. Godon and R. Moletta, Une carte d'identité moléculaire pour les microéboueurs, Biofutur, issue.178, pp.38-40, 1998.
DOI : 10.1016/s0294-3506(98)80086-0

C. Goeury, Numerical modelling of oil spill drifts in continental and estuarine waters, 2012.
URL : https://hal.archives-ouvertes.fr/pastel-00778087

D. 2. Goeury, . La, . Marine-;-r, and A. Woessner, Mers et océans, 2015.

A. Gricajeva, V. Bendikien?, and L. Kal?dien?, Lipase of Bacillus stratosphericus L1: Cloning, expression and characterization, International Journal of Biological Macromolecules, vol.92, pp.96-104, 2016.
DOI : 10.1016/j.ijbiomac.2016.07.015

E. J. Gudiña, A. I. Rodrigues, V. De-freitas, Z. Azevedo, J. A. Teixeira et al., Valorization of agro-industrial wastes towards the production of rhamnolipids, Bioresource Technology, vol.212, pp.144-150, 2016.

N. W. Gunther, A. Nunez, W. Fett, and D. K. Solaiman, Production of rhamnolipids by Pseudomonas chlororaphis, a nonpathogenic bacterium, Applied and environmental microbiology, vol.71, issue.5, pp.2288-2293, 2005.

E. Haba, M. J. Espuny, M. Busquets, and A. Manresa, Screening and production of rhamnolipids by Pseudomonas aeruginosa 47T2 NCIB 40044 from waste frying oils, Journal of Applied Microbiology, vol.88, issue.3, pp.379-387, 2000.

J. D. Haddock, J. R. Horton, and D. T. Gibson, Dihydroxylation and dechlorination of chlorinated biphenyls by purified biphenyl 2, 3-dioxygenase from Pseudomonas sp. strain LB400, Journal of bacteriology, vol.177, issue.1, pp.20-26, 1995.

H. H. Hahn and R. Pfeifer, The contribution of parked vehicle emissions to the pollution of urban run-off, Science of The Total Environment, pp.525-533, 1994.

T. A. Hall, BioEdit: a user-friendly biological sequence alignment editor and analysis program for Windows 95/98/NT. Nucleic acids symposium series, vol.41, pp.95-98, 1999.

L. Hall-stoodley, J. W. Costerton, and P. Stoodley, Bacterial biofilms: from the natural environment to infectious diseases, Nature reviews microbiology, vol.2, issue.2, p.95, 2004.

A. Hameed, S. Lin, W. Lai, M. Shahina, Y. Liu et al., Idiomarina tyrosinivorans sp. nov., isolated from estuarine surface water, International Journal of Systematic and Evolutionary Microbiology, vol.66, issue.12, pp.5384-5391, 2016.

I. W. Hamley, Lipopeptides: from self-assembly to bioactivity, Chemical Communications, vol.51, issue.41, pp.8574-8583, 2015.

M. B. Hansen, S. E. Nielsen, and K. Berg, Re-examination and further development of a precise and rapid dye method for measuring cell growth/cell kill, Journal of immunological methods, vol.119, issue.2, pp.203-210, 1989.

A. K. Haritash and C. P. Kaushik, Biodegradation aspects of Polycyclic Aromatic Hydrocarbons (PAHs): A review, Journal of Hazardous Materials, vol.169, issue.1, pp.1-15, 2009.

R. M. Harshey, J. R. Mireles, and A. Toguchi, Use of cyclic heptapeptides for the inhibition of biofilm formation, 2003.

P. Hasanizadeh, H. Moghimi, and J. Hamedi, Biosurfactant production by Mocur circinelloides on waste frying oil and possible uses in crude oil remediation, Water Science et Technology, vol.6, issue.78, pp.1706-1714, 2017.

S. Häussler, M. Nimtz, T. Domke, V. Wray, and I. Steinmetz, Purification and characterization of a cytotoxic exolipid of Burkholderia pseudomallei, Infection and immunity, vol.66, issue.4, pp.1588-1593, 1998.

S. Hazrati, R. Rostami, M. Farjaminezhad, and M. Fazlzadeh, Preliminary assessment of BTEX concentrations in indoor air of residential buildings and atmospheric ambient air in Ardabil, Iran. Atmospheric Environment, vol.132, pp.91-97, 2016.

I. M. Head, D. M. Jones, and S. R. Larter, Biological activity in the deep subsurface and the origin of heavy oil, Nature, vol.426, p.344, 2003.

I. M. Head, D. M. Jones, and W. F. Röling, Marine microorganisms make a meal of oil, Nature Reviews Microbiology, vol.4, p.173, 2006.

B. P. Hedlund, A. D. Geiselbrecht, and J. T. Staley, Marinobacter strain NCE312 has a Pseudomonas-like naphthalene dioxygenase, FEMS Microbiology Letters, vol.201, issue.1, pp.47-51, 2001.

M. Henkel, M. Geissler, F. Weggenmann, and R. Hausmann, Production of microbial biosurfactants: Status quo of rhamnolipid and surfactin towards large-scale production, Biotechnology journal, vol.12, issue.7, p.1600561, 2017.

D. Hentati, A. Chebbi, S. Loukil, S. Kchaou, J. Godon et al., Biodegradation of fluoranthene by a newly isolated strain of Bacillus stratosphericus from Mediterranean seawater of the Sfax fishing harbour, Tunisia. Environmental Science and Pollution Research, vol.23, issue.15, pp.15088-15100, 2016.

G. Hernandez-raquet, H. Budzinski, P. Caumette, P. Dabert, K. Le-menach et al., Molecular diversity studies of bacterial communities of oil polluted microbial mats from the Etang de Berre (France), FEMS microbiology ecology, vol.58, issue.3, pp.550-562, 2006.
URL : https://hal.archives-ouvertes.fr/hal-01841789

J. Hofemeister, B. Conrad, B. Adler, B. Hofemeister, J. Feesche et al., Genetic analysis of the biosynthesis of nonribosomal peptide-and polyketide-like antibiotics, iron uptake and biofilm formation by Bacillus subtilis A1/3, Molecular Genetics and Genomics, vol.272, issue.4, pp.363-378, 2004.

B. Hörmann, M. M. Müller, C. Syldatk, and R. Hausmann, Rhamnolipid production by Burkholderia plantarii DSM 9509T, European journal of lipid science and technology, vol.112, issue.6, pp.674-680, 2010.

Z. Huang, Y. Hu, L. Shou, and M. Song, Isolation and partial characterization of cyclic lipopeptide antibiotics produced by Paenibacillus ehimensis B7, BMC Microbiology, vol.13, issue.1, p.87, 2013.

C. Huber, I. Klimant, C. Krause, T. Werner, T. Mayr et al., Optical sensor for seawater salinity, Fresenius' journal of analytical chemistry, vol.368, issue.2-3, pp.196-202, 2000.

H. Abdelwahed and W. , Study on the regulation and biosynthesis of fengycin and plipastatin produced by Bacillus subtilis, vol.1, 2011.

Y. Hwang, B. Park, J. Lim, M. Kim, I. Song et al., Evaluation of Genetic and Developmental Toxicity of Surfactin C from <i>Bacillus subtilis</i> BC1212, Journal of Health Science, vol.54, issue.1, pp.101-106, 2008.

R. Ihaka and R. Gentleman, R: a language for data analysis and graphics, Journal of computational and graphical statistics, vol.5, issue.3, pp.299-314, 1996.

D. Imhoff-stuckle and N. Pfennig, Isolation and characterization of a nicotinic acid-degrading sulfate-reducing bacterium, Desulfococcus niacini sp. nov, Archives of microbiology, vol.136, issue.3, pp.194-198, 1983.

I. , Biological Evaluation of Medical Devices. Part 5: Tests for In Vitro Cytotoxicity. International Organization for Standardization, situ burning of spilled oil. IPIECA-IOGP Good Practice Guide Series, pp.10993-10998, 2009.

, Effects of Oil Pollution on the Marine Environment. Technical Information Paper (TIP) No. 13, International Tanker Owners Pollution Federation, ITOPF, 2011.

M. Jeanbille, Réponse des consortia microbiens benthiques à une contamination chronique aux hydrocarbures, 2015.

M. Jeanbille, J. Gury, R. Duran, J. Tronczynski, H. Agogué et al., Response of Core Microbial Consortia to Chronic Hydrocarbon Contaminations in Coastal Sediment Habitats, Frontiers in Microbiology, vol.7, 1637.
URL : https://hal.archives-ouvertes.fr/hal-01417951

T. C. Jeffries, M. L. Schmitz-fontes, D. P. Harrison, V. Van-dongen-vogels, B. D. Eyre et al., Bacterioplankton dynamics within a large anthropogenically impacted urban estuary, Frontiers in microbiology, vol.6, p.1438, 2016.

N. Jemil, A. Manresa, F. Rabanal, H. Ben-ayed, N. Hmidet et al., Structural characterization and identification of cyclic lipopeptides produced by Bacillus methylotrophicus DCS1 strain, Journal of Chromatography B, vol.1060, pp.374-386, 2017.

L. Jiang, C. Shen, X. Long, G. Zhang, and Q. Meng, Rhamnolipids elicit the same cytotoxic sensitivity between cancer cell and normal cell by reducing surface tension of culture medium, Applied Microbiology and Biotechnology, vol.98, issue.24, pp.10187-10196, 2014.

A. R. Johnsen and U. Karlson, PAH degradation capacity of soil microbial communities-does it depend on PAH exposure? Microbial ecology, vol.50, pp.488-495, 2005.

S. J. Joshi, S. J. Geetha, and A. J. Desai, Characterization and Application of Biosurfactant Produced by Bacillus licheniformis R2, Applied Biochemistry and Biotechnology, vol.177, issue.2, pp.346-361, 2015.

T. H. Jukes and C. R. Cantor, Evolution of protein molecules, Munro HN (ed) Mammalian protein metabolism, pp.21-132, 1969.

J. Guerra-de, O. Crispin-humberto, and G. , Properties of a biosurfactant produced by Bacillus pumilus using vinasse and waste frying oil as alternative carbon sources, Brazilian Archives of Biology and Technology, vol.56, issue.1, pp.155-160, 2013.

R. A. Kanaly and S. Harayama, Biodegradation of High-Molecular-Weight Polycyclic Aromatic Hydrocarbons by Bacteria, Journal of Bacteriology, vol.182, issue.8, pp.2059-2067, 2000.

M. Kanlayavattanakul and N. Lourith, Lipopeptides in cosmetics, International journal of cosmetic science, vol.32, issue.1, pp.1-8, 2010.

B. Karimi, M. Habibi, and M. Esvand, Biodegradation of naphthalene using Pseudomonas aeruginosa by up flow anoxic-aerobic continuous flow combined bioreactor, Journal of Environmental Health Science and Engineering, vol.13, issue.1, p.26, 2015.

A. Khopade, B. Ren, X. Liu, K. Mahadik, L. Zhang et al., Production and characterization of biosurfactant from marine Streptomyces species B3, Journal of Colloid and Interface Science, vol.367, issue.1, pp.311-318, 2012.

K. Kim, S. A. Jahan, E. Kabir, and R. J. Brown, A review of airborne polycyclic aromatic hydrocarbons (PAHs) and their human health effects, Environment International, vol.60, pp.71-80, 2013.

G. S. Kiran, B. Sabarathnam, N. Thajuddin, and J. Selvin, Production of Glycolipid Biosurfactant from Sponge-Associated Marine Actinobacterium Brachybacterium paraconglomeratum MSA21, Journal of Surfactants and Detergents, vol.17, issue.3, pp.531-542, 2014.

U. Kirso and N. Irha, Role of algae in fate of carcinogenic polycyclic aromatic hydrocarbons in the aquatic environment, Ecotoxicology and environmental safety, vol.41, issue.1, pp.83-89, 1998.

J. Kjeldahl, Neue methode zur bestimmung des stickstoffs in organischen körpern, Zeitschrift für analytische Chemie, vol.22, issue.1, pp.366-382, 1883.

B. Klein, W. Roether, B. B. Manca, D. Bregant, V. Beitzel et al., The large deep water transient in the Eastern Mediterranean, Deep Sea Research Part I: Oceanographic Research Papers, vol.46, pp.371-414, 1999.

R. J. Knechtel, A more economical method for the determination of chemical oxygen demand, 1978.

M. Korlevi?, J. Zucko, M. N. Dragi?, M. Bla?ina, E. Pustijanac et al., Bacterial diversity of polluted surface sediments in the northern Adriatic Sea, Systematic and applied microbiology, vol.38, issue.3, pp.189-197, 2015.

J. E. Kostka, O. Prakash, W. A. Overholt, S. J. Green, G. Freyer et al., Hydrocarbon degrading Bacteria and the bacterial community response in gulf of Mexico beach sands impacted by the deepwater horizon oil spill, Applied and Environmental Microbiology, vol.77, issue.22, pp.7962-7974, 2011.

A. P. Kumar, A. Janardhan, B. Viswanath, K. Monika, J. Jung et al., Evaluation of orange peel for biosurfactant production by Bacillus licheniformis and their ability to degrade naphthalene and crude oil, Biotech, vol.6, issue.3, p.43, 2016.

M. Kumar, V. León, A. D. Materano, and O. A. Ilzins, A halotolerant and thermotolerant Bacillus sp. degrades hydrocarbons and produces tensio-active emulsifying agent, World Journal of Microbiology and Biotechnology, vol.23, issue.2, pp.211-220, 2007.

S. Labana, M. Kapur, D. K. Malik, D. Prakash, and R. Jain, Diversity, biodegradation and bioremediation of polycyclic aromatic hydrocarbons, Environmental bioremediation technologies, pp.409-443, 2007.

T. Lacoue-labarthe, Incorporation des métaux dans les oeufs de la seiche commune Sepia officinalis et effets potentiels sur les fonctions digestives et immunitaires, 2007.

G. Lan, Q. Fan, Y. Liu, C. Chen, G. Li et al., Rhamnolipid production from waste cooking oil using Pseudomonas SWP-4, Biochemical Engineering Journal, vol.101, pp.44-54, 2015.

N. Lang, P. Delichere, and A. Tuel, Post-synthesis introduction of transition metals in surfactantcontaining MCM-41 materials, Microporous and mesoporous materials, vol.56, issue.2, pp.203-217, 2002.
URL : https://hal.archives-ouvertes.fr/hal-00007173

J. S. Latimer and J. G. Quinn, Aliphatic petroleum and biogenic hydrocarbons entering Narragansett Bay from tributaries under dry weather conditions, Estuaries, vol.21, issue.1, pp.91-107, 1998.
DOI : 10.2307/1352549

W. T. Lawrence, Physiology of the acute wound, Clinics in plastic surgery, vol.25, issue.3, pp.321-340, 1998.

L. Dû-lacoste and M. , Etude des phénomènes de biotransformation des hydrocarbures aromatiques polycycliques (HAP) par les organismes aquatiques (poissons) : relation expositiongénotoxicité, 2008.

J. Leahy and R. Colwell, Microbial Degradation of Hydrocarbons in the Environment, Microbiology Reviews, vol.54, issue.3, pp.305-315, 1990.

J. H. Leamon, W. L. Lee, K. R. Tartaro, J. R. Lanza, G. J. Sarkis et al., A massively parallel PicoTiterPlate? based platform for discrete picoliter-scale polymerase chain reactions, Electrophoresis, vol.24, issue.21, pp.3769-3777, 2003.

M. Lee, M. K. Kim, M. Vancanneyt, J. Swings, S. Kim et al., Tetragenococcus koreensis sp. nov., a novel rhamnolipid-producing bacterium, International journal of systematic and evolutionary microbiology, vol.55, issue.4, pp.1409-1413, 2005.

A. Lei, Z. Hu, Y. Wong, N. F. Tam, and .. , Removal of fluoranthene and pyrene by different microalgal species, Bioresource technology, vol.98, issue.2, pp.273-280, 2007.

J. Li, M. Deng, Y. Wang, and W. Chen, Production and characteristics of biosurfactant produced by Bacillus pseudomycoides BS6 utilizing soybean oil waste, International Biodeterioration & Biodegradation, vol.112, pp.72-79, 2016.

Y. Li, N. I. Haddad, S. Yang, and B. Mu, Variants of lipopeptides produced by Bacillus licheniformis HSN221 in different medium components evaluated by a rapid method ESI-MS, International Journal of Peptide Research and Therapeutics, vol.14, issue.3, pp.229-235, 2008.

A. O. Lima, A. Cabral, F. D. Andreote, A. Cavalett, M. L. Pessatti et al., Draft Genome Sequence of <span class="named-content genus-species" id="named-content-1">Bacillus stratosphericus</span> LAMA 585, Isolated from the Atlantic Deep Sea, Genome Announcements, issue.3, p.1, 2013.

M. Lin, X. Hu, W. Chen, H. Wang, and C. Wang, Biodegradation of phenanthrene by Pseudomonas sp. BZ-3, isolated from crude oil contaminated soil, International Biodeterioration & Biodegradation, vol.94, pp.176-181, 2014.

, Références Bibliographiques

A. B. Lins, A. M. Bione, T. C. Silva, D. G. De-souza, and G. M. Campos-takaki, Low-cost production of biosurfactant by Cunninghamella phaeospora using agro-industrial wastes, Microbes in the Spotlight: Recent Progress in the Understanding of Beneficial and Harmful Microorganisms ed Mendez Vilas A, pp.339-343, 2016.

B. Liu, M. Ju, J. Liu, W. Wu, and X. Li, Isolation, identification, and crude oil degradation characteristics of a high-temperature, hydrocarbon-degrading strain, Marine pollution bulletin, vol.106, issue.1-2, pp.301-307, 2016.

F. Liu, Y. Wang, C. Qu, Z. Zheng, J. Miao et al., The complete genome of hydrocarbon-degrading Pseudoalteromonas sp. NJ289 and its phylogenetic relationship, Acta Oceanologica Sinica, vol.36, issue.2, pp.88-93, 2017.

L. Liu, Y. Li, S. Li, N. Hu, Y. He et al., Comparison of nextgeneration sequencing systems, BioMed Research International, 2012.

D. Lohrer and N. Hancock, Marine soft sediments: more diversity than meets the eye, Water and Atmosphere, vol.3, p.12, 2004.

E. R. Long, D. D. Macdonald, S. L. Smith, and F. D. Calder, Incidence of adverse biological of chemical concentrations in marine and estuarine sediments, Environmental Management, vol.19, issue.1, pp.81-97, 1995.

A. Louati, B. Elleuch, M. Kallel, A. Saliot, J. Dagaut et al., Hydrocarbon contamination of coastal sediments from the Sfax area (Tunisia), Marine Pollution Bulletin, vol.42, issue.6, pp.444-451, 2001.

W. Ludwig, G. Kirchhof, N. Klugbauer, M. Weizenegger, D. Betzl et al., Complete 23S ribosomal RNA sequences of Grampositive bacteria with a low DNA G+ C content, Systematic and applied microbiology, vol.15, issue.4, pp.487-501, 1992.

E. H. Madslien, H. T. Rønning, T. Lindbäck, B. Hassel, M. A. Andersson et al., Lichenysin is produced by most Bacillus licheniformis strains, Journal of Applied Microbiology, vol.115, issue.4, pp.1068-1080, 2013.

E. R. Magalhães, F. L. Silva, M. A. Sousa, and E. S. Santos, Use of different agroindustrial waste and produced water for biosurfactant production, Biosciences Biotechnology Research Asia, vol.15, issue.1, pp.17-26, 2018.

B. Mahanty, K. Pakshirajan, and V. Dasu, Production and properties of a biosurfactant applied to polycyclic aromatic hydrocarbon solubilization, Applied Biochemistry and Biotechnology, vol.134, issue.2, pp.129-141, 2006.

A. Maiti, S. Das, and N. Bhattacharyya, Bioremediation of high molecular weight polycyclic aromatic hydrocarbons by Bacillus thuringiensis strain NA2, J Sci, vol.1, issue.4, pp.72-75, 2012.

R. S. Makkar, S. S. Cameotra, and I. M. Banat, Advances in utilization of renewable substrates for biosurfactant production, AMB express, vol.1, issue.1, p.5, 2011.

D. Maliji, Z. Olama, and H. Holail, Environmental studies on the microbial degradation of oil hydrocarbons and its application in Lebanese oil polluted coastal and marine ecosystem, Int J Curr Microbiol App Sci, vol.2, issue.6, pp.1-18, 2013.

S. M. Mandal, A. E. Barbosa, and O. L. Franco, Lipopeptides in microbial infection control: scope and reality for industry, Biotechnology Advances, vol.31, issue.2, pp.338-345, 2013.

S. Maneerat and K. Phetrong, Isolation of biosurfactant-producing marine bacteria and characteristics of selected biosurfactant, Songklanakarin J Sci Technol, vol.29, issue.3, pp.781-791, 2007.

P. Mani, G. Dineshkumar, T. Jayaseelan, K. Deepalakshmi, C. Ganesh-kumar et al., Antimicrobial activities of a promising glycolipid biosurfactant from a novel marine Staphylococcus saprophyticus SBPS 15, Biotech, vol.6, issue.3, p.163, 2016.

M. Marchand and R. Kantin, Contaminants chimiques en milieux aquatiques(Troisième partie, Présentation de quelques éléments caractéristiques), 1997.

R. Marchant and I. M. Banat, Microbial biosurfactants: challenges and opportunities for future exploitation, Trends in Biotechnology, vol.30, issue.11, pp.558-565, 2012.
DOI : 10.1016/j.tibtech.2012.07.003

M. Margulies, M. Egholm, W. E. Altman, S. Attiya, J. S. Bader et al., Genome sequencing in microfabricated highdensity picolitre reactors, Nature, vol.437, issue.7057, p.376, 2005.
DOI : 10.1038/nature03959

URL : https://www.nature.com/articles/nature03959.pdf

M. Markets, Markets and Markets, 2016.

P. Martin, Wound healing-aiming for perfect skin regeneration, Science, vol.276, issue.5309, pp.75-81, 1997.

N. Massu and G. Landmann, Connaissance des impacts du changement climatique sur la biodiversité en France métropolitaine, synthèse de la bibliographie, p.180, 2011.

J. C. Mata-sandoval, J. Karns, and A. Torrents, The influence of surfactants and biosurfactants on the bioavailability of hydrophobic organic pollutants in subsurface environments, Revista Internacional de Contaminación Ambiental, vol.16, issue.4, 2000.

D. J. Mcclements and C. E. Gumus, Natural emulsifiers-Biosurfactants, phospholipids, biopolymers, and colloidal particles: Molecular and physicochemical basis of functional performance, Advances in Colloid and Interface Science, vol.234, pp.3-26, 2016.

A. Mcelroy, J. Farrington, and J. Teal, Bioavailability of polycyclic aromatic hydrocarbons in the aquatic environment, 1989.

T. J. Mcgenity and . Timmis, Halophilic Hydrocarbon Degraders, Handbook of Hydrocarbon and Lipid Microbiology, pp.1939-1951, 2010.

A. J. Mearns, D. J. Reish, M. Bissell, A. M. Morrison, M. A. Rempel-hester et al., Effects of Pollution on Marine Organisms, Water Environment Research, vol.90, issue.10, pp.1206-1300, 2018.
DOI : 10.2175/106143007x218683

M. Jin, H. , H. Kim, K. Ok-jeon, and C. , Alteromonas naphthalenivorans sp. nov., a polycyclic aromatic hydrocarbon-degrading bacterium isolated from tidal-flat sediment, International Journal of Systematic and Evolutionary Microbiology, vol.65, issue.11, pp.4208-4214, 2015.

I. Mnif and D. Ghribi, Review lipopeptides biosurfactants: mean classes and new insights for industrial, biomedical, and environmental applications, Peptide Science, vol.104, issue.3, pp.129-147, 2015.
DOI : 10.1002/bip.22630

S. Mnif, M. Chamkha, M. Labat, and S. Sayadi, Simultaneous hydrocarbon biodegradation and biosurfactant production by oilfield-selected bacteria, Journal of applied microbiology, vol.111, issue.3, pp.525-536, 2011.
DOI : 10.1111/j.1365-2672.2011.05071.x

URL : https://hal.archives-ouvertes.fr/hal-00713642

S. Mnif, M. Chamkha, and S. Sayadi, Isolation and characterization of Halomonas sp. strain C2SS100, a hydrocarbon-degrading bacterium under hypersaline conditions, Journal of Applied Microbiology, vol.107, issue.3, pp.785-794, 2009.

S. Mnif, A. Zayen, F. Karray, V. Bru-adan, S. Loukil et al., Microbial population changes in anaerobic membrane bioreactor treating landfill leachate monitored by single-strand conformation polymorphism analysis of 16S rDNA gene fragments, International biodeterioration & biodegradation, vol.73, pp.50-59, 2012.

P. K. Mohan, G. Nakhla, and E. K. Yanful, Biodegradability of surfactants under aerobic, anoxic, and anaerobic conditions, Journal of Environmental Engineering, vol.132, issue.2, pp.279-283, 2006.
DOI : 10.1061/(asce)0733-9372(2006)132:2(279)

N. B. Mohd-zin, M. Yusof, B. Oslan, S. N. Wasoh, H. Tan et al., Utilization of acid pre-treated coconut dregs as a substrate for production of detergent compatible lipase by Bacillus stratosphericus, vol.7, p.131, 2017.

J. M. Monier and S. Cécillon, Les outils de biologie moléculaire et leur utilisation dans le domaine de la gestion des sites pollués-Synthèse, 2015.

J. J. Morelli and G. Szajer, Analysis of surfactants: Part I, Journal of Surfactants and Detergents, vol.3, issue.4, pp.539-552, 2000.

N. Morley, J. Burton, S. Tankere, and J. Martin, Distribution and behaviour of some dissolved trace metals in the western Mediterranean Sea, Deep Sea Research Part II: Topical Studies in Oceanography, vol.44, pp.675-691, 1997.

C. E. Morris, M. Bardin, O. Berge, P. Frey-klett, N. Fromin et al., Microbial biodiversity: approaches to experimental design and hypothesis testing in primary scientific literature from 1975 to 1999, Microbiology and Molecular Biology Reviews, vol.66, issue.4, pp.592-616, 2002.
URL : https://hal.archives-ouvertes.fr/hal-02126964

I. Moya-ramírez, M. García-román, and A. Fernández-arteaga, Waste Frying Oil Hydrolysis in a Reverse Micellar System, ACS Sustainable Chemistry & Engineering, vol.4, issue.3, pp.1025-1031, 2016.

S. Mukherjee, P. Das, C. Sivapathasekaran, and R. Sen, Antimicrobial biosurfactants from marine Bacillus circulans: extracellular synthesis and purification, Letters in Applied Microbiology, vol.48, issue.3, pp.281-288, 2009.

M. M. Müller, B. Hormann, M. Kugel, C. Syldatk, and R. Hausmann, Evaluation of rhamnolipid production capacity of Pseudomonas aeruginosa PAO1 in comparison to the rhamnolipid overproducer strains DSM 7108 and DSM 2874, Applied Microbiology and Biotechnology, vol.89, issue.3, pp.585-92, 2011.

M. M. Müller, B. Hörmann, M. Kugel, C. Syldatk, and R. Hausmann, Evaluation of rhamnolipid production capacity of Pseudomonas aeruginosa PAO1 in comparison to the rhamnolipid over-producer strains DSM 7108 and DSM 2874, Applied microbiology and biotechnology, vol.89, issue.3, pp.585-592, 2011.

C. N. Mulligan, Environmental applications for biosurfactants, Environmental pollution, vol.133, issue.2, pp.183-198, 2005.

K. Muthusamy, S. Gopalakrishnan, T. K. Ravi, and P. Sivachidambaram, Biosurfactants: properties, commercial production and application. Current science, pp.736-747, 2008.

Z. Najmi, G. Ebrahimipour, A. Franzetti, and I. M. Banat, In situ downstream strategies for costeffective bio/surfactant recovery, Biotechnology and Applied Biochemistry, vol.65, issue.4, pp.523-532, 2018.

K. Nakajima, A. Sato, Y. Takahara, and T. Iida, Microbial Oxidation of Isoprenoid Alkanes, Phytane, Norpristane and Farnesane, Agricultural and Biological Chemistry, vol.49, issue.7, pp.1993-2002, 1985.

M. Nakano, S. Iehata, R. Tanaka, and H. Maeda, Extracellular neutral lipids produced by the marine bacteria Marinobacter sp, Biocontrol science, vol.17, issue.2, pp.69-75, 2012.

S. Nalini and R. Parthasarathi, Biosurfactant production by Serratia rubidaea SNAU02 isolated from hydrocarbon contaminated soil and its physico-chemical characterization, Bioresource technology, vol.147, pp.619-622, 2013.

S. Namwong and S. Tanasupawat, Identification of Staphylococcus strain CH1-8 and its oil-degradation, Journal of Applied Pharmaceutical Science, vol.4, pp.24-029, 2014.

S. G. Nanjani and H. P. Soni, Characterization of an extremely halotolerant Staphylococcus arlettae HPSSN35C isolated from Dwarka Beach, India. Journal of Basic Microbiology, vol.54, issue.8, pp.843-850, 2013.

A. S. Nayak, M. Vijaykumar, and T. Karegoudar, Characterization of biosurfactant produced by Pseudoxanthomonas sp. PNK-04 and its application in bioremediation, International Biodeterioration & Biodegradation, vol.63, issue.1, pp.73-79, 2009.

G. Nedia, A. Chafai, S. M. Moncef, and Y. Chokri, Spatial distribution of heavy metals in the coastal zone of "Sfax-Kerkennah" plateau, Tunisia. Environmental Progress & Sustainable Energy, vol.30, issue.2, pp.221-233, 2011.

T. Nikitha, M. Satyaprakash, S. S. Vani, B. Sadhana, and S. Padal, A review on polycyclic aromatic hydrocarbons: their transport, fate and biodegradation in the environment, Int J Curr Microbiol Appl Sci, vol.6, issue.4, pp.1627-1639, 2017.

O. C. Nwinyi, O. O. Ajayi, and O. O. Amund, Degradation of polynuclear aromatic hydrocarbons by two strains of Pseudomonas, brazilian journal of microbiology, vol.47, issue.3, pp.551-562, 2016.

G. A. O&apos;toole, Microtiter dish biofilm formation assay, Journal of visualized experiments, issue.47, 2011.

P. D. O&apos;hara and L. A. Morandin, Effects of sheens associated with offshore oil and gas development on the feather microstructure of pelagic seabirds, Marine Pollution Bulletin, vol.60, issue.5, pp.672-678, 2010.

D. Onbasli and B. Aslim, Biosurfactant production in sugar beet molasses by some Pseudomonas spp, J Environ Biol, vol.30, issue.1, pp.161-163, 2009.

M. Ongena and P. Jacques, Bacillus lipopeptides: versatile weapons for plant disease biocontrol, Trends in microbiology, vol.16, issue.3, pp.115-125, 2008.
DOI : 10.1016/j.tim.2007.12.009

C. O. Onwosi and F. J. Odibo, Effects of carbon and nitrogen sources on rhamnolipid biosurfactant production by Pseudomonas nitroreducens isolated from soil, World Journal of Microbiology and Biotechnology, vol.28, issue.3, pp.937-942, 2012.

A. C. Ortmann and Y. Lu, Initial community and environment determine the response of bacterial communities to dispersant and oil contamination, Marine Pollution Bulletin, vol.90, issue.1, pp.106-114, 2015.

M. Pacwa-p?ociniczak, G. A. P?aza, Z. Piotrowska-seget, and S. S. Cameotra, Environmental applications of biosurfactants: recent advances, International journal of molecular sciences, vol.12, issue.1, pp.633-654, 2011.

M. Pacwa-p?ociniczak, G. A. P?aza, A. Poliwoda, and Z. Piotrowska-seget, Characterization of hydrocarbon-degrading and biosurfactant-producing Pseudomonas sp. P-1 strain as a potential tool for bioremediation of petroleum-contaminated soil, Environmental Science and Pollution Research, vol.21, issue.15, pp.9385-9395, 2014.

B. Padmapriya, S. Suganthi, and R. Anishya, Screening, optimization and production of biosurfactants by Candida species isolated from oil polluted soils, American-Eurasian Journal of Agricutural & Environmental Sciences, vol.13, issue.2, pp.227-233, 2013.

S. Pailan and P. Saha, Chemotaxis and degradation of organophosphate compound by a novel moderately thermo-halo tolerant Pseudomonas sp. strain BUR11: evidence for possible existence of two pathways for degradation, PeerJ, vol.3, 1378.

S. Paissé, F. Coulon, M. Goñi-urriza, L. Peperzak, T. J. Mcgenity et al., Structure of bacterial communities along a hydrocarbon contamination gradient in a coastal sediment, FEMS microbiology ecology, vol.66, issue.2, pp.295-305, 2008.

M. M. Palmisano, L. K. Nakamura, K. E. Duncan, C. A. Istock, and F. M. Cohan, Bacillus sonorensis sp. nov., a close relative of Bacillus licheniformis, isolated from soil in the Sonoran Desert, Arizona. International Journal of Systematic and Evolutionary Microbiology, vol.51, issue.5, pp.1671-1679, 2001.

S. L. Paralovo, G. C. Borillo, C. G. Barbosa, A. F. Godoi, C. I. Yamamoto et al., Observations of atmospheric monoaromatic hydrocarbons at urban, semi-urban and forest environments in the Amazon region, Atmospheric Environment, vol.128, pp.175-184, 2016.

S. Park, S. Y. Yoon, Y. Jung, S. Won, and J. Yoon, Gramella sediminilitoris sp. nov., isolated from a tidal flat, International Journal of Systematic and Evolutionary Microbiology, vol.66, issue.7, pp.2704-2710, 2016.
DOI : 10.1099/ijsem.0.001111

URL : http://ijs.microbiologyresearch.org/deliver/fulltext/ijsem/66/7/2704_ijsem001111.pdf?itemId=/content/journal/ijsem/10.1099/ijsem.0.001111&mimeType=pdf&isFastTrackArticle=

S. Patel, S. Ahmed, and J. S. Eswari, Therapeutic cyclic lipopeptides mining from microbes: latest strides and hurdles, World Journal of Microbiology and Biotechnology, vol.31, issue.8, pp.1177-1193, 2015.

S. Patil, J. Paradeshi, and B. Chaudhari, Suppression of charcoal rot in soybean by moderately halotolerant Pseudomonas aeruginosa GS-33 under saline conditions, Journal of Basic Microbiology, vol.56, issue.8, pp.889-899, 2016.

K. Patowary, R. Patowary, M. C. Kalita, and S. Deka, Characterization of Biosurfactant Produced during Degradation of Hydrocarbons Using Crude Oil As Sole Source of Carbon, Frontiers in Microbiology, vol.8, issue.279, 2017.

J. F. Patton and M. P. Dieter, Effects of petroleum hydrocarbons on hepatic function in the duck, Comparative Biochemistry and Physiology Part C: Comparative Pharmacology, vol.65, issue.1, pp.33-36, 1980.

H. Patzelt, Hydrocarbon degradation under hypersaline conditions. in: Adaptation to life at high salt concentrations in archaea, bacteria, and eukarya, pp.105-122, 2005.

Y. Pecci, F. Rivardo, M. G. Martinotti, and G. Allegrone, LC/ESI-MS/MS characterisation of lipopeptide biosurfactants produced by the Bacillus licheniformis V9T14 strain, Journal of Mass Spectrometry, vol.45, issue.7, pp.772-778, 2010.

R. Peng, A. Xiong, Y. Xue, X. Fu, F. Gao et al., Microbial biodegradation of polyaromatic hydrocarbons, FEMS microbiology reviews, vol.32, issue.6, pp.927-955, 2008.

S. Peng, The nutrient, total petroleum hydrocarbon and heavy metal contents in the seawater of Bohai Bay, China: Temporal-spatial variations, sources, pollution statuses, and ecological risks, Marine Pollution Bulletin, vol.95, issue.1, pp.445-451, 2015.

A. Perfumo, I. M. Banat, F. Canganella, and R. Marchant, Rhamnolipid production by a novel thermophilic hydrocarbon-degrading Pseudomonas aeruginosa AP02-1, Applied Microbiology and Biotechnology, vol.72, issue.1, p.132, 2005.

O. Pornsunthorntawee, N. Arttaweeporn, S. Paisanjit, P. Somboonthanate, M. Abe et al., Isolation and comparison of biosurfactants produced by Bacillus subtilis PT2 and Pseudomonas aeruginosa SP4 for microbial surfactant-enhanced oil recovery, Biochemical Engineering Journal, vol.42, issue.2, pp.172-179, 2008.

G. Potters, Marine Pollution, 2013.

S. M. Powell, J. P. Bowman, S. H. Ferguson, and I. Snape, The importance of soil characteristics to the structure of alkane-degrading bacterial communities on sub-Antarctic Macquarie Island, Soil Biology and Biochemistry, vol.42, issue.11, pp.2012-2021, 2010.

R. C. Prince, A. Gramain, T. J. Mcgenity, and . Timmis, Prokaryotic Hydrocarbon Degraders, Handbook of Hydrocarbon and Lipid Microbiology, pp.1669-1692, 2010.
DOI : 10.1007/978-3-319-60053-6_15-1

P. Smap-iii-tunisie, Stratégies de gestion intégrée des zones côtières pour les régions du KROUMIRIE ET MOGODS et les municipalités de Sfax, 2006.

J. Qi, B. Wang, J. Li, H. Ning, Y. Wang et al., Genetic determinants involved in the biodegradation of naphthalene and phenanthrene in Pseudomonas aeruginosa PAO1, Environmental Science and Pollution Research, vol.22, issue.9, pp.6743-6755, 2015.

M. A. Quail, M. Smith, P. Coupland, T. D. Otto, S. R. Harris et al., A tale of three next generation sequencing platforms: comparison of Ion Torrent, Pacific Biosciences and Illumina MiSeq sequencers, BMC genomics, vol.13, issue.1, p.341, 2012.

B. Quéguiner, Biogéochimie Côtière. Centre d'Océanologie de Marseille, Aix-Marseille Université; CNRS; LOPB UMR 6535, Laboratoire d'Océanographie et de Biogéochimie, p.49, 2009.

A. Rajasekar, T. G. Babu, S. K. Pandian, S. Maruthamuthu, N. Palaniswamy et al., Biodegradation and corrosion behavior of manganese oxidizer Bacillus cereus ACE4 in diesel transporting pipeline, Corrosion Science, vol.49, issue.6, pp.2694-2710, 2007.

J. Rajendhran and P. Gunasekaran, Microbial phylogeny and diversity: small subunit ribosomal RNA sequence analysis and beyond, Microbiological research, vol.166, issue.2, pp.99-110, 2011.
DOI : 10.1016/j.micres.2010.02.003

URL : https://doi.org/10.1016/j.micres.2010.02.003

V. Rajeswari, S. K. Priyadarshini, V. Saranya, P. Suguna, and R. Shenbagarathai, Immunostimulation by phospholipopeptide biosurfactant from Staphylococcus hominis in Oreochromis mossambicus, Fish & shellfish immunology, vol.48, pp.244-253, 2016.

K. Ramani, S. C. Jain, A. B. Mandal, and G. Sekaran, Microbial induced lipoprotein biosurfactant from slaughterhouse lipid waste and its application to the removal of metal ions from aqueous solution, Colloids and Surfaces B: Biointerfaces, vol.97, pp.254-263, 2012.

K. K. Randhawa and P. K. Rahman, Rhamnolipid biosurfactants-past, present, and future scenario of global market, Frontiers in Microbiology, vol.5, p.454, 2014.

L. Ranjard, L'apport des techniques de la biologie moléculaire à la connaissance de la biodiversité microbienne dans les sols et de ses fonctions, Innovations Agronomiques, issue.21, pp.31-43, 2012.

J. Readman, G. Fillmann, I. Tolosa, J. Bartocci, J. Villeneuve et al., Petroleum and PAH contamination of the Black Sea, Marine Pollution Bulletin, vol.44, issue.1, pp.48-62, 2002.

M. S. Reddy, S. Basha, H. Joshi, and G. Ramachandraiah, Seasonal distribution and contamination levels of total PHCs, PAHs and heavy metals in coastal waters of the Alang-Sosiya ship scrapping yard, Gulf of Cambay, India. Chemosphere, issue.11, pp.1587-1593, 2005.

H. Rehm and I. Reiff, Mechanisms and occurrence of microbial oxidation of long-chain alkanes, Reactors and Reactions, pp.175-215, 1981.

K. Rehmann, H. P. Noll, C. E. Steinberg, and A. A. Kettrup, Pyrene degradation by Mycobacterium sp. strain KR2, Chemosphere, vol.36, issue.14, pp.2977-2992, 1998.

R. Reis, G. Pacheco, A. Pereira, and D. Freire, Biosurfactants: production and applications, Biodegradation-Life of Science, 2013.

M. W. Rey, P. Ramaiya, B. A. Nelson, S. D. Brody-karpin, E. J. Zaretsky et al., Complete genome sequence of the industrial bacterium Bacillus licheniformis and comparisons with closely related Bacillus species, R77. Références Bibliographiques, vol.5, 2004.

T. ?ezanka, L. Siristova, and K. Sigler, Rhamnolipid-producing thermophilic bacteria of species Thermus and Meiothermus, Extremophiles, vol.15, issue.6, p.697, 2011.

J. Richir and S. Gobert, Trace elements in marine environments: occurrence, threats and monitoring with special focus on the costal mediterranean, Journal of Environmental and Analytical Toxicology, vol.6, issue.1, 2016.

F. Rivardo, R. J. Turner, G. Allegrone, H. Ceri, and M. G. Martinotti, Anti-adhesion activity of two biosurfactants produced by Bacillus spp. prevents biofilm formation of human bacterial pathogens, Applied Microbiology and Biotechnology, vol.83, issue.3, pp.541-553, 2009.

M. S. Roberts, L. K. Nakamura, and F. M. Cohan, Bacillus mojavensis sp. nov., Distinguishable from Bacillus subtilis by Sexual Isolation, Divergence in DNA Sequence, and Differences in Fatty Acid Composition, International Journal of Systematic Bacteriology, vol.44, issue.2, pp.256-264, 1994.

J. Rodier, C. Bazin, J. Broutin, P. Chambon, H. Champsaur et al., , 1996.

L. Rodrigues, I. M. Banat, J. Teixeira, and R. Oliveira, Biosurfactants: potential applications in medicine, Journal of Antimicrobial Chemotherapy, vol.57, issue.4, pp.609-618, 2006.

L. Rodrigues, H. C. Van-der-mei, J. Teixeira, and R. Oliveira, Influence of biosurfactants from probiotic bacteria on formation of biofilms on voice prostheses, Applied and environmental microbiology, vol.70, issue.7, pp.4408-4410, 2004.

E. Z. Ron and E. Rosenberg, Enhanced bioremediation of oil spills in the sea, Current Opinion in biotechnology, vol.27, pp.191-194, 2014.

M. Ronaghi, M. Uhlen, and P. Nyren, Real-time DNA sequencing using detection of pyrophosphate release, Analytic Biochemistry, vol.242, issue.1, pp.84-89, 1998.

M. Ronaghi, S. Karamohamed, B. Pettersson, M. Uhlen, and P. Nyren, A sequencing method based on real-time pyrophosphate, Science, vol.281, pp.363-365, 1996.

A. P. Rooney, N. P. Price, K. J. Ray, and T. Kuo, Isolation and characterization of rhamnolipidproducing bacterial strains from a biodiesel facility, FEMS microbiology letters, vol.295, issue.1, pp.82-87, 2009.

M. C. Rosano-hernández, H. Ramírez-saad, and L. Fernández-linares, Petroleum-influenced beach sediments of the Campeche Bank, Mexico: diversity and bacterial community structure assessment, Journal of environmental management, vol.95, pp.325-331, 2012.

E. Rosenberg and E. Ron, High-and low-molecular-mass microbial surfactants, Applied microbiology and biotechnology, vol.52, issue.2, pp.154-162, 1999.

E. Rosenberg, Z. Schwartz, A. Tenenbaum, C. Rubinovitz, R. Legmann et al., A MICROBIAL POLYMER THAT CHANGES THE SURFACE PROPERTIES OF LIMESTONE: EFFECT OF BIODISPERSAN IN GRINDING LIMESTONE AND MAKING PAPER, Journal of Dispersion Science and Technology, vol.10, issue.3, pp.241-250, 1989.

D. Rubio-ribeaux, R. F. Da-silva-andrade, G. S. Da-silva, R. A. De-holanda, M. A. Pele et al., Promising biosurfactant produced by a new Candida tropicalis UCP 1613 strain using substrates from renewable-resources, Africain Journal of Microbiology Research, vol.11, issue.23, pp.981-991, 2017.

Z. Sadouk, H. Hacene, and A. Tazerouti, Biosurfactants Production from Low Cost Substrate and Degradation of Diesel Oil by a Rhodococcus Strain, Oil & Gas Science and Technology-Rev. IFP, vol.63, issue.6, pp.747-753, 2008.
URL : https://hal.archives-ouvertes.fr/hal-02002052

R. Sahnoun, I. Mnif, H. Fetoui, R. Gdoura, K. Chaabouni et al., Evaluation of Bacillus subtilis SPB1 Lipopeptide Biosurfactant Toxicity Towards Mice, International Journal of Peptide Research and Therapeutics, vol.20, issue.3, pp.333-340, 2014.

H. Salehizadeh and S. Mohammadizad, Microbial enhanced oil recovery using biosurfactant produced by Alcaligenes faecalis, Iranian Journal of Biotechnology, vol.7, issue.4, pp.216-223, 2009.

A. Salihu, I. Abdulkadir, and M. N. Almustapha, An investigation for potential development on biosurfactants, Biotechnology and Molecular Biology Reviews, vol.4, issue.5, pp.111-117, 2009.

F. Salvà-serra, H. E. Jakobsson, A. Busquets, M. Gomila, D. Jaén-luchoro et al., Genome Sequences of Two Naphthalene-Degrading Strains of Pseudomonas balearica, Isolated from Polluted Marine Sediment and from an Oil Refinery Site, Genome announcements, vol.5, issue.14, pp.116-133, 2017.

S. Sana, S. Datta, D. Biswas, and D. Sengupta, Assessment of synergistic antibacterial activity of combined biosurfactants revealed by bacterial cell envelop damage, Biochimica et Biophysica Acta (BBA)-Biomembranes, vol.1860, issue.2, pp.579-585, 2018.

S. Santanu, A. Mazumder, S. Datta, and D. Biswas, Towards the development of an effective in vivo wound healing agent from Bacillus sp. derived biosurfactant using Catla catla fish fat, vol.7, pp.13668-13677, 2017.

D. Santos, R. Rufino, J. Luna, V. Santos, and L. Sarubbo, Biosurfactants: Multifunctional Biomolecules of the 21st Century, International Journal of Molecular Sciences, vol.17, issue.3, p.401, 2016.

A. Sarwar, M. N. Hassan, M. Imran, M. Iqbal, S. Majeed et al., Biocontrol activity of surfactin A purified from Bacillus NH-100 and NH-217 against rice bakanae disease, Microbiological Research, vol.209, pp.1-13, 2018.
DOI : 10.1016/j.micres.2018.01.006

URL : https://doi.org/10.1016/j.micres.2018.01.006

S. Sayadi, F. Zorgani, and R. Ellouz, Decolorization of olive mill waste-waters by free and immobilizedPhanerochaete chrysosporium cultures, Applied Biochemistry and Biotechnology, vol.56, issue.3, pp.265-276, 1996.
DOI : 10.1007/bf02786957

S. Schneiker, V. A. Santos, D. Bartels, T. Bekel, M. Brecht et al., Genome sequence of the ubiquitous hydrocarbon-degrading marine bacterium Alcanivorax borkumensis, Nature biotechnology, vol.24, issue.8, p.997, 2006.

B. Schuler, G. Meyer, D. Peña, O. C. Mullins, and L. Gross, Unraveling the Molecular Structures of Asphaltenes by Atomic Force Microscopy, Journal of the American Chemical Society, vol.137, issue.31, pp.9870-9876, 2015.

M. P. Schultz, J. A. Bendick, E. R. Holm, and W. M. Hertel, Economic impact of biofouling on a naval surface ship, Biofouling, vol.27, issue.1, pp.87-98, 2011.

K. K. Sekhon-randhawa and P. K. Rahman, Rhamnolipid biosurfactants-past, present, and future scenario of global market, Frontiers in microbiology, vol.5, p.454, 2014.

M. Sellami, A. Khlifi, F. Frikha, N. Miled, L. Belbahri et al., Agro-industrial waste based growth media optimization for biosurfactant production by Aneurinibacillus migulanus, Biotechnology and Food Sciences, vol.5, issue.6, pp.578-583, 2016.
DOI : 10.15414/jmbfs.2016.5.6.578-583

URL : https://www.jmbfs.org/wp-content/uploads/2016/05/jmbfs-1035-sellami.pdf

J. Seo, Y. Keum, and Q. Li, Bacterial degradation of aromatic compounds, International journal of environmental research and public health, vol.6, issue.1, pp.278-309, 2009.

E. ?epi?, M. Bricelj, and H. Leskov?ek, Degradation of fluoranthene by Pasteurella sp. IFA and Mycobacterium sp, Journal of applied microbiology, vol.85, issue.4, pp.746-754, 1998.

M. Serbaji, C. Azri, and K. Medhioub, Anthropogenic contributions to heavy metal distributions in the surface and sub-surface sediments of the northern coast of Sfax, Tunisia. International Journal of Environmental Research, vol.6, issue.3, pp.613-626, 2012.

M. Shahidul-islam and M. Tanaka, Impacts of pollution on coastal and marine ecosystems including coastal and marine fisheries and approach for management: a review and synthesis, Marine Pollution Bulletin, vol.48, issue.7, pp.624-649, 2004.

D. Sharma, B. S. Saharan, N. Chauhan, S. Procha, and S. Lal, Isolation and functional characterization of novel biosurfactant produced by Enterococcus faecium, SpringerPlus, vol.4, issue.1, p.4, 2015.
DOI : 10.1186/2193-1801-4-4

URL : https://springerplus.springeropen.com/track/pdf/10.1186/2193-1801-4-4

S. Shekhar, A. Sundaramanickam, and T. Balasubramanian, Biosurfactant Producing Microbes and their Potential Applications: A Review, Critical Reviews in Environmental Science and Technology, vol.45, issue.14, pp.1522-1554, 2015.
DOI : 10.1080/10643389.2014.955631

C. Shen, L. Jiang, H. Shao, C. You, G. Zhang et al., Targeted killing of myofibroblasts by biosurfactant di-rhamnolipid suggests a therapy against scar formation, Scientific Reports, vol.6, p.37553, 2016.

S. Shivaji, P. Chaturvedi, K. Suresh, G. S. Reddy, C. B. Dutt et al., Bacillus aerius sp. nov., Bacillus aerophilus sp. nov., Bacillus stratosphericus sp. nov. and Bacillus altitudinis sp. nov., isolated from cryogenic tubes used for collecting air samples from high altitudes, International Journal of Systematic and Evolutionary Microbiology, vol.56, issue.7, pp.1465-1473, 2006.

S. Shokralla, J. L. Spall, J. F. Gibson, and M. Hajibabaei, Next-generation sequencing technologies for environmental DNA research, Molecular ecology, vol.21, issue.8, pp.1794-1805, 2012.

M. A. Siddikee, P. S. Chauhan, R. Anandham, G. Han, and T. Sa, Isolation, characterization, and use for plant growth promotion under salt stress, of ACC deaminase-producing halotolerant bacteria derived from coastal soil, Journal of Microbiology and Biotechnology, vol.20, issue.1, pp.1577-1584, 2010.

S. N. Silva, C. B. Farias, R. D. Rufino, J. M. Luna, and L. A. Sarubbo, Glycerol as substrate for the production of biosurfactant by Pseudomonas aeruginosa UCP0992, Colloids and Surfaces B: Biointerfaces, vol.79, issue.1, pp.174-183, 2010.

S. L. Simonich, O. Motorykin, and N. Jariyasopit, PAH intermediates: Links between the atmosphere and biological systems, Chemico-Biological Interactions, vol.192, issue.1, pp.26-29, 2011.

D. N. Singh and A. K. Tripathi, Coal induced production of a rhamnolipid biosurfactant by Pseudomonas stutzeri, isolated from the formation water of Jharia coalbed, Bioresource technology, vol.128, pp.215-221, 2013.

P. Singh, Y. Patil, and V. Rale, Biosurfactant production: emerging trends and promising strategies, Journal of Applied Microbiology, vol.1, p.12

A. Souza, D. Rodriguez, D. Ribeaux, M. Luna, T. Lima-e-silva et al., Waste Soybean Oil and Corn Steep Liquor as Economic Substrates for Bioemulsifier and Biodiesel Production by Candida lipolytica UCP 0998, International Journal of Molecular Sciences, vol.17, issue.10, p.1608, 2016.

M. Srinivasa-reddy, S. Basha, H. V. Joshi, and G. Ramachandraiah, Seasonal distribution and contamination levels of total PHCs, PAHs and heavy metals in coastal waters of the AlangSosiya ship scrapping yard, Gulf of Cambay, India. Chemosphere, issue.11, pp.1587-1593, 2005.

M. I. Sriram, K. Kalishwaralal, V. Deepak, R. Gracerosepat, K. Srisakthi et al., Biofilm inhibition and antimicrobial action of lipopeptide biosurfactant produced by heavy metal tolerant strain Bacillus cereus NK1, Colloids and Surfaces B: Biointerfaces, vol.85, issue.2, pp.174-181, 2011.

T. Stipcevic, A. Piljac, and G. Piljac, Enhanced healing of full-thickness burn wounds using dirhamnolipid, Burns, vol.32, issue.1, pp.24-34, 2006.

P. Stoodley, K. Sauer, D. G. Davies, and J. W. Costerton, Biofilms as complex differentiated communities, Annual Reviews in Microbiology, vol.56, issue.1, pp.187-209, 2002.

W. A. Stubblefield, S. F. Brinkman, P. H. Davies, T. D. Garrison, J. R. Hockett et al., Effects of water hardness on the toxicity of manganese to developing brown trout (Salmo trutta), Environmental Toxicology and Chemistry, vol.16, issue.10, pp.2082-2089, 1997.

M. Y. Sun, K. A. Dafforn, M. V. Brown, and E. L. Johnston, Bacterial communities are sensitive indicators of contaminant stress, Marine Pollution Bulletin, vol.64, issue.5, pp.1029-1067, 2012.

H. Suthar and A. Nerurkar, Characterization of Biosurfactant Produced by Bacillus licheniformis TT42 Having Potential for Enhanced Oil Recovery, Applied Biochemistry and Biotechnology, vol.180, issue.2, pp.248-260, 2016.

D. Talluto, mpacts du déversement d'hydrocarbures sur les populations d'oiseaux marins, 2013.

K. Tamura, G. Stecher, and D. Peterson, MEGA6: molecular evolutionary genetics analysis version 6.0. Molecular Biology and Evolution, vol.30, pp.2725-2729, 2013.

T. Tan, B. Wang, and Z. Shao, Donghicola xiamenensis sp. nov., a marine bacterium isolated from seawater of the Taiwan Strait in China, International Journal of Systematic and Evolutionary Microbiology, vol.59, issue.5, pp.1143-1147, 2009.

A. Tapi, M. Chollet-imbert, B. Scherens, and P. Jacques, New approach for the detection of nonribosomal peptide synthetase genes in Bacillus strains by polymerase chain reaction, Applied microbiology and biotechnology, vol.85, issue.5, pp.1521-1531, 2010.

Y. H. Tapilatu, V. Grossi, M. Acquaviva, C. Militon, J. Bertrand et al., Isolation of hydrocarbon-degrading extremely halophilic archaea from an uncontaminated hypersaline pond, Extremophiles, vol.14, issue.2, pp.225-231, 2010.
URL : https://hal.archives-ouvertes.fr/insu-00687131

L. F. Tavares, P. M. Silva, M. Junqueira, D. C. Mariano, F. C. Nogueira et al., Characterization of rhamnolipids produced by wild-type and engineered Burkholderia kururiensis, Applied microbiology and biotechnology, vol.97, issue.5, pp.1909-1921, 2013.

M. Teramoto, M. Ohuchi, A. Hatmanti, Y. Darmayati, Y. Widyastuti et al., Oleibacter marinus gen. nov., sp. nov., a bacterium that degrades petroleum aliphatic hydrocarbons in a tropical marine environment, International Journal of Systematic and Evolutionary Microbiology, vol.61, issue.2, pp.375-380, 2011.

B. Thamdrup, D. E. Canfield, R. B. Sala, H. A. Jackson, R. W. Mooney et al., Benthic Respiration in Aquatic Sediments, Methods in Ecosystem Science, pp.86-103, 2000.

B. Tissot and D. Welte, Petroleum formation and occurance: A new approach to oil and gas exploration, 1978.

V. Torsvik, R. Sørheim, and J. Goksøyr, Total bacterial diversity in soil and sediment communities-a review, Journal of Industrial Microbiology, vol.17, issue.3-4, pp.170-178, 1996.

G. R. Tortella, M. C. Diez, and N. Durán, Fungal diversity and use in decomposition of environmental pollutants, Critical Reviews in Microbiology, vol.31, issue.4, pp.197-212, 2005.

T. and G. , ÉLÉMENTS DE GESTlON ÉCOSANlTAlRE DES ÉTANGS. La pisciculture en étang: actes du Congrès sur la Pisciculture en Étang, p.373, 1980.

J. D. Van-hamme, A. Singh, and O. P. Ward, Petroleum microbiology part 1: underlying biochemistry and physiology, Chimica oggi, issue.1, p.24, 2006.

E. Vasileva-tonkova, D. Galabova, E. Stoimenova, and Z. Lalchev, Production and properties of biosurfactants from a newly isolated Pseudomonas fluorescens HW-6 growing on hexadecane, Zeitschrift für Naturforschung C, pp.553-559, 2006.

V. Velma, S. Vutukuru, and P. B. Tchounwou, Ecotoxicology of hexavalent chromium in freshwater fish: a critical review, Reviews on environmental health, vol.24, issue.2, pp.129-146, 2009.

B. Vijaya, N. R. Jayalakshmi, and K. Manjunath, Enumeration of biosurfactant producing microorganisms from oil contaminated soil in and around Bangalore (India), International Journal of Current Science, vol.5, pp.86-94, 2013.

S. Vijayakumar and V. Saravanan, Biosurfactants-Types, Sources and Applications, Research Journal of Microbiology, vol.10, issue.5, p.181, 2015.

M. Vikas and G. Dwarakish, Coastal pollution: a review. Aquatic Procedia, vol.4, pp.381-388, 2015.

J. Vila, Z. López, J. Sabaté, C. Minguillón, A. M. Solanas et al., Identification of a novel metabolite in the degradation of pyrene by Mycobacterium sp. strain AP1: actions of the isolate on two-and three-ring polycyclic aromatic hydrocarbons, Applied and Environmental Microbiology, vol.67, issue.12, pp.5497-5505, 2001.

W. F. Vilela, S. G. Fonseca, F. Fantinatti-garboggini, V. M. Oliveira, and M. Nitschke, Production and Properties of a Surface-Active Lipopeptide Produced by a New Marine Brevibacterium luteolum Strain, Applied Biochemistry and Biotechnology, vol.174, issue.6, pp.2245-2256, 2014.

J. K. Volkman, D. G. Holdsworth, G. P. Neill, and H. Bavor, Identification of natural, anthropogenic and petroleum hydrocarbons in aquatic sediments, Science of the Total Environment, vol.112, issue.2-3, pp.203-219, 1992.

H. N. Wafi, Assessment of heavy metals contamination in the Mediterranean Sea along Gaza Coast-A case study of Gaza fishing harbor, 2015.

M. Wahl, F. Goecke, A. Labes, S. Dobretsov, and F. Weinberger, The Second Skin: Ecological Role of Epibiotic Biofilms on Marine Organisms, Frontiers in Microbiology, issue.292, p.3, 2012.

J. Wald, M. Hroudova, J. Jansa, B. Vrchotova, T. Macek et al., Pseudomonads rule degradation of polyaromatic hydrocarbons in aerated sediment, Frontiers in microbiology, vol.6, p.1268, 2015.

H. Wang and M. Frenklach, Transport properties of polycyclic aromatic hydrocarbons for flame modeling, Combustion and Flame, vol.96, issue.1, pp.163-170, 1994.

L. Wang, N. Qiao, F. Sun, and Z. Shao, Isolation, gene detection and solvent tolerance of benzene, toluene and xylene degrading bacteria from nearshore surface water and Pacific Ocean sediment, Extremophiles, vol.12, issue.3, pp.335-342, 2008.

W. Wang, L. Wang, and Z. Shao, Diversity and Abundance of Oil-Degrading Bacteria and Alkane Hydroxylase (alkB) Genes in the Subtropical Seawater of Xiamen Island, Microbial Ecology, vol.60, issue.2, pp.429-439, 2010.

D. M. Ward, C. M. Santegoeds, S. C. Nold, N. B. Ramsing, M. J. Ferris et al., Biodiversity within hot spring microbial mat communities: molecular monitoring of enrichment cultures, Antonie van Leeuwenhoek, vol.71, issue.1-2, pp.143-150, 1997.

D. Warshawsky, T. Cody, M. Radike, R. Reilman, B. Schumann et al., Biotransformation of benzo [a] pyrene and other polycyclic aromatic hydrocarbons and heterocyclic analogs by several green algae and other algal species under gold and white light, Chemico-biological interactions, vol.97, issue.2, pp.131-148, 1995.

L. Whang, P. G. Liu, C. Ma, and S. Cheng, Application of biosurfactants, rhamnolipid, and surfactin, for enhanced biodegradation of diesel-contaminated water and soil, Journal of Hazardous Materials, vol.151, issue.1, pp.155-163, 2008.

F. K. Wiese and P. C. Ryan, The extent of chronic marine oil pollution in southeastern Newfoundland waters assessed through beached bird surveys 1984-1999, Marine Pollution Bulletin, vol.46, issue.9, pp.1090-1101, 2003.

S. Winker and C. R. Woese, A definition of the domains Archaea, Bacteria and Eucarya in terms of small subunit ribosomal RNA characteristics, Systematic and Applied Microbiology, vol.14, issue.4, pp.305-310, 1991.

C. R. Woese, N. Won, K. Kim, J. H. Kang, S. R. Park et al., Exploring the impacts of anthropogenic disturbance on seawater and sediment microbial communities in Korean coastal waters using metagenomics analysis, International journal of environmental research and public health, vol.51, issue.2, p.130, 1987.

W. Wongwongsee, P. Chareanpat, and O. Pinyakong, Abilities and genes for PAH biodegradation of Bacteria isolated from mangrove sediments from the central of Thailand, Marine Pollution Bulltein, vol.74, issue.1, pp.95-104, 2013.

C. Wuchter, B. Abbas, M. J. Coolen, L. Herfort, J. Van-bleijswijk et al., Archaeal nitrification in the ocean, Proceedings of the National Academy of Sciences, vol.103, issue.33, pp.12317-12322, 2006.

W. Xia, Z. Luo, H. Dong, L. Yu, Q. Cui et al., Synthesis, Characterization, and Oil Recovery Application of Biosurfactant Produced by Indigenous Pseudomonas aeruginosa WJ-1 Using Waste Vegetable Oils, Applied Biochemistry and Biotechnology, vol.166, issue.5, pp.1148-1166, 2012.

Y. Xiao, W. Hui, Q. Wang, S. W. Roh, X. Shi et al., Pseudomonas caeni sp. nov., a denitrifying bacterium isolated from the sludge of an anaerobic ammoniumoxidizing bioreactor, International Journal of Systematic and Evolutionary Microbiology, vol.59, issue.10, pp.2594-2598, 2009.

H. X. Xu, H. Y. Wu, Y. P. Qiu, X. Q. Shi, G. H. He et al., Degradation of fluoranthene by a newly isolated strain of Herbaspirillum chlorophenolicum from activated sludge, Biodegradation, vol.22, issue.2, pp.335-345, 2011.

M. M. Yakimov, L. Giuliano, G. Gentile, E. Crisafi, T. N. Chernikova et al., Oleispira antarctica gen. nov., sp. nov., a novel hydrocarbonoclastic marine bacterium isolated from Antarctic coastal sea water, International Journal of Systematic and Evolutionary Microbiology, vol.53, issue.3, pp.779-785, 2003.

M. M. Yakimov, P. N. Golyshin, S. Lang, E. R. Moore, W. Abraham et al., Alcanivorax borkumensis gen. nov., sp. nov., a new, hydrocarbon-degrading and surfactant-producing marine bacterium, International Journal of Systematic and Evolutionary Microbiology, vol.48, issue.2, pp.339-348, 1998.

M. M. Yakimov, K. N. Timmis, V. Wray, and H. L. Fredrickson, Characterization of a new lipopeptide surfactant produced by thermotolerant and halotolerant subsurface Bacillus licheniformis BAS50, Applied and environmental microbiology, issue.5, pp.1706-1713, 1995.

X. Yan, J. Sims, B. Wang, and M. T. Hamann, Marine actinomycete Streptomyces sp. ISP2-49E, a new source of Rhamnolipid, Biochemical systematics and ecology, vol.55, pp.292-295, 2014.

Y. Yang, J. Wang, J. Liao, and S. Xie, Distribution of Naphtalene Dioxygenase Genes in Crude Oil-Contaminated Soils, Microbiol Ecology, vol.68, issue.4, pp.785-793, 2014.

H. Zaghden, M. Kallel, B. Elleuch, J. Oudot, and A. Saliot, Sources and distribution of aliphatic and polyaromatic hydrocarbons in sediments of Sfax, Marine Chemistry, vol.105, issue.1-2, pp.70-89, 2007.

H. Zaghden, M. M. Serbaji, A. Saliot, and S. Sayadi, The Tunisian Mediterranean coastline: potential threats from urban discharges Sfax-Tunisian Mediterranean coasts, Desalination and Water Treatment, vol.57, issue.52, pp.24765-24777, 2016.
URL : https://hal.archives-ouvertes.fr/hal-01498152

H. Zaghden, M. Tedetti, S. Sayadi, M. M. Serbaji, B. Elleuch et al., Origin and distribution of hydrocarbons and organic matter in the surficial sediments of the SfaxKerkennah channel, Marine pollution bulletin, vol.117, pp.414-428, 2017.
URL : https://hal.archives-ouvertes.fr/hal-01622004

B. Zenati, A. Chebbi, A. Badis, K. Eddouaouda, H. Boutoumi et al., A non-toxic microbial surfactant from Marinobacter hydrocarbonoclasticus SdK644 for crude oil solubilization enhancement, Ecotoxicology and Environmental Safety, vol.154, pp.100-107, 2018.

J. Zhang, E. Zhang, K. Scott, and J. G. Burgess, Enhanced Electricity Production by Use of Reconstituted Artificial Consortia of Estuarine Bacteria Grown as, Biofilms. Environmental Science & Technology, vol.46, issue.5, pp.2984-2992, 2012.

Z. Zhang and I. M. Lo, Biostimulation of petroleum-hydrocarbon-contaminated marine sediment with co-substrate: involved metabolic process and microbial community, Applied Microbiology and Biotechnology, vol.99, issue.13, pp.5683-96, 2015.

H. Zhou, J. Chen, Z. Yang, B. Qin, Y. Li et al., Biosurfactant production and characterization of Bacillus sp. ZG0427 isolated from oil-contaminated soil, Annals of Microbiology, vol.65, issue.4, pp.2255-2264, 2015.

X. Zhu, X. Ni, M. G. Waigi, J. Liu, K. Sun et al., Biodegradation of mixed PAHs by PAHdegrading endophytic bacteria, International journal of environmental research and public health, vol.13, issue.8, p.805, 2016.

W. Q. Zhuang, J. H. Tay, A. Maszenan, L. Krumholz, and S. L. Tay, Importance of Gram-positive naphthalene-degrading bacteria in oil-contaminated tropical marine sediments, Letters in applied microbiology, vol.36, issue.4, pp.251-257, 2003.

R. Zonta, M. Botter, D. Cassin, R. Pini, M. Scattolin et al., Sediment chemical contamination of a shallow water area close to the industrial zone of Porto Marghera, Marine Pollution Bulletin, vol.55, pp.529-542, 2007.

R. Zouari, D. Moalla-rekik, Z. Sahnoun, T. Rebai, S. Ellouze-chaabouni et al., Evaluation of dermal wound healing and in vitro antioxidant efficiency of Bacillus subtilis SPB1 biosurfactant, Biomedicine & Pharmacotherapy, vol.84, pp.878-891, 2016.

E. Zumstein, R. Moletta, and J. J. Godon, Examination of two years of community dynamics in an anaerobic bioreactor using fluorescence polymerase chain reaction (PCR) single-strand conformation polymorphism analysis, Environmental Microbiology, vol.2, issue.1, pp.69-78, 2000.