B. Aouizerate, D. Guehl, E. Cuny, A. Rougier, B. Bioulac et al., Pathophysiology of obsessive-compulsive disorder: a necessary link between phenomenology, neuropsychology, imagery and physiology, Prog Neurobiol, vol.72, pp.195-221, 2004.

V. J. Brown and E. M. Bowman, Discriminative cues indicating reward magnitude continue to determine reaction time of rats following lesions of the nucleus accumbens, Eur J Neurosci, vol.7, pp.2479-2485, 1995.

L. Carbonnell, C. Ramdani, C. Meckler, B. Burle, T. Hasbroucq et al., The N-40: an electrophysiological marker of response selection, Biol Psychol, vol.93, pp.231-236, 2013.
URL : https://hal.archives-ouvertes.fr/hal-01384090

J. Costermans, G. Lories, and C. Ansay, Confidence level and feeling of knowing in question answering: The weight of inferential processes, Journal of Experimental Psychology: Learning, Memory and Cognition, vol.18, issue.1, pp.142-150, 1992.

L. A. Fairbanks, M. J. Jorgensen, J. N. Bailey, S. E. Breidenthal, R. Grzywa et al., Heritability and genetic correlation of hair cortisol in vervet monkeys in low and higher stress environments, Psychoneuroendocrinology, vol.36, pp.1201-1208, 2011.

C. R. Fetsch, R. Kiani, W. T. Newsome, and M. N. Shadlen, Effects of cortical microstimulation on confidence in a perceptual decision, Neuron, vol.83, pp.797-804, 2014.

J. R. Folstein and C. Van-petten, Influence of cognitive control and mismatch on the N2 component of the ERP: a review, Psychophysiology, vol.45, pp.152-170, 2008.

R. O. Frost, K. J. Sher, and T. Geen, Psychopathology and personality characteristics of nonclinical compulsive checkers, Behav Res Ther, vol.24, pp.133-143, 1986.
DOI : 10.1016/0005-7967(86)90084-7

E. Fuchs, C. Kirschbaum, D. Benisch, and A. Bieser, Salivary cortisol: a non-invasive measure of hypothalamo-pituitary-adrenocortical activity in the squirrel monkey, Saimiri sciureus, Lab Anim, vol.31, pp.306-311, 1997.

M. A. Fullana, D. Mataix-cols, A. Caspi, H. Harrington, J. R. Grisham et al.,

, Obsessions and compulsions in the community: prevalence, interference, help-seeking, developmental stability, and co-occurring psychiatric conditions, Am J Psychiatry, vol.166, pp.329-336

C. Garcia-leal, A. C. Parente, C. M. Del-ben, F. S. Guimaraes, A. C. Moreira et al.,

, Anxiety and salivary cortisol in symptomatic and nonsymptomatic panic patients and healthy volunteers performing simulated public speaking, Psychiatry Res, vol.133, pp.239-252

W. J. Gehring, J. Himle, and L. G. Nisenson, Action-monitoring dysfunction in obsessivecompulsive disorder, Psychol Sci, vol.11, pp.1-6, 2000.
DOI : 10.1111/1467-9280.00206

B. S. Gershuny and K. J. Sher, Compulsive checking and anxiety in a nonclinical sample: Differences in cognition, behavior, personality, and affect, Journal of Psychopathology and Behavioral Assessment, vol.17, pp.19-38, 1995.

A. M. Graybiel and S. L. Rauch, Toward a neurobiology of obsessive-compulsive disorder, Neuron, vol.28, pp.343-347, 2000.

R. R. Hampton, Multiple demonstrations of metacognition in nonhumans: Converging evidence or multiple mechanisms?, Comp Cogn Behav Rev, vol.4, pp.17-28, 2009.

R. R. Hampton, A. Zivin, and E. A. Murray, Rhesus monkeys (Macaca mulatta) discriminate between knowing and not knowing and collect information as needed before acting, Anim Cogn, vol.7, pp.239-246, 2004.

J. P. Higham, A. B. Vitale, A. M. Rivera, J. E. Ayala, and D. Maestripieri, Measuring salivary analytes from free-ranging monkeys, Physiol Behav, vol.101, pp.601-607, 2010.

S. Johannes, B. M. Wieringa, W. Nager, D. Rada, R. Dengler et al., Discrepant target detection and action monitoring in obsessive-compulsive disorder, Psychiatry Res, vol.108, pp.101-110, 2001.

R. Johnson and E. Donchin, On how P300 amplitude varies with the utility of the eliciting stimuli, Electroencephalogr Clin Neurophysiol, vol.44, pp.424-437, 1978.

A. Kepecs and Z. F. Mainen, A computational framework for the study of confidence in humans and animals, Philos Trans R Soc Lond B Biol Sci, vol.367, pp.1322-1337, 2012.

A. Kepecs, N. Uchida, H. A. Zariwala, and Z. F. Mainen, Neural correlates, computation and behavioural impact of decision confidence, Nature, vol.455, pp.227-231, 2008.

R. Kiani and M. N. Shadlen, Representation of confidence associated with a decision by neurons in the parietal cortex, Science, vol.324, pp.759-764, 2009.

C. Kirschbaum and D. H. Hellhammer, Salivary cortisol in psychobiological research: an overview, Neuropsychobiology, vol.22, pp.150-169, 1989.

Y. Komura, A. Nikkuni, N. Hirashima, T. Uetake, and A. Miyamoto, Responses of pulvinar neurons reflect a subject's confidence in visual categorization, Nat Neurosci, vol.16, pp.749-755, 2013.

N. Kornell, L. K. Son, and H. S. Terrace, Transfer of metacognitive skills and hint seeking in monkeys, Psychol Sci, vol.18, pp.64-71, 2007.

C. K. Lutz, S. Tiefenbacher, M. J. Jorgensen, J. S. Meyer, and M. A. Novak, Techniques for collecting saliva from awake, unrestrained, adult monkeys for cortisol assay, Am J Primatol, vol.52, pp.93-99, 2000.

N. Maltby, D. F. Tolin, P. Worhunsky, O. Keefe, T. M. Kiehl et al., Dysfunctional action monitoring hyperactivates frontal-striatal circuits in obsessive-compulsive disorder: an eventrelated fMRI study, Neuroimage, vol.24, pp.495-503, 2005.

T. Michelet, B. Bioulac, N. Langbour, M. Goillandeau, D. Guehl et al., Electrophysiological Correlates of a Versatile Executive Control System in the Monkey Anterior Cingulate Cortex, Cereb Cortex, vol.26, pp.1684-1697, 2016.
URL : https://hal.archives-ouvertes.fr/hal-01155375

P. G. Middlebrooks and M. A. Sommer, Metacognition in monkeys during an oculomotor task, J Exp Psychol Learn Mem Cogn, vol.37, pp.325-337, 2011.

T. Nakao, A. Nakagawa, T. Yoshiura, E. Nakatani, M. Nabeyama et al., Brain activation of patients with obsessive-compulsive disorder during neuropsychological and symptom provocation tasks before and after symptom improvement: A functional magnetic resonance imaging study, Biological Psychiatry, vol.57, pp.901-910, 2005.

S. Nieuwenhuis, N. Yeung, W. Van-den-wildenberg, and K. R. Ridderinkhof, Electrophysiological correlates of anterior cingulate function in a go/no-go task: effects of response conflict and trial type frequency, Cogn Affect Behav Neurosci, vol.3, pp.17-26, 2003.

. Oxforddictionaries, Check-definition of check in English from the Oxford dictionary, 2016.

B. L. Pearson, P. G. Judge, and D. M. Reeder, Effectiveness of saliva collection and enzymeimmunoassay for the quantification of cortisol in socially housed baboons, Am J Primatol, vol.70, pp.1145-1151, 2008.

S. Rachman, A cognitive theory of compulsive checking, Behav Res Ther, vol.40, pp.625-639, 2002.

N. Ramnani and R. C. Miall, Instructed delay activity in the human prefrontal cortex is modulated by monetary reward expectation, Cereb Cortex, vol.13, pp.318-327, 2003.

S. L. Rauch, M. A. Jenike, N. M. Alpert, L. Baer, H. C. Breiter et al., Regional cerebral blood flow measured during symptom provocation in obsessive-compulsive disorder using oxygen 15-labeled carbon dioxide and positron emission tomography, Arch Gen Psychiatry, vol.51, pp.62-70, 1994.

G. Roper, S. Rachman, and R. Hodgson, An experiment on obsessional checking, Behav Res Ther, vol.11, pp.271-277, 1973.

J. Y. Rotge, D. Guehl, B. Dilharreguy, E. Cuny, J. Tignol et al., Provocation of obsessive-compulsive symptoms: a quantitative voxelbased meta-analysis of functional neuroimaging studies, J Psychiatry Neurosci, vol.33, pp.405-412, 2008.

J. Y. Rotge, N. Langbour, B. Dilharreguy, M. Bordessoulles, D. Guehl et al., Contextual and behavioral influences on uncertainty in obsessive-compulsive disorder, vol.62, pp.1-10, 2015.
URL : https://hal.archives-ouvertes.fr/hal-01155377

W. E. Shields, J. D. Smith, K. Guttmannova, and D. A. Washburn, Confidence judgments by humans and rhesus monkeys, J Gen Psychol, vol.132, pp.165-186, 2005.

W. E. Shields, J. D. Smith, and D. A. Washburn, Uncertain responses by humans and rhesus monkeys (Macaca mulatta) in a psychophysical same-different task, J Exp Psychol Gen, vol.126, pp.147-164, 1997.

J. D. Smith, J. Schull, J. Strote, K. Mcgee, R. Egnor et al., The uncertain response in the bottlenosed dolphin (Tursiops truncatus), J Exp Psychol Gen, vol.124, pp.391-408, 1995.

J. D. Smith, W. E. Shields, and D. A. Washburn, The comparative psychology of uncertainty monitoring and metacognition, Behav Brain Sci, vol.26, pp.340-373, 2003.

S. M. Stasik, K. Naragon-gainey, M. Chmielewski, and D. Watson, Core OCD symptoms: exploration of specificity and relations with psychopathology, J Anxiety Disord, vol.26, pp.859-870, 2012.

H. Szechtman, S. E. Ahmari, R. J. Beninger, D. Eilam, B. H. Harvey et al., Obsessive-compulsive disorder: Insights from animal models, Neurosci Biobehav Rev, 2016.

A. Szmalec, F. Verbruggen, A. Vandierendonck, W. De-baene, T. Verguts et al., Stimulus ambiguity elicits response conflict, Neurosci Lett, vol.435, pp.158-162, 2008.

S. Ursu, V. A. Stenger, M. K. Shear, M. R. Jones, C. Carter et al., Overactive action monitoring in obsessive-compulsive disorder: evidence from functional magnetic resonance imaging, Psychol Sci, vol.14, pp.853-868, 2003.

M. Watanabe, H. C. Cromwell, L. Tremblay, J. R. Hollerman, K. Hikosaka et al., Behavioral reactions reflecting differential reward expectations in monkeys, Exp Brain Res, vol.140, pp.511-518, 2001.

M. L. Welter, P. Burbaud, S. Fernandez-vidal, E. Bardinet, J. Coste et al., French Stimulation dans Trouble Obsessionnel Compulsif Study G. 2011. Basal ganglia dysfunction in OCD: subthalamic neuronal activity correlates with symptoms severity and predicts high-frequency stimulation efficacy, Transl Psychiatry, vol.1, p.5

G. F. Woodman, M. S. Kang, A. F. Rossi, and J. D. Schall, Nonhuman primate event-related potentials indexing covert shifts of attention, Proc Natl Acad Sci U S A, vol.104, pp.15111-15116, 2007.

A. Yildiz, O. T. Wolf, and C. Beste, Stress intensifies demands on response selection during action cascading processes, Psychoneuroendocrinology, vol.42, pp.178-187, 2014.

M. Abbruzzese, L. Bellodi, S. Ferri, and S. Scarone, Memory functioning in obsessivecompulsive disorder, Behavioural neurology, vol.6, pp.119-122, 1993.

D. Akkal, R. P. Dum, and P. L. Strick, Supplementary Motor Area and Presupplementary Motor Area: Targets of Basal Ganglia and Cerebellar Output, Journal of Neuroscience, vol.27, pp.10659-10673, 2007.

D. Akkal, B. Bioulac, J. Audin, and P. Burbaud, Comparison of neuronal activity in the rostral supplementary and cingulate motor areas during a task with cognitive and motor demands, Eur J Neurosci, vol.15, pp.887-904, 2002.

N. Albelda and J. D. , Current animal models of obsessive compulsive disorder: an update, NSC, vol.211, pp.83-106, 2012.

G. M. Alcolado and A. S. Radomsky, Believe in yourself: Manipulating beliefs about memory causes checking, Behaviour Research and Therapy, vol.49, pp.42-49, 2011.

G. E. Alexander and M. D. Crutcher, Preparation for movement: neural representations of intended direction in three motor areas of the monkey, J Neurophysiol, vol.64, pp.133-150, 1990.

W. H. Alexander and J. W. Brown, Medial prefrontal cortex as an action-outcome predictor, Nature Neuroscience, vol.14, pp.1338-1344, 2011.

A. H. Alkhatib, A. Dvorkin-gheva, and H. Szechtman, Quinpirole and 8-OH-DPAT induce compulsive checking behavior in male rats by acting on different functional parts of an OCD neurocircuit, Behav Pharmacol, vol.24, pp.65-73, 2013.

S. Amemori, . Amemori-k-i, M. L. Cantor, and A. M. Graybiel, A non-invasive head-holding device for chronic neural recordings in awake behaving monkeys, Journal of Neuroscience Methods, vol.240, pp.154-160, 2015.

, American Psychiatric Association (1980) Diagnostic and statistical manual of mental disorders, vol.3

, Diagnostic and statistical manual of mental disorders, American Psychiatric Association, 1994.

, Diagnostic and statistical manual of mental disorders (DSM-5®), American Psychiatric Association, 2013.

C. Amiez, Reward Encoding in the Monkey Anterior Cingulate Cortex, Cerebral Cortex, vol.16, pp.1040-1055, 2005.
URL : https://hal.archives-ouvertes.fr/inserm-00132137

C. Amiez and M. Petrides, Neuroimaging Evidence of the Anatomo-Functional Organization of the Human Cingulate Motor Areas, Cerebral Cortex, vol.24, pp.563-578, 2014.

C. Amiez, J. Joseph, and E. Procyk, Anterior cingulate error-related activity is modulated by predicted reward, European Journal of Neuroscience, vol.21, pp.3447-3452, 2005.
URL : https://hal.archives-ouvertes.fr/hal-00131438

B. Aouizerate, D. Guehl, E. Cuny, A. Rougier, B. Bioulac et al., Pathophysiology of obsessive-compulsive disorder: a necessary link between phenomenology, neuropsychology, imagery and physiology, Prog Neurobiol, vol.72, pp.195-221, 2004.

A. R. Aron, Progress in executive-function research: From tasks to functions to regions to networks, Current Directions in Psychological Science, 2008.

B. M. Basile, R. R. Hampton, S. J. Suomi, and E. A. Murray, An assessment of memory awareness in tufted capuchin monkeys (Cebus apella), Anim Cogn, vol.12, pp.169-180, 2009.

J. F. Bates and P. S. Goldman-rakic, Prefrontal connections of medial motor areas in the rhesus monkey, The Journal of Comparative Neurology, vol.336, pp.211-228, 1993.

S. Behseta, R. E. Kass, D. E. Moorman, and C. R. Olson, Testing equality of several functions: Analysis of single-unit firing-rate curves across multiple experimental conditions, Statistics in Medicine, vol.26, pp.3958-3975, 2007.

C. Belzung and P. Philippot, Anxiety from a Phylogenetic Perspective: Is there a Qualitative Difference between Human and Animal Anxiety?, Neural Plasticity, pp.1-17, 2007.

M. J. Beran and J. D. Smith, Information seeking by rhesus monkeys (Macaca mulatta) and capuchin monkeys (Cebus apella), Cognition, vol.120, pp.90-105, 2011.

M. J. Beran, B. M. Perdue, and J. D. Smith, What are my chances? Closing the gap in uncertainty monitoring between rhesus monkeys (Macaca mulatta) and capuchin monkeys (Cebus apella), Journal of Experimental Psychology: Animal Learning and Cognition, vol.40, pp.303-316, 2014.

M. H. Bloch, A. Landeros-weisenberger, M. C. Rosario, C. Pittenger, and J. F. Leckman, Metaanalysis of the symptom structure of obsessive-compulsive disorder, The American journal of psychiatry, vol.165, pp.1532-1542, 2008.

M. Botvinick, L. E. Nystrom, K. Fissell, C. S. Carter, and J. D. Cohen, Conflict monitoring versus selection-for-action in anterior cingulate cortex, Nature, vol.402, pp.179-181, 1999.

M. M. Botvinick, J. D. Cohen, and C. S. Carter, Conflict monitoring and anterior cingulate cortex: an update, Trends in Cognitive Sciences, vol.8, pp.539-546, 2004.

M. M. Botvinick, T. S. Braver, D. M. Barch, C. S. Carter, and J. D. Cohen, Conflict monitoring and cognitive control, Psychological Review, vol.108, p.624, 2001.

C. Bouras, E. Kovari, P. R. Hof, B. M. Riederer, and P. Giannakopoulos, Anterior cingulate cortex pathology in schizophrenia and bipolar disorder, Acta Neuropathol, vol.102, pp.373-379, 2001.

J. Bräuer, J. Call, and M. Tomasello, Visual perspective taking in dogs (Canis familiaris) in the presence of barriers, Applied Animal Behaviour Science, vol.88, pp.299-317, 2004.

V. B. Brooks, How does the limbic system assist motor learning? A limbic comparator hypothesis, Brain Behav Evol, vol.29, pp.29-53, 1986.

J. W. Brown, Conflict effects without conflict in anterior cingulate cortex: Multiple response effects and context specific representations, NeuroImage, vol.47, pp.334-341, 2009.

T. A. Brown, K. Moras, R. E. Zinbarg, and D. H. Barlow, Diagnostic and symptom distinguishability of generalized anxiety disorder and obsessive-compulsive disorder, Behavior Therapy, vol.24, pp.227-240, 1993.

V. J. Brown and E. M. Bowman, Discriminative cues indicating reward magnitude continue to determine reaction time of rats following lesions of the nucleus accumbens, Eur J Neurosci, vol.7, pp.2479-2485, 1995.

D. W. Bryden, E. E. Johnson, S. C. Tobia, V. Kashtelyan, and M. R. Roesch, Attention for Learning Signals in Anterior Cingulate Cortex, Journal of Neuroscience, vol.31, pp.18266-18274, 2011.

S. Budhani, A. A. Marsh, D. S. Pine, and R. Blair, Neural correlates of response reversal: Considering acquisition, NeuroImage, vol.34, pp.1754-1765, 2007.

A. Buge, R. Escourolle, G. Rancurel, and M. Poisson, Akinetic mutism and bicingular softening. 3 anatomo-clinical cases, Rev Neurol, vol.131, pp.121-131, 1975.

P. Burbaud, Neuronal activity correlated with checking behaviour in the subthalamic nucleus of patients with obsessive-compulsive disorder, Brain, vol.136, pp.304-317, 2013.
URL : https://hal.archives-ouvertes.fr/hal-00817570

G. Bush, B. A. Vogt, J. Holmes, A. M. Dale, D. Greve et al., Dorsal anterior cingulate cortex: a role in reward-based decision making, Proceedings of the National Academy of Sciences of the United States of America, vol.99, pp.523-528, 2002.

J. Call and M. Carpenter, Do apes and children know what they have seen?, Anim Cogn, vol.3, pp.207-220, 2001.

L. Carbonnell, C. Ramdani, C. Meckler, B. Burle, T. Hasbroucq et al., The N-40: an electrophysiological marker of response selection, Biol Psychol, vol.93, pp.231-236, 2013.
URL : https://hal.archives-ouvertes.fr/hal-01384090

P. Carruthers, Meta-cognition in animals: A skeptical look, Mind & Language, vol.23, pp.58-89, 2008.

C. S. Carter, T. S. Braver, D. M. Barch, M. M. Botvinick, D. Noll et al., Anterior cingulate cortex, error detection, and the online monitoring of performance, Science, vol.280, pp.747-749, 1998.

J. F. Cavanagh, T. O. Gründler, M. J. Frank, and J. J. Allen, Altered cingulate sub-region activation accounts for task-related dissociation in ERN amplitude as a function of obsessive-compulsive symptoms, Neuropsychologia, vol.48, pp.2098-2109, 2010.

S. R. Chamberlain, A. D. Blackwell, N. A. Fineberg, T. W. Robbins, and B. J. Sahakian, The neuropsychology of obsessive compulsive disorder: the importance of failures in cognitive and behavioural inhibition as candidate endophenotypic markers, Neuroscience & Biobehavioral Reviews, vol.29, pp.399-419, 2005.

S. Chassagnon, L. Minotti, S. Kremer, D. Hoffmann, and P. Kahane, Somatosensory, motor, and reaching/grasping responses to direct electrical stimulation of the human cingulate motor areas, Journal of Neurosurgery, vol.109, pp.593-604, 2008.
URL : https://hal.archives-ouvertes.fr/inserm-00378396

M. G. Coles, M. K. Scheffers, and C. B. Holroyd, Why is there an ERN/Ne on correct trials? Response representations, stimulus-related components, and the theory of errorprocessing, Biological Psychology, vol.56, pp.173-189, 2001.

J. Costermans, G. Lories, and C. Ansay, Confidence level and feeling of knowing in question answering: The weight of inferential processes, Journal of Experimental Psychology: Learning, Memory and Cognition, vol.18, issue.1, pp.142-150, 1992.

J. J. Couchman, M. Coutinho, M. J. Beran, and J. D. Smith, Beyond stimulus cues and reinforcement signals: A new approach to animal metacognition, Journal of Comparative Psychology, vol.124, pp.356-368, 2010.

J. D. Crystal and A. L. Foote, Evaluating information-seeking approaches to metacognition, Current zoology, vol.57, pp.531-542, 2011.

R. Cunnington, C. Windischberger, L. Deecke, and E. Moser, The Preparation and Execution of Self-Initiated and Externally-Triggered Movement: A Study of Event-Related fMRI, NeuroImage, vol.15, pp.373-385, 2002.

C. Cuttler and P. Graf, Checking-in on the memory deficit and meta-memory deficit theories of compulsive checking, Clinical psychology review, vol.29, pp.393-409, 2009.

C. Cuttler, V. Sirois-delisle, G. M. Alcolado, A. S. Radomsky, and S. Taylor, Diminished confidence in prospective memory causes doubts and urges to check, Journal of Behavior Therapy and Experimental Psychiatry, vol.44, pp.329-334, 2013.

S. Debener, Trial-by-Trial Coupling of Concurrent Electroencephalogram and Functional Magnetic Resonance Imaging Identifies the Dynamics of Performance Monitoring, Journal of Neuroscience, vol.25, pp.11730-11737, 2005.

L. Deecke and H. H. Kornhuber, An electrical sign of participation of the mesial 'supplementary' motor cortex in human voluntary finger movement, vol.2, pp.473-476, 1978.

M. P. Deiber, M. Honda, V. Ibañez, N. Sadato, and M. Hallett, Mesial motor areas in selfinitiated versus externally triggered movements examined with fMRI: effect of movement type and rate, Journal of Neurophysiology, vol.81, pp.3065-3077, 1999.

D. Sala, S. Marchetti, C. Spinnler, and H. , Right-sided anarchic (alien) hand: a longitudinal study, Neuropsychologia, vol.29, pp.1113-1127, 1991.

O. Devinsky, M. J. Morrell, and B. A. Vogt, Contributions of anterior cingulate cortex to behaviour, Brain, vol.118, pp.279-306, 1995.

M. Donoso, A. Collins, and E. Koechlin, Foundations of human reasoning in the prefrontal cortex, Science, vol.344, pp.1481-1486, 2014.

R. P. Dum and P. L. Strick, The origin of corticospinal projections from the premotor areas in the frontal lobe, The Journal of neuroscience : the official journal of the Society for Neuroscience, vol.11, pp.667-689, 1991.

R. P. Dum and P. L. Strick, Motor areas in the frontal lobe of the primate, Physiol Behav, vol.77, pp.677-682, 2003.

D. Durstewitz, N. M. Vittoz, S. B. Floresco, and J. K. Seamans, Abrupt Transitions between Prefrontal Neural Ensemble States Accompany Behavioral Transitions during Rule Learning, Neuron, vol.66, pp.438-448, 2010.

D. M. Eagle, C. Noschang, L. Angelo, C. Noble, J. O. Day et al., The dopamine D2/D3 receptor agonist quinpirole increases checking-like behaviour in an operant observing response task with uncertain reinforcement: A novel possible model of OCD, Behavioural Brain Research, pp.1-23, 2014.

R. B. Ebitz and M. L. Platt, Neuronal Activity in Primate Dorsal Anterior Cingulate Cortex Signals Task Conflict and Predicts Adjustments in Pupil-Linked Arousal, Neuron, vol.85, pp.628-640, 2015.

M. Economides, M. Guitart-masip, Z. Kurth-nelson, and R. J. Dolan, Anterior Cingulate Cortex Instigates Adaptive Switches in Choice by Integrating Immediate and Delayed Components of Value in Ventromedial Prefrontal Cortex, Journal of Neuroscience, vol.34, pp.3340-3349, 2014.

D. Eilam and H. Szechtman, Psychostimulant-Induced Behavior as an Animal Model of Obsessive-Compulsive Disorder: An Ethological Approach to the Form of Compulsive Rituals, CNS Spectrums, vol.10, pp.191-202, 2005.

D. Eilam, R. Zor, H. Szechtman, and H. Hermesh, Rituals, stereotypy and compulsive behavior in animals and humans, Neuroscience & Biobehavioral Reviews, vol.30, pp.456-471, 2006.

E. E. Emeric, J. W. Brown, M. Leslie, P. Pouget, V. Stuphorn et al., Performance Monitoring Local Field Potentials in the Medial Frontal Cortex of Primates: Anterior Cingulate Cortex, Journal of Neurophysiology, vol.99, pp.759-772, 2008.

T. Endrass and M. Ullsperger, Specificity of performance monitoring changes in obsessivecompulsive disorder, Neurosci Biobehav Rev, vol.46, pp.124-138, 2014.

L. A. Fairbanks, M. J. Jorgensen, J. N. Bailey, S. E. Breidenthal, R. Grzywa et al., Heritability and genetic correlation of hair cortisol in vervet monkeys in low and higher stress environments, Psychoneuroendocrinology, vol.36, pp.1201-1208, 2011.

M. Falkenstein, J. Hohnsbein, J. Hoormann, and L. Blanke, Effects of crossmodal divided attention on late ERP components. II. Error processing in choice reaction tasks, Electroencephalogr Clin Neurophysiol, vol.78, pp.447-455, 1991.

J. R. Fedota, J. E. Hardee, K. Pérez-edgar, and J. C. Thompson, Representation of response alternatives in human presupplementary motor area_ Multi-voxel pattern analysis in a go/no-go task, Neuropsychologia, vol.56, pp.110-118, 2014.

C. R. Fetsch, R. Kiani, W. T. Newsome, and M. N. Shadlen, Effects of cortical microstimulation on confidence in a perceptual decision, Neuron, vol.83, pp.797-804, 2014.

K. D. Fitzgerald, R. C. Welsh, W. J. Gehring, J. L. Abelson, J. A. Himle et al., Error-related hyperactivity of the anterior cingulate cortex in obsessive-compulsive disorder, Biological Psychiatry, vol.57, pp.287-294, 2005.

J. R. Folstein and C. Van-petten, Influence of cognitive control and mismatch on the N2 component of the ERP: a review, Psychophysiology, vol.45, pp.152-170, 2008.

A. L. Foote and J. D. Crystal, Metacognition in the Rat, Current Biology, vol.17, pp.551-555, 2007.

I. Fried, A. Katz, G. Mccarthy, K. J. Sass, P. Williamson et al., Functional organization of human supplementary motor cortex studied by electrical stimulation, The Journal of neuroscience : the official journal of the Society for Neuroscience, vol.11, pp.3656-3666, 1991.

K. J. Friston, C. D. Frith, R. E. Passingham, P. F. Liddle, and R. S. Frackowiak, Motor practice and neurophysiological adaptation in the cerebellum: a positron tomography study, Proc Biol Sci, vol.248, pp.223-228, 1992.

R. O. Frost, K. J. Sher, and T. Geen, Psychopathology and personality characteristics of nonclinical compulsive checkers, Behav Res Ther, vol.24, pp.133-143, 1986.

E. Fuchs, C. Kirschbaum, D. Benisch, and A. Bieser, Salivary cortisol: a non-invasive measure of hypothalamo-pituitary-adrenocortical activity in the squirrel monkey, Saimiri sciureus, Lab Anim, vol.31, pp.306-311, 1997.

M. A. Fullana, D. Mataix-cols, A. Caspi, H. Harrington, J. R. Grisham et al., Obsessions and compulsions in the community: prevalence, interference, helpseeking, developmental stability, and co-occurring psychiatric conditions, Am J Psychiatry, vol.166, pp.329-336, 2009.

G. J. Gage, H. Parikh, and T. C. Marzullo, The cingulate cortex does everything, Annals of Improbable Research, vol.3, p.55, 2008.

H. Garavan, T. J. Ross, J. Kaufman, and E. A. Stein, A midline dissociation between errorprocessing and response-conflict monitoring, NeuroImage, vol.20, pp.1132-1139, 2003.

C. Garcia-leal, A. C. Parente, C. M. Del-ben, F. S. Guimaraes, A. C. Moreira et al., Anxiety and salivary cortisol in symptomatic and nonsymptomatic panic patients and healthy volunteers performing simulated public speaking, Psychiatry Res, vol.133, pp.239-252, 2005.

W. J. Gehring, The Error-Related Negativity:An Event-Related Brain Potential Accompanying Errors, vol.27, pp.1-77, 1990.

W. J. Gehring, J. Himle, and L. G. Nisenson, Action-monitoring dysfunction in obsessivecompulsive disorder, Psychol Sci, vol.11, pp.1-6, 2000.

H. Gemba, K. Sasaki, and V. B. Brooks, Error' potentials in limbic cortex (anterior cingulate area 24) of monkeys during motor learning, Neuroscience Letters, vol.70, pp.223-227, 1986.

C. Gerloff, B. Corwell, R. Chen, M. Hallett, and L. G. Cohen, Stimulation over the human supplementary motor area interferes with the organization of future elements in complex motor sequences, Brain, vol.120, pp.1587-1602, 1997.

B. S. Gershuny and K. J. Sher, Compulsive checking and anxiety in a nonclinical sample: Differences in cognition, behavior, personality, and affect, Journal of Psychopathology and Behavioral Assessment, vol.17, pp.19-38, 1995.

W. K. Goodman, L. H. Price, S. A. Rasmussen, C. Mazure, P. Delgado et al., The Yale-Brown Obsessive Compulsive Scale. II. Validity, Archives of general psychiatry, vol.46, pp.1012-1016, 1989.

W. K. Goodman, L. H. Price, S. A. Rasmussen, C. Mazure, R. L. Fleischmann et al., The Yale-Brown Obsessive Compulsive Scale. I. Development, use, and reliability, Archives of general psychiatry, vol.46, pp.1006-1011, 1989.

A. M. Graybiel and S. L. Rauch, Toward a neurobiology of obsessive-compulsive disorder, Neuron, vol.28, pp.343-347, 2000.

M. Graziano and T. N. Aflalo, Mapping Behavioral Repertoire onto the Cortex, Neuron, vol.56, pp.239-251, 2007.

R. Grützmann, T. Endrass, C. Kaufmann, A. E. Eichele, T. Kathmann et al., Presupplementary Motor Area Contributes to Altered Error Monitoring in ObsessiveCompulsive Disorder, vol.BPS, pp.1-10, 2015.

U. Halsband, Y. Matsuzaka, and J. Tanji, Neuronal activity in the primate supplementary, pre-supplementary and premotor cortex during externally and internally instructed sequential movements, Neuroscience research, vol.20, pp.149-155, 1994.

R. R. Hampton, Rhesus monkeys know when they remember, Proc Natl Acad Sci, vol.98, pp.5359-5362, 2001.

R. R. Hampton, Multiple demonstrations of metacognition in nonhumans: Converging evidence or multiple mechanisms?, Comp Cogn Behav Rev, vol.4, pp.17-28, 2009.

R. R. Hampton, A. Zivin, and E. A. Murray, Rhesus monkeys (Macaca mulatta) discriminate between knowing and not knowing and collect information as needed before acting, Anim Cogn, vol.7, pp.239-246, 2004.

J. P. Higham, A. B. Vitale, A. M. Rivera, J. E. Ayala, and D. Maestripieri, Measuring salivary analytes from free-ranging monkeys, Physiol Behav, vol.101, pp.601-607, 2010.

O. Hikosaka and M. Isoda, Switching from automatic to controlled behavior: cortico-basal ganglia mechanisms, Trends Cogn Sci, vol.14, pp.154-161, 2010.

A. L. Hinds, E. Z. Woody, M. Van-ameringen, L. A. Schmidt, and H. Szechtman, When Too Much Is Not Enough: Obsessive-Compulsive Disorder as a Pathology of Stopping, Rather than Starting, Plos ONE, vol.7, p.30586, 2012.

R. J. Hodgson and S. Rachman, Obsessional-compulsive complaints, Behaviour Research and Therapy, vol.15, pp.389-395, 1977.

E. Hollander, A. Braun, and D. Simeon, Should OCD leave the anxiety disorders in DSM-V? The case for obsessive compulsive-related disorders, Depression and Anxiety, vol.25, pp.317-329, 2008.

C. B. Holroyd and M. Coles, The neural basis of human error processing: Reinforcement learning, dopamine, and the error-related negativity, Psychological Review, vol.109, pp.679-709, 2002.

C. B. Holroyd and N. Yeung, Motivation of extended behaviors by anterior cingulate cortex, Trends in Cognitive Sciences, vol.16, pp.121-127, 2012.
DOI : 10.1016/j.tics.2011.12.008

C. B. Holroyd, S. Nieuwenhuis, N. Yeung, L. Nystrom, R. B. Mars et al., Dorsal anterior cingulate cortex shows fMRI response to internal and external error signals, Nature Neuroscience, vol.7, pp.497-498, 2004.
DOI : 10.1038/nn1238

URL : http://dare.ubvu.vu.nl/bitstream/1871/17407/2/Holroyd%20Nature%20Neuroscience%207%285%29%202004%20u.pdf

E. Hoshi, Differential Roles of Neuronal Activity in the Supplementary and Presupplementary Motor Areas: From Information Retrieval to Motor Planning and Execution, Journal of Neurophysiology, vol.92, pp.3482-3499, 2004.

A. Hyafil, C. Summerfield, and E. Koechlin, Two Mechanisms for Task Switching in the Prefrontal Cortex, Journal of Neuroscience, vol.29, pp.5135-5142, 2009.

J. M. Hyman, J. Whitman, E. Emberly, T. S. Woodward, and J. K. Seamans, Action and Outcome Activity State Patterns in the Anterior Cingulate Cortex, Cerebral Cortex, vol.23, pp.1257-1268, 2013.
DOI : 10.1093/cercor/bhs104

URL : https://academic.oup.com/cercor/article-pdf/23/6/1257/1182247/bhs104.pdf

M. Inase, H. Tokuno, A. Nambu, T. Akazawa, and M. Takada, Corticostriatal and corticosubthalamic input zones from the presupplementary motor area in the macaque monkey: comparison with the input zones from the supplementary motor area, Brain Research, vol.833, pp.191-201, 1999.

A. Inman and S. J. Shettleworth, Detecting metamemory in nonverbal subjects: A test with pigeons, Journal of Experimental Psychology: Animal Behavior Processes, vol.25, pp.389-395, 1999.
DOI : 10.1037/0097-7403.25.3.389

M. Isoda and O. Hikosaka, Switching from automatic to controlled action by monkey medial frontal cortex, Nat Neurosci, vol.10, pp.240-248, 2007.
DOI : 10.1038/nn1830

Y. Isomura and M. Takada, Neural mechanisms of versatile functions in primate anterior cingulate cortex, Rev Neurosci, vol.15, pp.279-291, 2004.
DOI : 10.1515/revneuro.2004.15.4.279

S. Ito, V. Stuphorn, J. W. Brown, and J. D. Schall, Performance monitoring by the anterior cingulate cortex during saccade countermanding, Science, vol.302, pp.120-122, 2003.
DOI : 10.1126/science.1087847

N. Jaafari, B. Aouizerate, J. Tignol, W. El-hage, I. Wassouf et al., The Relationship between Insight and Uncertainty in Obsessive-Compulsive Disorder, vol.44, pp.272-276, 2011.
DOI : 10.1159/000323607

M. Jahanshahi, I. H. Jenkins, R. G. Brown, C. D. Marsden, R. E. Passingham et al., Selfinitiated versus externally triggered movements. I. An investigation using measurement of regional cerebral blood flow with PET and movement-related potentials in normal and Parkinson's disease subjects, Brain, vol.118, pp.913-933, 1995.

J. Jastrow and C. S. Peirce, On small differences of sensation, pp.1-11, 1884.

I. H. Jenkins, M. Jahanshahi, M. Jueptner, R. E. Passingham, and D. J. Brooks, Self-initiated versus externally triggered movements. II. The effect of movement predictability on regional cerebral blood flow, Brain, vol.123, pp.1216-1228, 2000.
DOI : 10.1093/brain/123.6.1216

URL : https://academic.oup.com/brain/article-pdf/123/6/1216/17873791/1231216.pdf

R. Johnson and E. Donchin, On how P300 amplitude varies with the utility of the eliciting stimuli, Electroencephalogr Clin Neurophysiol, vol.44, pp.424-437, 1978.

M. P. Karlsson, D. Tervo, and A. Y. Karpova, Network Resets in Medial Prefrontal Cortex Mark the Onset of Behavioral Uncertainty, Science, vol.338, pp.135-139, 2012.

T. Kawai, H. Yamada, N. Sato, M. Takada, and M. Matsumoto, Roles of the Lateral Habenula and Anterior Cingulate Cortex in Negative Outcome Monitoring and Behavioral Adjustment in Nonhuman Primates, pp.1-13, 2015.

S. W. Kennerley, M. E. Walton, T. Behrens, M. J. Buckley, and M. Rushworth, Optimal decision making and the anterior cingulate cortex, Nature Neuroscience, vol.9, pp.940-947, 2006.
DOI : 10.1038/nn1724

A. Kepecs and Z. F. Mainen, A computational framework for the study of confidence in humans and animals, Philos Trans R Soc Lond B Biol Sci, vol.367, pp.1322-1337, 2012.

A. Kepecs, N. Uchida, H. A. Zariwala, and Z. F. Mainen, Neural correlates, computation and behavioural impact of decision confidence, Nature, vol.455, pp.227-231, 2008.

M. Khamassi, R. Quilodran, P. Enel, P. F. Dominey, and E. Procyk, Behavioral Regulation and the Modulation of Information Coding in the Lateral Prefrontal and Cingulate Cortex, Cereb Cortex, vol.25, pp.3197-3218, 2015.
URL : https://hal.archives-ouvertes.fr/hal-01219972

R. Kiani and M. N. Shadlen, Representation of confidence associated with a decision by neurons in the parietal cortex, Science, vol.324, pp.759-764, 2009.

R. Kiani, L. Corthell, and M. N. Shadlen, Choice Certainty Is Informed by Both Evidence and Decision Time, Neuron, vol.84, pp.1329-1342, 2014.

C. Kirschbaum and D. H. Hellhammer, Salivary cortisol in psychobiological research: an overview, Neuropsychobiology, vol.22, pp.150-169, 1989.

N. Kolling, T. Behrens, R. B. Mars, and M. Rushworth, Neural mechanisms of foraging, Science, vol.336, pp.95-98, 2012.

Y. Komura, A. Nikkuni, N. Hirashima, T. Uetake, and A. Miyamoto, Responses of pulvinar neurons reflect a subject's confidence in visual categorization, Nat Neurosci, vol.16, pp.749-755, 2013.

N. Kornell, L. K. Son, and H. S. Terrace, Transfer of metacognitive skills and hint seeking in monkeys, Psychol Sci, vol.18, pp.64-71, 2007.

H. W. Krohne, The concept of coping modes: Relating cognitive person variables to actual coping behavior, Advances in Behaviour Research and Therapy, vol.11, pp.235-248, 1989.

C. Kunimoto, J. Miller, and H. Pashler, Confidence and Accuracy of Near-Threshold Discrimination Responses, Consciousness and Cognition, vol.10, pp.294-340, 2001.

M. Kuwabara, F. A. Mansouri, M. J. Buckley, and K. Tanaka, Cognitive Control Functions of Anterior Cingulate Cortex in Macaque Monkeys Performing a Wisconsin Card Sorting Test Analog, Journal of Neuroscience, vol.34, pp.7531-7547, 2014.

D. Kvitsiani, S. Ranade, B. Hangya, H. Taniguchi, and J. Z. Huang, Distinct behavioural and network correlates of two interneuron types in prefrontal cortex, Nature, vol.498, pp.363-366, 2013.

A. Lak, G. M. Costa, E. Romberg, A. A. Koulakov, Z. F. Mainen et al., Orbitofrontal Cortex Is Required for Optimal Waiting Based on Decision Confidence, Neuron, vol.84, pp.190-201, 2014.

, Vérification-définition de vérification en français du dictionnaire Larrousse, 2016.

H. C. Lau, Attention to Intention, Science, vol.303, pp.1208-1210, 2004.

L. Pelley and M. E. , Metacognitive monkeys or associative animals? Simple reinforcement learning explains uncertainty in nonhuman animals, Journal of Experimental Psychology: Learning, Memory, and Cognition, vol.38, pp.686-708, 2012.

J. F. Leckman, D. E. Grice, J. Boardman, H. Zhang, A. Vitale et al., Symptoms of obsessive-compulsive disorder, American Journal of Psychiatry, vol.154, pp.911-917, 1997.

B. Levant, D. E. Grigoriadis, and E. B. Desouza, 3H]quinpirole binding to putative D2 and D3 dopamine receptors in rat brain and pituitary gland: a quantitative autoradiographic study, The Journal of pharmacology and experimental therapeutics, vol.264, pp.991-1001, 1993.

C. M. Leyland and W. K. Honig, Maintenance of behavior controlling the duration of discriminative stimuli, Journal of the Experimental Analysis of Behavior, vol.24, pp.207-214, 1975.

C. Lind and M. J. Boschen, Intolerance of uncertainty mediates the relationship between responsibility beliefs and compulsive checking, Journal of Anxiety Disorders, vol.23, pp.1047-1052, 2009.

M. T. Lu, J. B. Preston, and P. L. Strick, Journal of Comparative Neurology, vol.341, pp.375-392, 1994.

G. Luppino, M. Matelli, and R. Camarda, Corticocortical connections of area F3 (SMAproper) and area F6 (pre-SMA) in the macaque monkey, Journal of Comparative Neurology, 1993.

G. Luppino, M. Matelli, R. M. Camarda, V. Gallese, and G. Rizzolatti, Multiple representations of body movements in mesial area 6 and the adjacent cingulate cortex: an intracortical microstimulation study in the macaque monkey, The Journal of Comparative Neurology, vol.311, pp.463-482, 1991.

C. K. Lutz, S. Tiefenbacher, M. J. Jorgensen, J. S. Meyer, and M. A. Novak, Techniques for collecting saliva from awake, unrestrained, adult monkeys for cortisol assay, Am J Primatol, vol.52, pp.93-99, 2000.

J. M. Macpherson, C. Marangoz, T. S. Miles, and M. Wiesendanger, Microstimulation of the supplementary motor area (SMA) in the awake monkey, Exp Brain Res, vol.45, pp.410-416, 1982.

T. V. Maia, R. E. Cooney, and B. S. Peterson, The neural bases of obsessive-compulsive disorder in children and adults, Development and Psychopathology, vol.20, p.1251, 2008.

H. L. Marsh and S. E. Macdonald, Information seeking by orangutans: a generalized search strategy?, Animal Cognition, vol.15, pp.293-304, 2011.

D. Mataix-cols and S. Wooderson, Distinct Neural Correlates of Washing, Checking, and Hoarding SymptomDimensions in Obsessive-compulsive Disorder, Archives of General Psychiatry, vol.61, pp.564-576, 2004.

D. Mataix-cols, R. Mcd, and J. F. Leckman, A multidimensional model of obsessive-compulsive disorder, Am J Psychiatry, vol.162, pp.228-238, 2005.

M. Matelli, G. Luppino, and G. Rizzolatti, Architecture of superior and mesial area 6 and the adjacent cingulate cortex in the macaque monkey, The Journal of Comparative Neurology, vol.311, pp.445-462, 1991.

K. Matsumoto, W. Suzuki, and K. Tanaka, Neuronal correlates of goal-based motor selection in the prefrontal cortex, Science, vol.301, pp.229-232, 2003.

M. Matsumoto, K. Matsumoto, H. Abe, and K. Tanaka, Medial prefrontal cell activity signaling prediction errors of action values, Nature Neuroscience, vol.10, pp.647-656, 2007.

Y. Matsuzaka and J. Tanji, Changing directions of forthcoming arm movements: neuronal activity in the presupplementary and supplementary motor area of monkey cerebral cortex, Journal of Neurophysiology, vol.76, pp.2327-2342, 1996.

Y. Matsuzaka, H. Aizawa, and J. Tanji, A motor area rostral to the supplementary motor area (presupplementary motor area) in the monkey: neuronal activity during a learned motor task, J Neurophysiol, vol.68, pp.653-662, 1992.

S. Mcmahon, K. Macpherson, and W. A. Roberts, Dogs choose a human informant: Metacognition in canines, Behavioural processes, vol.85, pp.293-298, 2010.

R. J. Mcnally and P. A. Kohlbeck, Reality monitoring in obsessive-compulsive disorder, Behav Res Ther, vol.31, pp.249-253, 1993.

K. Merten and A. Nieder, Comparison of abstract decision encoding in the monkey prefrontal cortex, the presupplementary, and cingulate motor areas, Journal of Neurophysiology, vol.110, pp.19-32, 2013.

J. Metcalfe, Drawing the line on metacognition, Behav Brain Sci, vol.26, pp.350-351, 2003.

T. Michelet, B. Bioulac, D. Guehl, L. Escola, and P. Burbaud, Impact of Commitment on Performance Evaluation in the Rostral Cingulate Motor Area, Journal of Neuroscience, vol.27, pp.7482-7489, 2007.

T. Michelet, B. Bioulac, D. Guehl, M. Goillandeau, and P. Burbaud, Single medial prefrontal neurons cope with error, Plos ONE, vol.4, p.6240, 2009.

T. Michelet, B. Bioulac, N. Langbour, M. Goillandeau, D. Guehl et al., Electrophysiological Correlates of a Versatile Executive Control System in the Monkey Anterior Cingulate Cortex, Cereb Cortex, vol.26, pp.1684-1697, 2016.
URL : https://hal.archives-ouvertes.fr/hal-01155375

P. G. Middlebrooks and M. A. Sommer, Metacognition in monkeys during an oculomotor task, J Exp Psychol Learn Mem Cogn, vol.37, pp.325-337, 2011.

W. H. Miltner, C. H. Braun, and M. G. Coles, Event-related brain potentials following incorrect feedback in a time-estimation task: evidence for a "generic" neural system for error detection, J Cogn Neurosci, vol.9, pp.788-798, 1997.

A. R. Mitz and S. P. Wise, The somatotopic organization of the supplementary motor area: intracortical microstimulation mapping, J Neurosci, vol.7, pp.1010-1021, 1987.

K. F. Muakkassa and P. L. Strick, Frontal lobe inputs to primate motor cortex: evidence for four somatotopically organized 'premotor' areas, Brain Research, pp.176-182, 1979.

P. Nachev, C. Kennard, and M. Husain, Functional role of the supplementary and presupplementary motor areas, Nature Reviews Neuroscience, vol.9, pp.856-869, 2008.

P. Nachev, G. Rees, A. Parton, C. Kennard, and M. Husain, Human Medial Frontal Cortex. Current Biology, vol.15, pp.122-128, 2005.

P. Nachev, H. Wydell, K. O. Neill, M. Husain, and C. Kennard, The role of the presupplementary motor area in the control of action, Human Brain Mapping Journal, vol.36, pp.155-163, 2007.

K. Nakamura, K. Sakai, and O. Hikosaka, Neuronal activity in medial frontal cortex during learning of sequential procedures, Journal of Neurophysiology, vol.80, pp.2671-2687, 1998.

N. S. Narayanan, J. F. Cavanagh, M. J. Frank, and M. Laubach, Common medial frontal mechanisms of adaptive control in humans and rodents, Nature Publishing Group, vol.16, pp.1888-1895, 2013.

M. Nedeljkovic and M. Kyrios, Confidence in memory and other cognitive processes in obsessive-compulsive disorder, Behaviour Research and Therapy, vol.45, pp.2899-2914, 2007.

G. Nemeth, K. Hegedus, and L. Molnar, Akinetic mutism associated with bicingular lesions: clinicopathological and functional anatomical correlates, Eur Arch Psychiatry Neurol Sci, vol.237, pp.218-222, 1988.

S. Nieuwenhuis, N. Yeung, W. Van-den-wildenberg, and K. R. Ridderinkhof, Electrophysiological correlates of anterior cingulate function in a go/no-go task: effects of response conflict and trial type frequency, Cogn Affect Behav Neurosci, vol.3, pp.17-26, 2003.

H. Niki and M. Watanabe, Prefrontal and cingulate unit activity during timing behavior in the monkey, Brain Research, vol.171, pp.213-224, 1979.

C. R. Oehrn, S. Hanslmayr, J. Fell, L. Deuker, N. A. Kremers et al., Neural Communication Patterns Underlying Conflict Detection, Resolution, and Adaptation, Journal of Neuroscience, vol.34, pp.10438-10452, 2014.

J. V. Pardo, P. J. Pardo, K. W. Janer, and M. E. Raichle, The anterior cingulate cortex mediates processing selection in the Stroop attentional conflict paradigm, Proc Natl Acad Sci U S A, vol.87, pp.256-259, 1990.

J. Parvizi, V. Rangarajan, W. R. Shirer, N. Desai, and M. D. Greicius, The Will to Persevere Induced by Electrical Stimulation of the Human Cingulate Gyrus, Neuron, vol.80, pp.1359-1367, 2013.

A. Paukner, J. R. Anderson, and K. Fujita, Redundant food searches by capuchin monkeys (Cebus apella): a failure of metacognition?, Animal Cognition, vol.9, pp.110-117, 2005.

T. Paus, Primate anterior cingulate cortex: where motor control, drive and cognition interface, Nat Rev Neurosci, vol.2, pp.417-424, 2001.

B. L. Pearson, P. G. Judge, and D. M. Reeder, Effectiveness of saliva collection and enzymeimmunoassay for the quantification of cortisol in socially housed baboons, Am J Primatol, vol.70, pp.1145-1151, 2008.

D. Perani, C. Colombo, S. Bressi, A. Bonfanti, F. Grassi et al., 18F]FDG PET study in obsessive-compulsive disorder. A clinical/metabolic correlation study after treatment, The British Journal of Psychiatry, vol.166, pp.244-250, 1995.

C. J. Perry and A. B. Barron, Honey bees selectively avoid difficult choices, Proceedings of the National Academy of Sciences, vol.110, pp.19155-19159, 2013.

N. Persaud, P. Mcleod, and A. Cowey, Post-decision wagering objectively measures awareness, Nat Neurosci, vol.10, pp.257-261, 2007.

D. K. Peterson and G. F. Pitz, Confidence, uncertainty, and the use of information, Journal of Experimental Psychology: Learning, Memory, and Cognition, vol.14, pp.85-92, 1988.

N. Picard and P. L. Strick, Motor areas of the medial wall: a review of their location and functional activation, Cerebral cortex, vol.6, pp.342-353, 1991.

N. Picard and P. L. Strick, Imaging the premotor areas, Curr Opin Neurobiol, vol.11, pp.663-672, 2001.

R. K. Pitman, A cybernetic model of obsessive-compulsive psychopathology, Comprehensive Psychiatry, vol.28, pp.334-343, 1987.

R. K. Pitman, Animal models of compulsive behavior, Biological Psychiatry, vol.26, pp.189-198, 1989.

A. Pouget, J. Drugowitsch, and A. Kepecs, Confidence and certainty: distinct probabilistic quantities for different goals, Nature Neuroscience, vol.19, pp.366-374, 2016.

E. Procyk, Y. L. Tanaka, and J. P. Joseph, Anterior cingulate activity during routine and nonroutine sequential behaviors in macaques, Nat Neurosci, vol.3, pp.502-508, 2000.
URL : https://hal.archives-ouvertes.fr/inserm-00132133

E. Procyk, C. R. Wilson, F. M. Stoll, M. C. Faraut, M. Petrides et al., Midcingulate Motor Map and Feedback Detection: Converging Data from Humans and Monkeys, Cereb Cortex, vol.26, pp.467-476, 2016.

R. Quilodran, M. Rothé, and E. Procyk, Behavioral Shifts and Action Valuation in the Anterior Cingulate Cortex, Neuron, vol.57, pp.314-325, 2008.
URL : https://hal.archives-ouvertes.fr/inserm-00906686

S. Rachman, A cognitive theory of compulsive checking, Behav Res Ther, vol.40, pp.625-639, 2002.

A. S. Radomsky, S. Rachman, and D. Hammond, Memory bias, confidence and responsibility in compulsive checking, Behaviour Research and Therapy, vol.39, pp.813-822, 2001.

A. S. Radomsky, P. T. Gilchrist, and D. Dussault, Repeated checking really does cause memory distrust, Behaviour Research and Therapy, vol.44, pp.305-316, 2006.

M. E. Raichle, J. A. Fiez, T. O. Videen, A. M. Macleod, J. V. Pardo et al., Practice-related changes in human brain functional anatomy during nonmotor learning, Cerebral cortex, vol.4, pp.8-26, 1991.

N. Ramnani and R. C. Miall, Instructed delay activity in the human prefrontal cortex is modulated by monetary reward expectation, Cereb Cortex, vol.13, pp.318-327, 2003.

R. Li-cs, Imaging Response Inhibition in a Stop-Signal Task: Neural Correlates Independent of Signal Monitoring and Post-Response Processing, Journal of Neuroscience, vol.26, pp.186-192, 2006.

G. F. Reed, Obsessional personality disorder and remembering, The British Journal of Psychiatry, vol.130, pp.177-183, 1977.

K. R. Ridderinkhof, M. Ullsperger, E. A. Crone, and S. Nieuwenhuis, The role of the medial frontal cortex in cognitive control, Science, vol.306, pp.443-447, 2004.

W. A. Roberts, M. C. Feeney, N. Mcmillan, K. Macpherson, E. Musolino et al., Do Pigeons (Columba livia) Study for a Test, Journal of Experimental Psychology: Animal Behavior Processes, vol.35, pp.129-142, 2009.
DOI : 10.1037/a0013722

P. E. Roland, B. Larsen, N. A. Lassen, and E. Skinhoj, Supplementary motor area and other cortical areas in organization of voluntary movements in man, J Neurophysiol, vol.43, pp.118-136, 1980.

R. Romo and W. Schultz, Neuronal activity preceding self-initiated or externally timed arm movements in area 6 of monkey cortex, Exp Brain Res, vol.67, pp.656-662, 1987.

G. Roper, S. Rachman, and R. Hodgson, An experiment on obsessional checking, Behav Res Ther, vol.11, pp.271-277, 1973.
DOI : 10.1016/0005-7967(73)90003-x

J. Y. Rotge, N. Langbour, B. Dilharreguy, M. Bordessoulles, D. Guehl et al., Contextual and behavioral influences on uncertainty in obsessive-compulsive disorder, vol.62, pp.1-10, 2015.
DOI : 10.1016/j.cortex.2012.12.010

URL : https://hal.archives-ouvertes.fr/hal-01155377

J. Y. Rotge, B. Aouizerate, V. Amestoy, V. Lambrecq, N. Langbour et al., The associative and limbic thalamus in the pathophysiology of obsessive-compulsive disorder: an experimental study in the monkey, Transl Psychiatry, vol.2, p.161, 2012.
URL : https://hal.archives-ouvertes.fr/hal-01155556

J. Y. Rotgé, A. H. Clair, N. Jaafari, E. G. Hantouche, A. Pelissolo et al., A challenging task for assessment of checking behaviors in obsessive-compulsive disorder, Acta Psychiatr Scand, vol.117, pp.465-473, 2008.

A. M. Ruscio, D. J. Stein, W. T. Chiu, and R. C. Kessler, The epidemiology of obsessivecompulsive disorder in the National Comorbidity Survey Replication, Mol Psychiatry, vol.15, pp.53-63, 2008.

M. F. Rushworth, K. A. Hadland, T. Paus, and P. K. Sipila, Role of the human medial frontal cortex in task switching: a combined fMRI and TMS study, J Neurophysiol, vol.87, pp.2577-2592, 2002.

M. F. Rushworth, M. E. Walton, S. W. Kennerley, and D. M. Bannerman, Action sets and decisions in the medial frontal cortex, Trends Cogn Sci, vol.8, pp.410-417, 2004.

M. Rushworth, K. A. Hadland, D. Gaffan, and R. E. Passingham, The effect of cingulate cortex lesions on task switching and working memory, Journal of cognitive neuroscience, vol.15, pp.338-353, 2003.

M. Rushworth, M. P. Noonan, E. D. Boorman, M. E. Walton, and T. E. Behrens, Frontal Cortex and Reward-Guided Learning and Decision-Making, Neuron, vol.70, pp.1054-1069, 2011.
DOI : 10.1016/j.neuron.2011.05.014

URL : https://doi.org/10.1016/j.neuron.2011.05.014

P. Salgado-pineda, C. Junqué, P. Vendrell, I. Baeza, N. Bargalló et al., Decreased cerebral activation during CPT performance, NeuroImage, vol.21, pp.840-847, 2004.
DOI : 10.1016/j.neuroimage.2003.10.027

J. H. Sandell, C. G. Gross, and M. H. Bornstein, Color categories in macaques, J Comp Physiol Psychol, vol.93, pp.626-635, 1979.
DOI : 10.1037/h0077594

J. I. Sanders, B. Hangya, and A. Kepecs, Signatures of a Statistical Computation in the Human Sense of Confidence, Neuron, vol.90, pp.499-506, 2016.

K. W. Scangos, R. Aronberg, and V. Stuphorn, Performance monitoring by presupplementary and supplementary motor area during an arm movement countermanding task, Journal of Neurophysiology, vol.109, pp.1928-1939, 2013.
DOI : 10.1152/jn.00688.2012

URL : https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3628008/pdf

J. D. Schall, V. Stuphorn, and J. W. Brown, Monitoring and control of action by the frontal lobes, Neuron, vol.36, pp.309-322, 2002.

J. M. Schwartz, Neuroanatomical aspects of cognitive-behavioural therapy response in obsessive-compulsive disorder. An evolving perspective on brain and behaviour. The British journal of psychiatry Supplement, pp.38-44, 1997.

J. K. Seamans, S. B. Floresco, and A. G. Phillips, Functional differences between the prelimbic and anterior cingulate regions of the rat prefrontal cortex, Behavioral Neuroscience, vol.109, pp.1063-1073, 1995.

J. Sergent, E. Zuck, S. Terriah, and B. Macdonald, Distributed neural network underlying musical sight-reading and keyboard performance, Science, vol.257, pp.106-109, 1992.
DOI : 10.1126/science.1621084

A. Shenhav, M. M. Botvinick, and J. D. Cohen, The expected value of control: an integrative theory of anterior cingulate cortex function, Neuron, vol.79, pp.217-240, 2013.

A. Shenhav, J. D. Cohen, and M. M. Botvinick, Dorsal anterior cingulate cortex and the value of control, Nature Publishing Group, vol.19, pp.1286-1291, 2016.

A. Shenhav, M. A. Straccia, J. D. Cohen, and M. M. Botvinick, Anterior cingulate engagement in a foraging context reflects choice difficulty, not foraging value, Nature Publishing Group, vol.17, pp.1249-1254, 2014.

K. J. Sher, R. O. Frost, and R. Otto, Cognitive deficits in compulsive checkers: an exploratory study, Behav Res Ther, vol.21, pp.357-363, 1983.

S. A. Sheth, M. K. Mian, S. R. Patel, W. F. Asaad, Z. M. Williams et al., Human dorsal anterior cingulate cortex neurons mediate ongoing behavioural adaptation, Nature, vol.488, pp.218-221, 2012.

M. Shidara and B. J. Richmond, Anterior cingulate: single neuronal signals related to degree of reward expectancy, Science, vol.296, pp.1709-1711, 2002.

W. E. Shields, J. D. Smith, and D. A. Washburn, Uncertain responses by humans and rhesus monkeys (Macaca mulatta) in a psychophysical same-different task, J Exp Psychol Gen, vol.126, pp.147-164, 1997.

W. E. Shields, J. D. Smith, K. Guttmannova, and D. A. Washburn, Confidence judgments by humans and rhesus monkeys, J Gen Psychol, vol.132, pp.165-186, 2005.

K. Shima and J. Tanji, Role for cingulate motor area cells in voluntary movement selection based on reward, Science, vol.282, pp.1335-1338, 1998.

K. Shima and J. Tanji, Both supplementary and presupplementary motor areas are crucial for the temporal organization of multiple movements, Journal of Neurophysiology, vol.80, pp.3247-3260, 1998.

K. Shima, H. Mushiake, N. Saito, and J. Tanji, Role for cells in the presupplementary motor area in updating motor plans, Proceedings of the National Academy of Sciences of the United States of America, vol.93, pp.8694-8698, 1996.

K. Shima, K. Aya, H. Mushiake, M. Inase, H. Aizawa et al., Two movement-related foci in the primate cingulate cortex observed in signal-triggered and self-paced forelimb movements, J Neurophysiol, vol.65, pp.188-202, 1991.

J. D. Smith, J. J. Couchman, and M. J. Beran, Animal metacognition: A tale of two comparative psychologies, Journal of Comparative Psychology, vol.128, pp.115-131, 2014.

J. D. Smith, J. Schull, J. Strote, K. Mcgee, R. Egnor et al., The uncertain response in the bottlenosed dolphin (Tursiops truncatus), J Exp Psychol Gen, vol.124, pp.391-408, 1995.

L. M. Sole, S. J. Shettleworth, and P. J. Bennett, Uncertainty in pigeons, Psychon Bull Rev, vol.10, pp.738-745, 2003.

L. K. Son and N. Kornell, Metaconfidence Judgments in Rhesus Macaques: Explicit Versus Implicit Mechanisms, pp.296-320, 2008.

S. M. Stasik, K. Naragon-gainey, M. Chmielewski, and D. Watson, Core OCD symptoms: exploration of specificity and relations with psychopathology, J Anxiety Disord, vol.26, pp.859-870, 2012.

M. B. Stein, D. R. Forde, G. Anderson, and J. R. Walker, Obsessive-compulsive disorder in the community: an epidemiologic survey with clinical reappraisal, Am J Psychiatry, vol.154, pp.1120-1126, 1997.

F. M. Stoll, V. Fontanier, and E. Procyk, Specific frontal neural dynamics contribute to decisions to check, Nature communications, vol.7, pp.1-14, 2016.

C. Suda-king, Do orangutans (Pongo pygmaeus) know when they do not remember?, Anim Cogn, vol.11, pp.21-42, 2007.

D. Swick and A. U. Turken, Dissociation between conflict detection and error monitoring in the human anterior cingulate cortex, Proceedings of the National Academy of Sciences of the United States of America, vol.99, pp.16354-16359, 2002.

H. Szechtman, W. Sulis, and D. Eilam, Quinpirole induces compulsive checking behavior in rats: a potential animal model of obsessive-compulsive disorder (OCD), Behavioral Neuroscience, vol.112, pp.1475-1485, 1998.

A. Szmalec, F. Verbruggen, A. Vandierendonck, W. De-baene, T. Verguts et al., Stimulus ambiguity elicits response conflict, Neurosci Lett, vol.435, pp.158-162, 2008.
DOI : 10.1016/j.neulet.2008.02.023

H. Tabu, T. Mima, T. Aso, R. Takahashi, and H. Fukuyama, Functional relevance of presupplementary motor areas for the choice to stop during Stop signal task, Neuroscience research, vol.70, pp.277-284, 2011.

J. Talairach, J. Bancaud, S. Geier, M. Bordas-ferrer, A. Bonis et al., The cingulate gyrus and human behaviour, Electroencephalography and clinical neurophysiology, vol.34, pp.45-52, 1973.
DOI : 10.1016/0013-4694(73)90149-1

J. Tanji, Sequential organization of multiple movements: involvement of cortical motor areas, Annual Review of Neuroscience, vol.24, pp.631-651, 2001.

J. Tanji and K. Shima, Role for supplementary motor area cells in planning several movements ahead, Nature, vol.371, pp.413-416, 1994.

R. Thompson, Underwater frequency discrimination in the bottlenosed dolphin (1140 kHz) and the human (1-8 kHz), J Acoust Soc Am, vol.57, p.943, 1975.

E. L. Thorndike, Animal Intelligence. Experimental Studies. The Journal of Nervous and Mental Disease, vol.39, p.357, 1912.

D. F. Tolin, J. S. Abramowitz, B. D. Brigidi, and E. B. Foa, Intolerance of uncertainty in obsessive-compulsive disorder, Journal of Anxiety Disorders, vol.17, pp.233-242, 2003.

D. F. Tolin, J. S. Abramowitz, B. D. Brigidi, N. Amir, G. P. Street et al., Memory and memory confidence in obsessive-compulsive disorder, Behaviour Research and Therapy, vol.39, pp.913-927, 2001.

J. M. Tyszka, S. T. Grafton, W. Chew, R. P. Woods, and P. M. Colletti, Parceling of mesial frontal motor areas during ideation and movement using functional magnetic resonance imaging at 1.5 tesla, Annals of neurology, vol.35, pp.746-749, 1994.

M. Ullsperger and D. Y. Von-cramon, Subprocesses of Performance Monitoring: A Dissociation of Error Processing and Response Competition Revealed by EventRelated fMRI and ERPs, NeuroImage, vol.14, pp.1387-1401, 2001.

S. Ursu, V. A. Stenger, M. K. Shear, M. R. Jones, C. Carter et al., Overactive action monitoring in obsessive-compulsive disorder: evidence from functional magnetic resonance imaging, van den Hout M, Kindt M, vol.14, pp.301-316, 2003.
DOI : 10.1111/1467-9280.24411

V. Van-veen and C. S. Carter, The anterior cingulate as a conflict monitor: fMRI and ERP studies, Physiol Behav, vol.77, pp.477-482, 2002.

D. Vickers, J. Burt, P. Smith, and M. Brown, Experimental paradigms emphasising state or process limitations: I Effects on speed-accuracy tradeoffs, Acta Psychologica, vol.59, pp.129-161, 1985.
DOI : 10.1016/0001-6918(85)90017-4

D. Vickers, P. Smith, J. Burt, and M. Brown, Experimental paradigms emphasising state or process limitations: II effects on confidence, Acta Psychologica, vol.59, pp.163-193, 1985.
DOI : 10.1016/0001-6918(85)90018-6

A. Q. Vining and H. L. Marsh, Information seeking in capuchins (Cebus apella): A rudimentary form of metacognition?, Animal Cognition, vol.18, pp.667-681, 2015.

M. E. Walton, D. M. Bannerman, and M. Rushworth, The role of rat medial frontal cortex in effort-based decision making, Journal of Neuroscience, vol.22, pp.10996-11003, 2002.

M. E. Walton, S. W. Kennerley, D. M. Bannerman, P. Phillips, and M. Rushworth, Weighing up the benefits of work: Behavioral and neural analyses of effort-related decision making, Neural Networks, vol.19, pp.1302-1314, 2006.

D. A. Washburn, J. D. Smith, and W. E. Shields, Rhesus monkeys (macaca mulatta) immediately generalize the uncertain response, Journal of Experimental Psychology: Animal Behavior Processes, vol.32, pp.185-189, 2006.
DOI : 10.1037/0097-7403.32.2.185

D. A. Washburn, J. P. Gulledge, M. J. Beran, and J. D. Smith, With his memory magnetically erased, a monkey knows he is uncertain, Biology Letters, vol.6, pp.160-162, 2010.

M. Watanabe, H. C. Cromwell, L. Tremblay, J. R. Hollerman, K. Hikosaka et al., Behavioral reactions reflecting differential reward expectations in monkeys, Experimental Brain Research, vol.140, pp.511-518, 2001.

S. Yazawa, A. Ikeda, T. Kunieda, S. Ohara, T. Mima et al., Human presupplementary motor area is active before voluntary movement: subdural recording of Bereitschaftspotential from medial frontal cortex, Experimental Brain Research, vol.131, pp.165-177, 2000.

A. Yildiz, O. T. Wolf, and C. Beste, Stress intensifies demands on response selection during action cascading processes, Psychoneuroendocrinology, vol.42, pp.178-187, 2014.

P. Zadicario, S. Ronen, and D. Eilam, Modulation of quinpirole-induced compulsive-like behavior in rats by environmental changes: Implications for OCD rituals and for exploration and navigation, BMC Neuroscience, vol.8, pp.1-18, 2007.

N. Zarr and J. W. Brown, Hierarchical error representation in medial prefrontal cortex, NeuroImage, vol.124, pp.238-247, 2016.