. Al-safadi, area and southern Bahr el Jebel. It was first found in the southern region of Sudan (Southern Sudan country) by Longstaff (1914) who obtained no more than 5 specimens; the sites now reported show L, Outside the mainland of Africa, this species is possibly present in Yemen, 1977.

. Brown, Habitat: It is found in permanent standing and running waters of different sizes (IUCN, 2017) including lakes, rivers, khors, flooded areas and irrigation canals; a wide variety of macrophytes grew at these sites, 1984.

B. Semi, aquatic snails Pila wernei, p.1851

, It is also known under the name of Ampullaria wernei, P. wernei belongs to the wide family of Ampullariidae, 1902.

. Longstaff-;-in-brown, Philippi, 1851 in Köhler and Glaubrecht, 2006) Geographic range: Pila wernei is generally known from scattered localities over a wide area in Africa, from Mali to Somalia, but it represents a native species in Cameroon, 1914.

. Brown, Darfur and Bahr el Jebel, 1984.

, 1213 palynomorphs out of the total counted specimens have been classified into four main groups in order to help in the paleoclimate interpretation. The identified groups are : aquatic, tropical, savanna, and arid indicators

. Latrubesse, This group includes Chomotriletes, 2009.

G. Worobiec and . Scafati, Plates 4.2 and 4.3 show some representative pollens of this group. The tropical climate indicators group includes Proxapertites, Lecaniella forma (Worobiec, 2014), Spirogyra (zygospore, 1993.

, Cymbopogon schoenanthus (El Ghazali, 1993) and Monolites alveolatus, Silene burchellii, 1993.

. Volkheimer, Classopollis spp, Leiotriletes adriennis (Uzodimma, 2013), and Erica arbora, 1993.

A. Galil and M. Y. , Geology and Mineralization related to anorogenic igneous complex of Northern Nuba Mountains and Northern Kordofan, 2008.

A. Abdu, The Crescentic dunes of northern Sudan, East African Geographical Review, vol.14, pp.61-71, 1976.

P. Abell and M. Williams, Oxygen and carbon isotope ratios in gastropod shells as indicators of paleoenvironments in Afar region of Ethiopia, Palaeogeography, Palaeoclimatology, Palaeoecology, vol.74, pp.265-278, 1989.

P. I. Abell, Oxygen isotope ratios in modern African gastropod shells: A data base for paleoclimatology, Isotope Geoscience Section), vol.58, pp.183-193, 1985.

P. I. Abell and P. Hoelzmann, Holocene palaeoclimates in northwestern Sudan: stable isotope studies on molluscs, Global and Planetary Change, vol.26, pp.1-12, 2000.

J. S. Adams, M. J. Kraus, and S. L. Wing, Evaluating the use of weathering indices for determining mean annual precipitation in the ancient stratigraphic record. Palaeogeography, Palaeoclimatology, Palaeoecology, vol.309, pp.358-366, 2011.

A. J. Alonso-zarza and B. J. Jones, Root calcrete formation on Quaternary karstic surfaces of Grand Cayman, Geologica Acta, vol.5, pp.77-88, 2007.

A. M. Alonso-zarza, Palaeoenvironmental significance of palustrine carbonates and calcretes in the geological record, Earth Science Reviews, vol.60, pp.261-298, 2003.

A. M. Alonso-zarza, J. F. Genise, and M. Verde, Sedimentology, diagenesis and ichnology of Cretaceous and Palaeogene calcretes and palustrine carbonates from Uruguay, Sedimentary Geology, vol.236, pp.45-61, 2011.

A. M. Alonso-zarza, V. P. Wright, A. M. Alonso-zarza, J. F. Genise, and M. Verde, Sedimentology, diagenesis and ichnology of Cretaceous and Palaeogene calcretes and palustrine carbonates from Uruguay, Sedimentary Geology, vol.236, pp.45-61, 2010.

M. M. Al-safadi, Freshwater molluscs of Yemen Arab Republic, IUCN, 2017. The IUCN Red List of Threatened Species: Radix natalensis, 1990.

A. V. Arakel, Quaternary vadose Calcretes revisited, Journal of Australian Geology & Geophysics, vol.16, issue.3, pp.223-229, 1996.

H. W. Arz, F. Lamy, F. Pätzold, J. Müller, P. J. Prins et al., Mediterranean Moisture Source for an Early-Holocene Humid Period in the Northern Red Sea, Science, vol.300, pp.118-121, 2003.

D. Ayliffe, M. Williams, and F. Sheldon, Stable carbon and oxygen isotopic composition of earlyHolocene gastropods from Wadi Mansurab, north-central Sudan, The Holocene, vol.6, pp.157-169, 1996.

R. A. Bagnold, Grain size distributions and depositional processes, Visher, vol.39, pp.1074-1106, 1956.

E. M. Bakker and J. Maley, Late Quaternary Palaeoenvironments of the Sahara Region. A.A. Balkema-Rotterdam, Orstom Fond Documentaire, Palaeoclimatology of Africa and the surrounding Islands, vol.11, 1977.

M. A. Berke, T. C. Johnson, J. P. Werne, S. Schouten, and J. S. Sinninghe-damsté, A mid-Holocene thermal maximum at the end of the African Humid Period, Earth and Planetary Science Letters, vol.351, pp.95-104, 2012.

M. J. Bishop, A new species of Caracolus (Pulmonata: Camaenidae) from the Oligocene of Nebraska and the biotic history of the American camaenid land snails, Zoological Journal of the Linnean Society, vol.67, pp.269-284, 1979.

S. Boggs, Principle of Sedimentology and Stratigraphy, vol.662, 2006.

D. A. Bolaji, C. A. Edokpayi, O. B. Samuel, R. O. Akinnigbagbe, and A. A. Ajulo, Morphological characteristics and Salinity tolerance of Melanoides tuberculatus (Muller, 1774), World Journal of Biological Research, vol.4, issue.2, pp.1-11, 2011.

F. Bouchette, M. Schuster, J. Ghienne, C. Denamiel, C. Roquin et al., Hydrodynamics in Holocene Lake Mega-Chad, Quaternary Research, vol.73, pp.226-236, 2010.
URL : https://hal.archives-ouvertes.fr/hal-00475633

R. S. Bradley, Recent Developments in Quaternary Paleoclimatology, Climate and Geoscience, pp.165-171, 1989.

D. S. Brown, T. Fison, V. R. Southgate, and C. A. Wright, Aquatic snails of the Jonglei region, southern Sudan, and transmission of trematode parasites, Hydrobiologia, vol.110, pp.247-271, 1984.

D. S. Brown, D. Rollinson, D. S. Brown, T. Fison, V. R. Southgate et al., Aquatic snails of the Jonglei region, southern Sudan, and transmission of trematode parasites, Hydrobiologia, vol.110, pp.247-271, 1982.

D. S. Brown and M. D. Gallagher, Freshwater snails of Oman, South Eastern Arabia, IUCN, 2017. The IUCN Red List of Threatened Species: Radix natalensis, 1985.

D. S. Brown and C. A. Wright, The IUCN Red List of Threatened Species: Radix natalensis, 1980.

F. M. Chambers, B. Van-geel, and M. Van-der-linden, Considerations for the preparation of peat samples for palynology, and for the counting of pollen and non-pollen palynomorphs, Mires and Peat, vol.7, pp.1-14, 2014.

X. Y. Chen, M. J. Lintern, and I. C. Roach, Calcrete characteristics, distribution and use in mineral exploration. Cooperative Research Centre for Landscape Environments and Mineral Exploration, vol.160, 2002.

O. I. Chiaghanam, K. C. Chiadikobi, O. N. Ikegwuonu, and A. O. Omoboriowo, Palynology, Source Rock Potential and Thermal Maturity of Eocene Nanka Formation (Ameki Group, Anambra Basin: An Investigation of Agulu Lake, vol.2, pp.87-97, 2014.

E. J. Ciolkosz and R. R. Dobos, Color and mottling in Pennsylvania soil. Agronomy series number 108. The Pennsylvania state university, 1990.

J. M. Cole, S. L. Goldstein, P. B. Hemming, S. R. Grousset, and F. E. , Contrasting compositions of Saharan dust in the eastern Atlantic Ocean during the last deglaciation and African Humid Period, Earth and Planetary Science Letters, vol.278, pp.257-266, 2009.

P. Collier, G. Conway, and T. Venables, Climate change and Africa, Oxford Review of Economic Policy, vol.24, issue.2, pp.337-353, 2008.

A. Creese and W. Pokam, Report of a project funded by the UK Department for International Development (DFID) and the Natural Environment Research Council (NERC) for the developing countries and the advance of scientific research, 2016.

O. G. Dávalos-Álvarez, Á. F. Nieto-samaniego, S. A. Alaniz-Álvarez, E. Martínez-hernández, and Y. E. Ramírez-arriaga, Estratigrafía cenozoica de la región de Tehuacán y su relación con el sector norte de la falla de Oaxaca, Revista Mexicana de Ciencias Geológicas, vol.24, issue.2, pp.197-215, 2007.

A. Dawelbeit, Hydrogeological Evaluation of Sug'a El Jamal Basin, 2010.

K. N. Kock and C. T. Wolmarans, Distribution and habitats of Melanoides tuberculata (Müller, 1774) and M. victoriae (Dohrn, 1865) (Mollusca: Prosobranchia: Thiaridae) in South Africa, Water SA, vol.35, issue.5, pp.713-720, 2009.

K. N. De-kock, C. T. Wolmarans, and M. Bornman, Distribution and habitats of Biomphalaria pfeifferi, snail intermediate host of Schistosoma mansoni, South Africa. Water SA, vol.30, pp.29-36, 2004.

M. Dehghani and H. Akhani, Pollen morphological studies in subfamily Suaedoideae (Chenopodiaceae), Grana, vol.48, pp.79-101, 2009.

P. B. Demenocal and J. E. Tierney, Green Sahara: African Humid Periods paced by Earth's orbital changes, Nature Education, 2012.

P. Demenocal, J. Ortiz, T. Guilderson, J. Adkins, M. Sarnthein et al., Abrupt onset and termination of the African Humid Period: rapid climate responses to gradual insolation forcing, Quaternary Science Reviews, vol.19, pp.347-361, 2000.

D. Demske, P. E. Tarasov, and T. Nakagawa, Atlas of pollen, spores and further non-pollen palynomorphs recorded in the glacial-interg laciallate Quaternary sediments of Lake Suigetsu, central Japan, Quaternary International, vol.290, issue.291, pp.164-238, 2006.

R. R. Dobos, E. J. Ciolkosz, and W. J. Waltman, Color and mottling in Pennsylvania soil. Agronomy series number 108. The Pennsylvania state university, 1990.

J. Dupouy, D. Rousseau, G. Dussart, M. V. Liaud, and H. Nassi, Correspondence analysis of shell morphology in the African freshwater snail Bkomphalaria pfeiierk (Kraus 1848) (Pulmonata: Gastropoda, Biological Journal of the Linnean Society, vol.50, pp.329-338, 1993.

J. M. Edmond, The Distribution of the Kordofan Sands, Anglo-Egyptian Sudan". Geol. Magazine, vol.79, issue.1, pp.18-30, 1942.

D. N. Edwards, The archeology of Sudan and Nubia, Annual Review of Anthropology, vol.36, pp.211-239, 2007.

A. Eisawi and E. Schrank, Upper Cretaceous to Neogene palynology of the Melut basin, southeast Sudan. Palunology, vol.32, pp.101-129, 2008.

G. E. El-ghazali, A study of pollen flora of Sudan, Review of Palaeobotany and Palynology, vol.76, pp.99-345, 1993.

A. A. El-shafie, K. A. Zeinelabdein, and A. A. Eisawi, Paleoclimatic evolution of central Sudan during the Late Miocene to Pleistocene. Saudi Society for Geosciences, Arab J Geosci, 2011.

Y. A. El-amier, Morphological studies of the pollen grains for some hydrophytes in coastal Mediterranean lakes, Egypt. Egyptian Journal of basic and applied Sciences, vol.2, pp.132-138, 2015.

M. Elhadi, Assessment and evaluation of the Water-bearing formations in central-north Kordofan and West White Nile States, 2012.

A. B. Elhag and S. M. Elzien, Structures controls on groundwater occurrence and flow in crystalline bedrocks: a case study of the El Obeid area, Western Sudan. Global Advanced Research Journal of Environmental Science and Toxicology, vol.2, pp.2315-5140, 2013.

A. A. Elmansour, Assessment of the groundwater of En Nuhud basin, 2017.

A. A. Elmansour and O. A. Omer, Crustal architecture, geodynamics and evolution of En Nuhud basin area, Kordofan region, Sudan. International Jurnal of advanced Geoscience, vol.5, issue.2, pp.121-132, 2017.
DOI : 10.14419/ijag.v5i2.8468

URL : https://2fwww.sciencepubco.com/index.php/IJAG/article/download/8468/2928

A. Elsheikh, M. G. Abdelsalam, and K. Mickus, Geology and geophysics of the West Nubian Paleolake and the Northern Darfur Megalake (WNPL-NDML): Implication for groundwater resources in Darfur, northwestern Sudan, Journal of African Earth Sciences, vol.61, pp.82-93, 2011.

, Organic matter decline. Sustainable agriculture and soil conservation (SoCo), 2009.

O. A. Fadipe, P. F. Carey, and S. A. Adekola, Provenance, diagenesis and reservoir quality of the lower Cretaceous sandstone of the Orange basin, South Africa. South African Journal of Geology, vol.114, issue.3-4, pp.433-448, 2011.

K. Faegri and J. Iversen, Late Quaternary palaeoenvironments of the southern Cape, South Africa: palynological evidence from three coastal wetlands, 1989.

R. G. Fairbanks, R. A. Mortlock, T. Chiu, L. Cao, A. Kaplan et al., Radiocarbon calibration curve spanning 0 to 50,000 years BP based on paired 230 Th/ 234 U/ 238 U and 14 C dates on pristine corals, Quaternary Science Reviews, vol.24, pp.1781-1796, 2005.

G. Faure and T. M. Mensing, Isotopes: principles and applications, vol.897, 2005.

B. Ferry, T. Eglin, A. Bispo, É. Dambrine, and C. Chenu, Soil organic matter of forests and climate and atmosphere changes (English translation of, Matières organiques des sols forestiers et changements climatique et atmosphérique »), 2014.

J. Filatoff, Jurassic palynology of the Perth Basin, Western Australia. Palaeontographica Bd. 154 Abt. B, 1975.

V. Foerster, A. Junginger, O. Langkamp, T. Gebru, A. Asrat et al., Climatic change recorded in the sediments of the Chew Bahir basin, southern Ethiopia, during the last 45,000 years, Quaternary International, vol.274, pp.25-37, 2012.

R. L. Folk and W. C. Ward, Brazos River Bar: a study in the significance of grain size parameters, J. Sediment. Petrol, vol.27, pp.3-26, 1957.

R. Folk, Longitudinal dunes of the northwestern edge of the Simpson Desert, northern Territory, Australia, 1. Geomorphology and grain size relationships, Sedimentology, vol.16, pp.5-54, 1974.

P. Freytet, Petrography and paleo-environment of continental carbonate deposits with particular reference to the Upper Cretaceous and Lower Eocene of Languedoc (Southern France), Sedimentary Geology, vol.10, pp.25-60, 1973.

P. Freytet, J. Plaziat, and E. P. Verrecchia, A classification of rhizogenic (root-formed) calcretes, with examples from the Upper Jurassic-Lower Cretaceous of Spain and Upper Cretaceous of southern France-Discussion, Sedimentary geology, vol.110, pp.299-303, 1997.

P. Freytet and E. P. Verrecchia, Lacustrine and palustrine carbonate petrography: an overview, Journal of Paleolimnology, vol.27, pp.221-237, 2002.

F. Gasse, Hydrological changes in the African tropics since the Last Glacial Maximum, Quaternary Science Reviews, vol.19, pp.189-211, 2000.

F. Gasse, F. Chalié, A. Vincens, M. A. Williams, and D. Williamson, Climatic patterns in equatorial and southern Africa from 30,000 to 10,000 years ago reconstructed from terrestrial and nearshore proxy data, Quaternary Science Reviews, vol.27, pp.2316-2340, 2008.

. Geotechnica, Elobeid Water Supply Project, Feasibility study of Groundwater Resources, 1985.

S. Gilmore and K. D. Hill, Relationships of the Wollemi Pine (Wollemia nobilis) and a molecular phylogeny of the Araucariaceae, Telopea, vol.7, issue.3, pp.275-291, 1997.

M. A. Ginaya, A. Elkrail, and . A. Farawa, Numerical Modelling of the Groundwater flow in Ennahud Basin, Northern Kordofan-Sudan. Research and Applications (IJERA), vol.3, pp.217-222, 2013.

M. A. Ginaya, . Sudan, T. C. Goji, and A. E. Ayodele, Foliar epidermal and Pollen Characters in the genus Cola Schott. & Endl. in Nigeria, Ministry of Sciences & Technology, vol.2, pp.57-63, 2005.

G. A. Goodfriend, M. Magaritz, and J. R. Gat, Stable isotope composition of land snail body water and its relation to environmental waters and shell carbonate, Geochimica et Cosmochimica Acta, vol.53, pp.3215-3221, 1989.

W. D. Gosling, C. S. Miller, and D. A. Livingstone, Atlas of the tropical West African pollen flora, Review of Palaeobotany and Palynology, vol.199, pp.1-135, 2013.

W. Gossel, A. M. Ebraheem, and P. Wycisk, A very large scale GIS-based groundwater flow model for the Nubian sandstone aquifer in Eastern Sahara, 2004.

H. Journal12, , pp.698-713

Y. Hamann, W. Ehrmann, G. Schmiedl, G. Kuhnt, and T. , Modern and late Quaternary clay mineral distribution in the area of the SE Mediterranean Sea, Quaternary Research, vol.71, pp.453-464, 2009.

M. A. Hamdan, A. A. Refaat, E. Abu-anwar, and N. A. Shallaly, Source of the aeolian dune sand of Toshka area, southeastern Western Desert, Egypt. Aeolian Research, vol.17, pp.275-289, 2014.

F. Hassan, Chronology of the Khartoum 'Mesolithic' and 'Neolithic' and related sites in the Sudan: statistical analysis and comparisons with Egypt, The African Archaeological Review, vol.4, pp.83-102, 1986.

M. Hesse and R. Zetter, The fossil pollen record of Araceae, Plant Systematics and Evolution, vol.263, pp.93-115, 2007.

A. Hewaidy, O. R. Elshahat, and S. Kamal, Stratigraphy, facies analysis and depositional environments of the Upper Unit of Abu, Journal of African Earth Sciences, vol.139, pp.26-37, 2018.

C. L. Hill, Geologic Contexts of the Acheulian (Middle Pleistocene) in the Eastern Sahara, Geoarchaeology: An International Journal, vol.16, pp.65-94, 2001.

P. Hoelzmann, B. Keding, H. Berke, S. Kröpelin, and H. Kruse, Environmental change and archeology: lake evaluation and human occupation in the Eastern Sahara during the Holocene, Palaeogeography, Palaeoclimatology, Palaeoecology, vol.169, pp.193-217, 2001.

P. Hoelzmann, A. Schwalb, N. Roberts, P. Cooper, and A. Burgess, Hydrological response of an east-Saharan palaeolake (NW Sudan) to early-Holocene climate, The Holocene, vol.20, issue.4, pp.537-549, 2010.

H. Hong, C. Wang, K. Zeng, Y. Gu, Y. Wu et al., Geochemical constraints on provenance of the mid-Pleistocene red earth sediments in subtropical China, Sedimentary Geology, vol.290, pp.97-108, 2013.

R. Idohou, C. A. Djagoun, B. Kassa, A. E. Assogbadjo, C. Codjia et al., Soil factors affecting density of three giant land snail species in different habitats of DassaZoumè district (Central Benin), 2013.

. Iucn, A. Jorgensen, T. K. Kristensen, A. Stensgaard, and D. Van-damme, The IUCN Red List of Threatened Species: Pila wernei

D. Van-damme, A. Jorgensen, T. K. Kristensen, C. Lange, A. Stensgaard et al., The IUCN Red List of Threatened Species: Bulinus truncatus, IUCN, 2012.

J. F. , Early Khartoum ceramics in the Wadi Howar (Northwest Sudan), in : Recent research into the Stone Age of northeastern Africa, The IUCN Red List of Threatened Species: Radix natalensis, vol.7, pp.77-87, 2000.

S. J. Jung, G. R. Davies, G. M. Ganssen, and D. Kroon, Stepwise Holocene aridifcation in NE Africa deduced from dust-borne radiogenic isotope records, Earth and Planetary Science Letters, vol.221, pp.27-37, 2004.

B. G. Karkanis, Hydrochemical Facies of Grounwater in the Western Provinces of Sudan, 1966.

S. J. Kemp, M. A. Ellis, I. Mouteney, and S. Kender, Palaeoclimatic implications of high resolution clay mineral assemblages preceding and across the onset of the Palaeocene-Eocene Thermal Maximum, Clay Minerals, vol.51, pp.793-813, 2016.

P. Kershaw and B. Wagstaff, The southern conifer family Araucariaeacae: History, Status, and Value for Paleoenvironmental Reconstruction, Annu. Rev. Ecol. Syst, vol.32, pp.397-414, 2001.

C. F. Klappa, Rhizoliths in terrestrial carbonates: classification, recognition, genesis and significance, Sedimentology, vol.27, pp.611-627, 1980.

F. Köhler and M. Glaubrecht, The Types of Ampullariidae Gray, 1824 (Mollusca, Gastropoda) in the Malacological Collection of the Natural History Museum, Berlin: an annotated catalogue with lectotype designations, Mitt. Mus. Nat. kd. Berlin, Zool. Reihe, vol.82, issue.1, pp.198-215, 2006.

S. Kröpelin and I. Soulié-märche, Charophyte remains from Wadi Howar as evidence for deep MidHolocene freshwater lakes in the Eastern Sahara of Northwest Sudan, Quaternary research, vol.36, pp.210-223, 1991.

S. Kröpelin, D. Verschuren, A. Lézine, H. Eggermont, C. Cocquyt et al., Climate-Driven Ecosystem Succession in the Sahara: The Past 6000 Years, Science, vol.320, pp.765-768, 2008.

A. Kumar, New palynological evidence for the age of the Beda Formation, Palaeontologia Electronica, vol.19, pp.1-14, 2016.

R. Kuper and S. Kröpelin, Climate-Controlled Holocene Occupation in the Sahara: Motor of Africa's Evolution, Science, vol.313, pp.803-807, 2006.

J. Lario, S. Sanshez-moral, V. Fernandez, A. Jimeno, and M. Menendez, Paleoenvironmental Evolution of the Blue Nile (central Sudan) During the Early and Mid-Holocene (MesolithicNeolithic Transition), Quaternary Science Reviews, vol.16, pp.583-588, 1997.

L. M. Larsson, V. Vajda, and E. S. Rasmussen, Early Miocene pollen and spores from western Jylland, Denmark-environmental and climatic implications, GFF, vol.128, pp.261-272, 2006.

A. Lézine, W. Zheng, P. Braconnot, and G. Krinner, A late Holocene pollen and climate record from Lake Yoa, northern Chad, Clim. Past Discuss, vol.7, pp.2413-2444, 2011.

W. Lotfy and L. M. Lotfy, Synopsis of the Egyptian freshwater snail fauna, Folia Malacol, vol.23, issue.1, pp.19-40, 2015.

Y. Lu, Y. Hautevelle, and R. Michels, Determination of the molecular signature of fossil conifers by experimental palaeochemotaxonomy-Part 1: The Araucariaceae family, Biogeosciences Discuss, vol.9, pp.10513-10550, 2012.
URL : https://hal.archives-ouvertes.fr/hal-01303028

J. Lukavský, Algae, Cyanobacteria and Chytridiales of Èerné Lake inthe Bohemian Forest, Silva Gabreta, vol.15, issue.1, pp.1-48, 2009.

J. Madhavaraju, S. Ramasamy, A. Ruffell, and S. P. Mohan, Clay mineralogy of the Late Cretaceous and early Tertiary successions of the Cauvery Basin (southeastern India): implications for sediment source and palaeoclimates at the K/T boundary, Cretaceous Research, vol.23, pp.153-163, 2002.

J. Madhavaraju, E. Ramírez-montoya, R. Monreal, C. M. González-león, T. Pi-puig et al., Paleoclimate, paleoweathering and paleoredox conditions of Lower Cretaceous shales from the Mural Limestone, Tuape section, Constraints from clay mineralogy and geochemistry: Revista Mexicana de Ciencias Geológicas, vol.33, pp.34-48, 2016.

H. Y. Madukwe and R. A. Obasi, Geochemistry, Classification and maturity of the Sandstone facies of the Abeokuta Formation, south west Nigeria, European Journal of Basic and Applied Sciences, vol.3, issue.2, pp.2059-3058, 2016.

M. Magendantz, The biology of Biomphalaria choanomphala and B. sudanica in relation to their role in the transmission of Schistosoma mansoni in Lake Victoria at Mwanza, Tanzania. Bull. Org. mond. Santé (Bull. Wld. Hlth Org.), vol.47, pp.331-342, 1972.

E. A. Malek, D. S. Brown, T. Fison, V. R. Southgate, and C. A. Wright, Distribution of the intermediate hosts of bilharziasis in relation to hydrography. With special reference to the Nile basin and the Sudan, Hydrobiologia, vol.110, pp.247-271, 1958.

A. W. Mann and R. C. Horwitz, Palaeoenvironmental significance of palustrine carbonates and calcretes in the geological record, Earth Science Reviews, vol.60, pp.261-298, 1979.

K. Manning and A. Timpson, The demographic response to Holocene climate change in the Sahara, Quaternary Science Reviews, vol.101, pp.28-35, 2014.

A. E. Marks, A. S. Mohammed-ali, and A. M. Khabir, The Wavy Line and the Dotted Wavy Line Pottery in the Prehistory of the Central Nile and the Sahara-Sahel Belt, African Archaeological Review, vol.20, issue.1, pp.25-58, 1991.

A. S. Mohammed-ali, A. S. Mohammed-ali, and A. M. Khabir, The Wavy Line and the Dotted Wavy Line Pottery in the Prehistory of the Central Nile and the Sahara-Sahel Belt, African Archaeological Review, vol.20, issue.1, pp.25-58, 1991.

A. S. Mohammed-ali and A. M. Khabir, The Wavy Line and the Dotted Wavy Line Pottery in the Prehistory of the Central Nile and the Sahara-Sahel Belt, African Archaeological Review, vol.20, issue.1, pp.25-58, 2003.

P. Monga, M. Kumar, V. Prasad, and Y. Joshi, Palynostratigraphy, palynofacies and depositional environment of a lignite-bearing succession at Surkha Mine, Cambay Basin, north-western India, Acta Palaeobotanica, vol.55, issue.2, pp.183-207

R. Muscheler, J. Beer, G. Wagner, C. Laj, C. Kissel et al., Changes in the carbon cycle during the last deglaciation as indicated by the comparison of 10 Be and 14 C records, Earth and Planetary Science Letters, vol.219, pp.325-340, 2004.
URL : https://hal.archives-ouvertes.fr/in2p3-00025055

O. Nawari and K. Schetelig, Chemical, mineralogical and fabric properties of Kordofan tropical black soils (Sudan), Journal of Aft-lean Earth Sciences, vol.14, issue.4, pp.545-55, 1992.

G. T. Ndifon and F. M. Ukoli, Ecology of freshwater snails in south-western Nigeria. I: Distribution and habitat preferences, Hydrobiologia, vol.171, pp.231-253, 1989.

H. W. Nesbitt and G. M. Young, Early Proterozoic climates and plate motions inferred from major elements chemistry of lutites, Nature, vol.299, pp.715-717, 1982.

P. Netterberg, Palaeoenvironmental significance of palustrine carbonates and calcretes in the geological record, Earth Science Reviews, vol.60, pp.261-298, 1969.

E. Neubert, Annotated checklist of the terrestrial and freshwater molluscs of the Arabian Peninsula with descriptions of new species, IUCN, 2017. The IUCN Red List of Threatened Species: Radix natalensis, 1998.

. J. Ngaira, Impact of climate change on agriculture in Africa by 2030, Scientific Research and Essays, vol.2, issue.7, pp.238-243, 2007.

S. Nicholson and H. Flohn, African Environmental and Climatic changes and the general atmospheric circulation in Late Pleistocene and Holocene, Climatic Change, vol.2, pp.313-348, 1980.

K. Nicoll, Recent environmental change and prehistoric human activity in Egypt and Northern Sudan, Quaternary Science Reviews, vol.23, pp.561-580, 2004.

C. A. Offord, Pushed to the limit: consequences of climate change for the Araucariaceae: a relictual rain forest family, Annals of Botany, vol.108, pp.347-357, 2011.

I. N. Oloto, Y. Yikarebogha, and A. O. Omoboriowo, Palynological studies of Upper Cretaceous succession of Herwa-1 well, central Chad Basin, Nigeria. International Journal of Innovative Research in Science, Engineering and Technology, vol.2, issue.4, pp.880-889, 2013.

S. O. Oni and A. S. Olatuji, Depositional environments signatures, maturity and source weathering of Niger ddelta sediments from an oil well in southeastern Delta State, Eurasian Journal of Soil Scince, vol.6, issue.3, pp.259-274, 2017.

U. S. Onoduku and E. Okosun, Palynology, Palynostratigraphy and Paleoenvironmental Analysis of Maiganga Coal Mine, Gombe Formation, Nigeria. Universal Journal of Geoscience, vol.2, issue.3, pp.93-103, 2014.

H. J. Pachur, H. J. Pachur, and S. Kröpelin, Wadi Howar: Paleoclimatic Evidence from an Extinct River System in the Southeastern Sahara, SCIENCE, vol.237, pp.298-300, 1974.

H. J. Pachur and P. Hoelzmann, Paleoclimatic Implications of Late Quaternary Lacustrine Sediments in Western Nubia, Sudan. Quaternary Research, vol.36, pp.257-216, 1991.

H. J. Pachur and S. Kröpelin, Wadi Howar: Paleoclimatic Evidence from an Extinct River System in the Southeastern Sahara, J. Veg. Sci, vol.237, pp.793-804, 1987.

A. Perveen, A Palynological Survey of Aquatic Flora of Karachi-Pakistan, Tr. J. of Botany, vol.23, pp.309-317, 1999.

J. Peters, The Late Prehistory of the Eastern Sahel-the Mesolithic and Neolithic of Shaqadod, 1991.

H. I. Petersen, S. Lindström, H. P. Nytoft, and P. Rosenberg, Composition, peat-forming vegetation and kerogen paraffinicity of Cenozoic coals: Relationship to variations in the petroleum generation potential (Hydrogen Index), International Journal of Coal Geology, vol.78, pp.119-134, 2009.

H. A. Pilsbry, J. Bequaert, K. N. De-kock, and C. T. Wolmarans, Distribution and habitats of Melanoides tuberculata (Müller, 1774) and M. victoriae (Dohrn, 1865) (Mollusca: Prosobranchia: Thiaridae) in South Africa, Water SA, vol.35, issue.5, pp.713-720, 1927.

P. E. Potter, H. Y. Madukwe, and R. A. Obasi, Geochemistry, Classification and maturity of the Sandstone facies of the Abeokuta Formation, south west Nigeria, European Journal of Basic and Applied Sciences, vol.86, issue.4, pp.2059-3058, 1978.

S. Priene and H. Lavee, Soil organic matter and degradation. Soil conservation and protection for Europe (SCOPE), 2015.

S. Qamar, A. Saif, and J. Altaf, Identification of the species of genus Zootecus on the basis of morphology, Journal of Biodiversity and Environmental Sciences, vol.11, issue.3, pp.122-127, 2017.

L. J. Quick, Late Quaternary palaeoenvironments of the southern Cape, South Africa: palynological evidence from three coastal wetlands, 2013.

M. R. Rao, A. Sahni, R. S. Rana, and P. Verma, Palynostratigraphy and depositional environment of Vastan Lignite Mine, J. Earth Syst. Sci, vol.122, issue.2, pp.289-307, 2013.

S. K. Raut, G. M. Barker, S. K. Tan, and M. E. Low, An update on the West African Limicolaria flammea (Müller, 1774) in Singapore, and its distinction from the confamilial Achatina fulica Bowdich, 1822 (Mollusca: Grastropoda: Achatinidae), Nature in Singapore, vol.4, pp.311-317, 2002.

J. B. Riding, M. J. Leng, S. Kender, S. P. Hesselbo, and S. Feist-burkhardt, Isotopic and palynological evidence for a new Early Jurassic environmental perturbation, Palaeogeography, Palaeoclimatology, Palaeoecology, vol.374, pp.16-27, 2013.
DOI : 10.1016/j.palaeo.2012.10.019

URL : https://doi.org/10.1016/j.palaeo.2012.10.019

H. G. Rodis, A. Hassan, and L. Wahadan, Groundwater Geology of Kordofan Province-Sudan, Contribution to the hydrology of Africa and Mediterranean Region. United State government printing office, 1968.

D. Rodrigues, P. I. Abell, and S. Kröpelin, Seasonality in the early Holocene climate of Northwest Sudan: interpretation of Etheria elliptica shell isotopic data, Global and Planetary Change, vol.26, pp.181-187, 2000.

K. Rozanski, L. Araguás-araguás, and R. Gonfiantini, Isotopic patterns in modern global precipitation, Geophysical Monograph, vol.78, pp.1-36, 1993.
DOI : 10.1029/gm078p0001

V. Rull, A quantitative palynological record from the Early Miocene of western Venezuela, with emphasis on mangroves, Palynology, vol.25, pp.109-126, 2001.

R. Salama, Directorate for groundwater and Wadi water resources, Ministry of irrigation and water resources, 1976.

M. Salard-cheboldaeff, Intertropical African Palynostratigraphy from Cretaceous to Late Quaternary times, Journal of African Earth Sciences, vol.11, pp.1-9, 1990.

L. Scavati, D. L. Melendi, and W. Volkheimer, A Danian subtropical lacustrine palynobiota from South America (Bororó Formation, Geologica Acta, vol.7, issue.12, pp.35-61, 2009.

E. Schrank, Nonmarine Cretaceous palynology of northern Kordofan, Sudan, with notes on fossil Salviniales (water ferns), Geol. Rundschau, vol.83, pp.773-786, 1994.

R. Schultheiß, O. W. Ndeo, M. Malikwisha, C. Marek, U. Bößneck et al., Freshwater molluscs of the Eastern Congo: notes on taxonomy, biogeography and conservation, African Invertebrates, vol.52, issue.2, pp.265-284, 2011.

L. Scott, B. Cooremans, J. S. De-wet, and J. C. Vogel, Holocene environmental changes in Namibia inferred from pollen analysis of swamp and lake deposits, Holocene, vol.1, pp.8-13, 1991.

T. Shanahan, N. P. Mckay, K. A. Hughen, J. T. Overpeck, B. Otto-bliesner et al., The time-transgressive termination of the African Humid Period, Nature Geoscience, vol.8, pp.140-144, 2015.

N. D. Sheldon, G. J. Retallack, and S. Tanaka, Geochemical climofunctions from North American soils and application to paleosols across the Eocene-Oligocene boundary in Oregon, Journal of Geology, vol.110, pp.687-696, 2002.

N. D. Sheldon and N. J. Tabor, Quantitative paleoenvironmental and paleoclimatic reconstruction using paleosols, Earth-Science Reviews, vol.95, pp.1-52, 2009.

N. D. Smith, The Braided Stream Depositional Environment: Comparison of the Platte River with Some Silurian Clastic Rocks, vol.81, pp.2993-3014, 1970.

K. R. Smythe and M. D. Gallagher, The IUCN Red List of Threatened Species: Radix natalensis, 1977.

E. A. Soares, R. Dino, D. P. Soares, L. Antonioli, and M. A. Da-silva, New sedimentological and palynological data from surface Miocene strata in the central Amazonas Basin area, Brazilian Journal of Geology, vol.45, issue.3, pp.337-357, 2015.

. Strojexport, Geophysical Investigations of Groundwater Structures, Western of Kordofan Province and Eastern part of Darfur Province, Fourth stage. Ministry of Agriculture food and Natural Resources, Rural Water Corporation, 1976.

S. Stukins, D. W. Jolley, D. Mcilroy, and A. J. Hartley, Middle Jurassic vegetation dynamics from allochthonous palynological assemblages: An example from a marginal marine depositional setting; Lajas Formation, Palaeogeography, Palaeoclimatology, Palaeoecology, vol.392, pp.117-127, 2013.

B. I. Szabo, C. V. Haynes, and T. A. Maxwell, Ages of Quaternary pluvial episodes determined by uranium-series and radiocarbon dating of lacustrine deposits of Eastern Sahara, Palaeogeography, Palaeoclimatology, Palaeoecology, vol.113, pp.227-242, 1995.

S. K. Tan and G. R. Clements, Limicolaria flammea (Müller, 1774), another potentially invasive African land snail in tropical Asia, Tropical Conservation Science, vol.4, issue.1, pp.97-102, 2011.

S. K. Tan and M. E. Low, An update on the West African Limicolaria flammea (Müller, 1774) in Singapore, and its distinction from the confamilial Achatina fulica Bowdich, 1822 (Mollusca: Grastropoda: Achatinidae), Nature in Singapore, vol.4, pp.311-317, 2011.

S. M. Tchouatcha, N. Ricard, M. M. Said, D. A. Ekodeck, and E. G. , Existence of "late continental" deposits in the Mbere and Djerem sedimentary basins (North Cameroon): Palynologic and stratigraphic evidence, Journal of Geology and Mining Research, vol.2, issue.6, pp.159-169, 2010.

R. J. Telford and H. F. Lamb, Groundwater-Mediated response to Holocene climatic change recorded by the Diatom Stratigraphy of an Ethiopian Crater Lake, Quaternary Research, vol.52, pp.63-75, 1999.

J. E. Tierney, S. C. Lewis, B. I. Cook, A. N. Legrande, and G. A. Schmidt, Model, proxy and isotopic perspectives on the East African Humid Period, Earth and Planetary Science Letters, vol.307, pp.103-112, 2011.

J. E. Tierney, J. M. Russell, J. S. Sinninghe-damsté, Y. Huang, and D. Verschuren, Late Quaternary behavior of the East African monsoon and the importance of the Congo Air Boundary, Quaternary Science Reviews, vol.30, pp.798-807, 2011.

J. A. Tóth, K. Lajtha, Z. Kotroczó, Z. Krakomperger, B. Caldwell et al., The effect of climate change on soil Organic Matter decomposition, Acta Silv. Lign. Hung, vol.3, pp.75-85, 2007.

A. Traverse, , 2008.

S. K. Tripathi, M. Kumar, and D. Srivastava, Palynology of Lower Palaeogene (ThanetianYpresian) coastal deposits from the Barmer Basin (Akli Formation, Palaeoenvironmental and palaeoclimatic implications. Geologica Acta, vol.7, pp.147-160, 2009.

M. E. Tucker, , p.234, 2003.

M. E. Tucker and V. P. Wright, Carbonate sedimentology. Blackwell Science, vol.482, 1990.

D. E. Uzodimma, Palynostratigraphy, Age Determination and Depositional Environments of the Imo Shale Exposures at the Okigwe/Port Harcourt Express Road Junction Okigwe, Southeastern Nigeria, Greener Journal of Physical Sciences, vol.3, issue.7, pp.255-272, 2013.

H. Van-der-schalie, A new species of Caracolus (Pulmonata: Camaenidae) from the Oligocene of Nebraska and the biotic history of the American camaenid land snails, Zoological Journal of the Linnean Society, vol.67, pp.269-284, 1948.

G. Visher, Grain size distributions and depositional processes, Journal of Sedimentary Petrology, vol.39, pp.1074-1106, 1969.

W. Volkhemer, O. W. Rauhut, M. E. Quattrocchio, and M. A. Martinez, Jurasic paleoclimate in Argentina, a review, Revista de la Asociación Geológica Argentina, vol.63, issue.4, pp.549-556, 2008.

B. Volkoff, Red and lateritic soils: World scenario, Managing Red and Lateritic Soils for Sustainable Agriculture, vol.1, pp.57-74, 1988.

M. Walker, Quaternary dating methods, p.286, 2005.

M. J. Warren, Tectonic Inversion and Petroleum System Implications in the Rifts of Central Africa. Frontiers + Innovation CSPG CSEG CWLS Convention, pp.461-464, 2009.

G. Webbe, The biology of Biomphalaria choanomphala and B. sudanica in relation to their role in the transmission of Schistosoma mansoni in Lake Victoria at Mwanza, Tanzania. Bull. Org. mond. Santé (Bull. Wld. Hlth Org.), vol.47, pp.331-342, 1962.

A. J. Whiteman, The Geology of the Sudan Republic, 1971.

M. A. Williams, M. Talbot, P. Aharon, Y. Salaam, F. Williams et al., Abrupt return of the summer monsoon 15,000 years ago: new supporting evidence from the lower White Nile valley and Lake Albert, Quaternary Science Reviews, vol.25, pp.2651-2665, 2006.

M. Williams, Climate Change in Deserts (past, present and future), 2014.

M. A. Williams, Late Pleistocene and Holocene environments in the Nile basin, Global and Planetary Change, vol.69, pp.1-15, 2009.

M. A. Williams, D. Adamson, B. Cock, and R. Mcevedy, Late Quaternary environments in the White Nile region, Global and Planetary Change, vol.26, pp.305-316, 2000.

M. A. Williams and D. A. Adamson, Late Pleistocene desiccation along the White Nile, Nature, vol.248, pp.584-586, 1974.

S. N. Williams, P. J. Hunter, D. S. Brown, T. Fison, V. R. Southgate et al., Aquatic snails of the Jonglei region, southern Sudan, and transmission of trematode parasites, Hydrobiologia, vol.110, pp.247-271, 1968.

M. J. Wilson, The origin and formation of clay minerals in soils: past, present, and feature prospectives, Clay minerals, vol.34, pp.7-25, 1999.

J. C. Woodward, M. G. Macklin, D. Michael, D. Krom, and M. A. Williams, The Nile: Evolution, Quaternary River Environments and Material Fluxes, 2007.

E. Worobiec, Fossil zygospores of Zygnemataceae and other microremains of freshwater algae from two Miocene palaeosinkholes in the Opole region, Acta Palaeobotanica, vol.54, pp.113-157, 2014.

E. Worobiec and P. Gedl, Spore-pollen and phytoplankton analysis of the Upper Miocene deposits from Józefina, Geological Quarterly, vol.54, issue.1, pp.41-54, 2010.

E. Worobiec and G. Worobiec, Miocene palynoflora from the KRAM-P 218 leaf assemblage from the Be?chatów Lignite Mine (Central Poland), Acta Palaeobotanica, vol.56, issue.2, pp.499-517, 2016.

. J. Ybert, Atlas de pollen de Côte D'Ivoire. Initiations-Documentations Techniques No 40. Office de la Recherche Scientifique et Technique Outre-Mer, 1979.

R. Zetter, M. Hesse, and A. Frosch-radivo, Early Eocene zona-aperturate pollen grains of the Proxapertites type with affinity to Araceae, Review of Paleaobotany and Palynology, vol.117, pp.267-279, 2011.