M. Abramoff, P. Magalhaes, and S. Ram, Image Processing with ImageJ. Biophotonics International, vol.11, pp.36-42, 2004.

E. L. Allan, S. T. Ambrose, N. B. Richoux, and P. W. Froneman, Determining spatial changes in the diet of nearshore suspension-feeders along the South African coastline: Stable isotope and fatty acid signatures, Estuarine Coastal and Shelf Science, vol.87, pp.463-471, 2010.

W. G. Ambrose, M. L. Carroll, M. Greenacre, S. R. Thorrold, and K. W. Mcmahon, Variation in Serripes groenlandicus (Bivalvia) growth in a Norwegian high-Arctic fjord: evidence for local-and large-scale climatic forcing, Global Change Biology, vol.12, pp.1595-1607, 2006.

W. G. Ambrose, P. E. Renaud, W. L. Locke, F. R. Cottier, J. Berge et al., Growth line deposition and variability in growth of two circumpolar bivalves (Serripes groenlandicus and Clinocardium ciliatum), Polar Biology, vol.35, pp.345-354, 2012.

M. Anderson, R. Gorley, and K. Clarke, PERMANOVA+ for PRIMER: guide to software and statistical methods, 2008.

K. R. Arrigo, D. K. Perovich, R. S. Pickart, Z. W. Brown, G. L. Van-dijken et al., Massive Phytoplankton Blooms Under Arctic Sea Ice. Science, vol.336, pp.1408-1408, 2012.

K. R. Arrigo, G. Van-dijken, and S. Pabi, Impact of a shrinking Arctic ice cover on marine primary production, Geophysical Research Letters, vol.35, 2008.

K. M. Attard, K. Hancke, M. K. Sejr, and R. N. Glud, Benthic primary production and mineralization in a High Arctic fjord: in situ assessments by aquatic eddy covariance, Marine Ecology Progress Series, vol.554, pp.35-50, 2016.

P. Bachelard-s-;-maloine, A. Barats, D. Amouroux, L. Chauvaud, C. Pecheyran et al., High frequency Barium profiles in shells of the Great Scallop Pecten maximus: a methodical long-term and multi-site survey in Western Europe, Quelques aspects des notions de modèle et de justification des modèles, vol.6, pp.157-170, 1979.

B. Barber and N. Blake, Energy storage and utilization in relation to gametogenesis in Argopecten irradians concentricus (Say), Journal of Experimental Marine Biology and Ecology, vol.52, pp.121-134, 1981.

G. Barnathan, Non-methylene-interrupted fatty acids from marine invertebrates: Occurrence, characterization and biological properties, Biochimie, vol.91, pp.671-678, 2009.
DOI : 10.1016/j.biochi.2009.03.020

I. Bartsch, M. Paar, S. Fredriksen, and C. Wiencke, Changes in kelp forest biomass and depth distribution at Kongsfjorden (Spitsbergen) between 1996/98 and 2012-2014 reflect Arctic warming, European Journal of Phycology, vol.50, pp.105-105, 2015.
DOI : 10.1007/s00300-015-1870-1

S. J. Batenburg, G. J. Reichart, T. Jilbert, M. Janse, F. P. Wesselingh et al., Interannual climate variability in the Miocene: High resolution trace element and stable isotope ratios in giant clams, Palaeogeography Palaeoclimatology Palaeoecology, vol.306, pp.75-81, 2011.

A. Bec, M. Perga, A. Koussoroplis, G. Bardoux, C. Desvilettes et al., Assessing the reliability of fatty acid-specific stable isotope analysis for trophic studies, Methods in Ecology and Evolution, vol.2, pp.651-659, 2011.

L. Beierlein, O. Salvigsen, B. R. Schone, A. Mackensen, and T. Brey, The seasonal water temperature cycle in the Arctic Dicksonfjord (Svalbard) during the Holocene Climate Optimum derived from subfossil Arctica islandica shells, Holocene, vol.25, pp.1197-1207, 2015.

J. Bendtsen, K. Gustafsson, S. Rysgaard, and T. Vang, Physical conditions, dynamics and model simulations during the ice-free period of the Young Sound/Tyrolerfjord system, Meddelelser om Grønland Bioscience, vol.58, 2007.

J. J. Beukema, E. Knol, and G. C. Cadee, Effects of temperature on the length of the annual growing-season in the tellinid bivalve Macoma balthica living on tidal flats in the Dutch Wadden Sea, Journal of Experimental Marine Biology and Ecology, vol.90, pp.129-144, 1985.

J. T. Bhaskar, S. C. Tripathy, P. Sabu, C. M. Laluraj, and S. Rajan, Variation of phytoplankton assemblages of Kongsfjorden in early autumn 2012: a microscopic and pigment ratio-based assessment, Environmental Monitoring and Assessment, vol.188, 2016.

K. L. Bice, M. A. Arthur, and L. Marincovich, Late Paleocene Arctic Ocean shallow-marine temperatures from mollusc stable isotopes, Paleoceanography, vol.11, pp.241-249, 1996.

S. R. Birkely, O. Grahl-nielsen, and B. Gulliksen, Temporal variations and anatomical distributions of fatty acids in the bivalve Mya truncata, L. 1758, from Isfjorden, Spitsbergen. Polar Biology, vol.26, pp.83-92, 2003.

B. A. Black, Climate-driven synchrony across tree, bivalve, and rockfish growthincrement chronologies of the northeast Pacific, Marine Ecology Progress Series, vol.378, pp.37-46, 2009.

J. Bland and D. Altman, Cronbach's alpha, British Medical Journal, vol.314, p.572, 1997.

M. E. Blicher, S. Rysgaard, and M. K. Sejr, Growth and production of sea urchin Strongylocentrotus droebachiensis in a high-Arctic fjord, and growth along a climatic gradient (64 to 77 degrees N), Marine Ecology Progress Series, vol.346, pp.314-314, 2007.

E. Bligh and W. Dyer, A rapid method of total lipid extraction and purification, Canadian Journal of Biochemistry and Physiology, vol.37, pp.911-917, 1959.

E. W. Born, S. Rysgaard, G. Ehlmé, and M. Sejr, Underwater observations of foraging freeliving Atlantic walruses (Odobenus rosmarus rosmarus) and estimates of their food consumption, Polar Biology, vol.26, pp.348-357, 2003.

L. Bougeois, M. De-rafelis, G. J. Reichart, L. J. De-nooijer, F. Nicollin et al., A high resolution study of trace elements and stable isotopes in oyster shells to estimate Central Asian Middle Eocene seasonality, Chem Geol, vol.363, pp.200-212, 2014.
URL : https://hal.archives-ouvertes.fr/insu-00913050

T. Brey, Population dynamics in benthic invertebrates. A virtual handbook, 2001.

S. Brockington and A. Clarke, The relative influence of temperature and food on the metabolism of a marine invertebrate, Journal of Experimental Marine Biology and Ecology, vol.258, pp.87-99, 2001.

T. A. Brown and S. T. Belt, Identification of the sea ice diatom biomarker IP25 in Arctic benthic macrofauna: direct evidence for a sea ice diatom diet in Arctic heterotrophs, Polar Biology, vol.35, pp.131-137, 2012.

T. A. Brown, D. J. Yurkowski, S. H. Ferguson, C. Alexander, and S. T. Belt, H-Print: a new chemical fingerprinting approach for distinguishing primary production sources in Arctic ecosystems, Environmental Chemistry Letters, vol.12, pp.387-392, 2014.

C. Buchholz and C. Wiencke, Working on a baseline for the Kongsfjorden food web: production and properties of macroalgal particulate organic matter (POM), Polar Biology Budge SM, vol.22, pp.759-801, 2006.

S. M. Budge, S. W. Wang, T. E. Hollmén, and M. J. Wooller, Carbon isotopic fractionation in eider adipose tissue varies with fatty acid structure: implications for trophic studies, The Journal of Experimental Biology, vol.214, pp.3790-3800, 2011.
DOI : 10.1242/jeb.057596

URL : http://jeb.biologists.org/content/214/22/3790.full.pdf

S. M. Budge, M. J. Wooller, A. M. Springer, S. J. Iverson, C. P. Mcroy et al., Tracing carbon flow in an arctic marine food web using fatty acid-stable isotope analysis, Oecologia, vol.157, pp.117-129, 2008.
DOI : 10.1007/s00442-008-1053-7

M. Cabanellas-reboredo, S. Deudero, and A. Blanco, Stable-isotope signatures (delta c13 and delta n-15) of different tissues of Pinna nobilis Linnaeus, 1758 (Bivalvia): Isotopic variations among tissues and between seasons, Journal of Molluscan Studies, vol.75, pp.343-349, 2009.

S. E. Campana, Accuracy, precision and quality control in age determination, including a review of the use and abuse of age validation methods, Journal of Fish Biology, vol.59, pp.197-242, 2001.

S. E. Campana and C. M. Jones, Radiocarbon from nuclear testing applied to age validation of black drum, Pogonias cromis, Fishery Bulletin, vol.96, pp.185-192, 1998.

E. Canobbio, Atlas des pôles, 2007.

B. Carell, S. Forberg, E. Grundelius, L. Henrikson, A. Johnels et al., Can mussel shells reveal environmental history, Ambio, vol.16, pp.2-10, 1987.

M. L. Carroll, W. G. Ambrose, B. S. Levin, W. L. Locke, G. A. Henkes et al., Pan-Svalbard growth rate variability and environmental regulation in the Arctic bivalve Serripes groenlandicus, Journal of Marine Systems, vol.88, pp.239-251, 2011.

M. L. Carroll, W. G. Ambrose, B. S. Levin, S. K. Ryan, A. R. Ratner et al., Climatic regulation of Clinocardium ciliatum (Bivalvia) growth in the northwestern Barents Sea, Palaeogeography Palaeoclimatology Palaeoecology, vol.302, pp.10-20, 2011.

M. L. Carroll, W. G. Ambrose, V. Locke, . Wl, S. K. Ryan et al., Bivalve growth rate and isotopic variability across the Barents Sea Polar Front, Journal of Marine Systems, vol.130, pp.167-180, 2014.

M. L. Carroll, J. Carroll, B. J. Johnson, G. A. Henkes, K. W. Mcmahon et al., Bivalves as indicators of environmental variation and potential anthropogenic impacts in the southern Barents Sea, Biogeochemistry of Marine Systems, vol.59, pp.193-206, 2003.

L. Chauvaud, A. Lorrain, R. B. Dunbar, Y. M. Paulet, G. Thouzeau et al., Shell of the Great Scallop Pecten maximus as a highfrequency archive of paleoenvironmental changes, Geochemistry Geophysics Geosystems, vol.6, 2005.
URL : https://hal.archives-ouvertes.fr/hal-00091298

M. E. Chrpa, Morphology of Astarte borealis (mollusca: Bivalvia) of Camden Bay, Northern Alaska Florida Atlantic University Clarke A (1988) Seasonality in the Antarctic marine environment, Comparative Biochemistry and Physiology B-Biochemistry & Molecular Biology, vol.90, pp.461-473, 2013.

K. Clarke and R. Gorley, PRIMER v6: user manual/tutorial. PRIMER-E, Plymouth Clarke KR (1993) Nonparametric multivariate analyses of changes in community structure, Australian Journal of Ecology, vol.18, pp.117-143, 2006.

R. A. Collins and R. H. Cruickshank, The seven deadly sins of DNA barcoding, Molecular Ecology Resources, vol.13, pp.969-975, 2013.

J. C. Comiso, C. L. Parkinson, R. Gersten, and L. Stock, Accelerated decline in the Arctic Sea ice cover, Geophysical Research Letters, vol.35, 2008.

F. R. Cottier, F. Nilsen, M. E. Inall, S. Gerland, V. Tverberg et al., Wintertime warming of an Arctic shelf in response to large-scale atmospheric circulation, Geophysical Research Letters, vol.34, 2007.

E. L. Crockett, The cold but not hard fats in ectotherms: consequences of lipid restructuring on susceptibility of biological membranes to peroxidation, a review, Journal of Comparative Physiology B-Biochemical Systemic and Environmental Physiology, vol.178, pp.795-809, 2008.

J. Dalsgaard, M. St-john, G. Kattner, D. Muller-navarra, and W. Hagen, Fatty acid trophic markers in the pelagic marine environment, Advances in Marine Biology, vol.46, pp.225-340, 2003.

C. Dang, X. De-montaudouin, M. Gam, C. Paroissin, N. Bru et al., The Manila clam population in Arcachon Bay (SW France): Can it be kept sustainable, Journal of Sea Research, vol.63, pp.108-118, 2010.
URL : https://hal.archives-ouvertes.fr/hal-00868095

C. B. Davenport, Growth lines in fossil pectens as indicators of past climates, Journal of Paleontology, vol.12, pp.514-515, 1938.

F. De-saussure, Grande Bibliothèque Payot, Paris Dekker R, Beukema JJ (1999) Relations of summer and winter temperatures with dynamics and growth of two bivalves, Tellina tenuis and Abra tenuis, on the northern edge of their intertidal distribution, Journal of Sea Research, vol.42, pp.207-220, 1967.

C. M. Del-rio, N. Wolf, S. A. Carleton, and L. Z. Gannes, Isotopic ecology ten years after a call for more laboratory experiments, Biological Reviews, vol.84, pp.91-111, 2009.

M. J. Deniro and S. Epstein, Influence of diet on the distribution of nitrogen isotopes in animals, Geochimica Et Cosmochimica Acta, vol.45, pp.341-351, 1981.

D. L. Dettman, K. W. Flessa, P. D. Roopnarine, B. R. Schone, and D. H. Goodwin, The use of oxygen isotope variation in shells of estuarine mollusks as a quantitative record of seasonal and annual Colorado River discharge, Geochimica Et Cosmochimica Acta, vol.68, pp.1253-1263, 2004.

J. R. Dodd, Environmental control of strontium and magnesium in Mytilus, Geochimica Et Cosmochimica Acta, vol.29, 1965.

M. J. Dunbar, Arctic and Subarctic Marine Ecology: Immediate Problems Arctic, vol.6, pp.75-90, 1953.

M. J. Dunbar, Ecological Development in Polar Regions. A study in evolution, 1968.

K. H. Dunton, J. L. Goodall, S. V. Schonberg, J. M. Grebmeier, and D. R. Maidment, Multidecadal synthesis of benthic-pelagic coupling in the western arctic: Role of crossshelf advective processes, Deep-Sea Research Part Ii-Topical Studies in Oceanography, vol.52, pp.3462-3477, 2005.

M. Elliot, P. B. Linsley, B. K. Howe, and S. S. , Environmental controls on the stable isotopic composition of Mercenaria mercenaria: Potential application to paleoenvironmental studies, Geochemistry Geophysics Geosystems, vol.4, 2003.

M. Elliot, K. Welsh, C. Chilcott, M. Mcculloch, J. Chappell et al., Profiles of trace elements and stable isotopes derived from giant long-lived Tridacna gigas bivalves: Potential applications in paleoclimate studies, Palaeogeography Palaeoclimatology Palaeoecology, vol.280, pp.132-142, 2009.

S. Epstein, R. Buchsbaum, H. A. Lowenstam, and H. C. Urey, Revised carbonate-water isotopic temperature scale, Geological Society of America Bulletin, vol.64, pp.1315-1325, 1953.

D. Ezgeta-balic, M. Najdek, M. Peharda, and M. Blazina, Seasonal fatty acid profile analysis to trace origin of food sources of four commercially important bivalves, Aquaculture, vol.334, pp.89-100, 2012.

T. Farkas, Adaptation of fatty-acid compositions to temperature-Study on planktonic crustaceans, Comparative Biochemistry and Physiology BBiochemistry & Molecular Biology, vol.64, pp.71-76, 1979.

T. W. Fileman, D. W. Pond, R. G. Barlow, and R. Mantoura, Vertical profiles of pigments, fatty acids and amino acids: Evidence for undegraded diatomaceous material sedimenting to the deep ocean in the Bellingshausen Sea, Antarctica. Deep Sea Research Part I: Oceanographic Research Papers, vol.45, pp.333-346, 1998.

N. S. Fisher, R. Guillard, and D. C. Bankston, The accumulation of Barium by marinephytoplankton grown in culture, Journal of Marine Research, vol.49, pp.339-354, 1991.

O. Folmer, M. Black, W. Hoeh, R. Lutz, and R. Vrijenhoek, DNA primers for amplification of mitochondrial cytochrome c oxidase subunit I from diverse Metazoan invertebrates, Molecular marine biology and biotechnology, vol.3, pp.294-299, 1994.

R. L. France and R. H. Peters, Ecosystem differences in the trophic enrichment of C-13 in aquatic food webs, Canadian Journal of Fisheries and Aquatic Sciences, vol.54, pp.1255-1258, 1997.

S. Fredriksen, Food web studies in a Norwegian kelp forest based on stable isotope (delta C-13 and delta N-15) analysis, Marine Ecology Progress Series, vol.260, pp.71-81, 2003.

P. S. Freitas, L. J. Clarke, H. Kennedy, C. A. Richardson, and F. Abrantes, Environmental and biological controls on elemental (Mg/Ca, Sr/Ca and Mn/Ca) ratios in shells of the king scallop Pecten maximus, Geochimica Et Cosmochimica Acta, vol.70, pp.5119-5133, 2006.

Y. Frenot, Régions polaires : quels enjeux ? Le Pommier, 2006.

B. Gaillard, B. Gaillard, T. Meziane, R. Tremblay, P. Archambault et al., Use of filter-feeding bivalves to monitor pelagic-benthic coupling in Arctic, revision) Trophic resources of the bivalve Astarte elliptica in a subarctic fjord: a multi-biomarkers approach, 2016.

B. Gaillard, T. Meziane, R. Tremblay, P. Archambault, K. Layton et al., Dietary tracers in Bathyarca glacialis from contrasting trophic regions in the Canadian Arctic, Marine Ecology Progress Series, vol.536, pp.175-186, 2015.

J. Gayon, F. Dehairs, A. Lorrain, D. Steenmans, W. Baeyens et al., Barium uptake into the shells of the common mussel (Mytilus edulis) and the potential for estuarine paleo-chemistry reconstruction, Gachelin G (ed) Les organismes modèles dans la recherche médicale. Presses Universitaires de France Gillikin DP, vol.70, pp.395-407, 2006.

D. P. Gillikin, A. Lorrain, Y. M. Paulet, L. Andre, and F. Dehairs, Synchronous barium peaks in high-resolution profiles of calcite and aragonite marine bivalve shells, GeoMarine Letters, vol.28, pp.351-358, 2008.
URL : https://hal.archives-ouvertes.fr/hal-00452787

G. Team, GNU Image Manipulation Program 2.8.10, 2014.

M. I. Gladyshev, N. N. Sushchik, G. S. Kalachova, and O. N. Makhutova, Stable Isotope Composition of Fatty Acids in Organisms of Different Trophic Levels in the Yenisei River, PloS One, vol.7, 2012.

M. I. Gladyshev, N. N. Sushchik, O. N. Makhutova, and G. S. Kalachova, Trophic fractionation of isotope composition of polyunsaturated fatty acids in the trophic chain of a river ecosystem, Doklady Biochemistry and Biophysics, vol.454, pp.4-5, 2014.

R. Glud, S. Rysgaard, M. Kühl, and J. Hansen, (eds) Carbon cycling in Arctic marine ecosystems: Case study Young Sound, Meddelelser om Grønland Bioscience, vol.58, 2007.

R. N. Glud, M. Kuhl, F. Wenzhofer, and S. Rysgaard, ): importance for ecosystem primary production, Marine Ecology Progress Series, vol.238, pp.15-29, 2002.

R. N. Glud, S. Rysgaard, and . Greenland, (eds) Carbon cycling in Arctic marine ecosystems: Case study Young Sound, Meddelelser om Grønland Bioscience, vol.58, 2007.

R. N. Glud, J. Woelfel, U. Karsten, M. Kuhl, and S. Rysgaard, Benthic microalgal production in the Arctic: applied methods and status of the current database, Botanica Marina, vol.52, pp.559-571, 2009.

S. Golubic and J. Schneider, Preliminary evaluation of potential stable isotope and trace element productivity proxies in the oyster Crassostrea gigas, Palaeogeography Palaeoclimatology Palaeoecology, vol.373, pp.88-97, 1979.

D. H. Goodwin, B. R. Schone, and D. L. Dettman, Resolution and fidelity of oxygen isotopes as paleotemperature proxies in bivalve mollusk shells: Models and observations, Palaios, vol.18, pp.110-125, 2003.

E. Gosling, M. Gosselin, M. Levasseur, P. A. Wheeler, R. A. Horner et al., New measurements of phytoplankton and ice algal production in the Arctic Ocean, Deep-Sea Research Part II-Topical Studies in Oceanography, vol.44, p.1623, 1997.

R. R. Gradinger, B. A. Bluhm, R. R. Hopcroft, A. V. Gebruk, K. Kosobokova et al., Marine Life in the Arctic, McIntyre AD (ed) Life in the World's Oceans, 2010.

M. Graeve, G. Kattner, and D. Piepenburg, Lipids in Arctic benthos: Does the fatty acid and alcohol composition reflect feeding and trophic interactions, Polar Biology, vol.18, pp.53-61, 1997.

C. Graham, L. Oxtoby, S. W. Wang, S. M. Budge, and M. J. Wooller, Sourcing fatty acids to juvenile polar cod (Boreogadus saida) in the Beaufort Sea using compound-specific stable carbon isotope analyses, Polar Biology, vol.37, pp.697-705, 2014.

J. M. Grebmeier and J. P. Barry, The influence of oceanographic processes on pelagicbenthic coupling in polar regions: A benthic perspective, Journal of Marine Systems, vol.2, pp.495-518, 1991.

J. M. Grebmeier, J. E. Overland, S. E. Moore, E. V. Farley, E. C. Carmack et al., A major ecosystem shift in the northern Bering Sea, Science, vol.311, pp.1461-1464, 2006.

D. R. Grocke and D. P. Gillikin, Advances in mollusc sclerochronology and sclerochemistry: tools for understanding climate and environment, Geo-Marine Letters, vol.28, pp.265-268, 2008.

E. L. Grossman and T. L. Ku, Oxygen and carbon isotope fractionation in biogenic aragonite-temperature effects, Chem Geol, vol.59, pp.59-74, 1986.

A. A. Gusev and L. V. Rudinskaya, Shell form, growth, and production of Astarte borealis (Schumacher, 1817) (Astartidae, Bivalvia) in the southeastern Baltic Sea, Oceanology, vol.54, pp.458-464, 2014.

E. N. Hegseth, Primary production of the northern Barents Sea, Polar Research, vol.17, pp.113-123, 1998.

E. N. Hegseth and V. Tverberg, Effect of Atlantic water inflow on timing of the phytoplankton spring bloom in a high Arctic fjord (Kongsfjorden, Svalbard), Journal of Marine Systems, vol.113, pp.94-105, 2013.

O. Heilmayer, C. Honnen, U. Jacob, M. Chiantore, R. Cattaneo-vietti et al., Temperature effects on summer growth rates in the Antarctic scallop, Adamussium colbecki. Polar Biology, vol.28, pp.523-527, 2005.

J. P. Hendry, W. T. Perkins, and T. Bane, Short-term environmental change in a Jurassic lagoon deduced from geochemical trends in aragonite bivalve shells, Geological Society of America Bulletin, vol.113, pp.790-798, 2001.

K. A. Hobson, A. Fisk, N. Karnovsky, M. Holst, J. M. Gagnon et al., A stable isotope (delta C-13, delta N-15) model for the North Water food web: implications for evaluating trophodynamics and the flow of energy and contaminants, Deep-Sea Research Part II-Topical Studies in Oceanography, vol.49, pp.5131-5150, 2002.

K. A. Hobson, D. M. Schell, D. Renouf, and E. Noseworthy, Stable carbon and nitrogen isotopic fractionation between diet and tissues of captive seals: Implications for dietary reconstructions involving marine mammals, Canadian Journal of Fisheries and Aquatic Sciences, vol.53, pp.528-533, 1996.

H. Hodal, S. Falk-petersen, H. Hop, S. Kristiansen, and M. Reigstad, Spring bloom dynamics in Kongsfjorden, Svalbard: nutrients, phytoplankton, protozoans and primary production, Polar Biology, vol.35, pp.191-203, 2012.

H. Hop, T. Pearson, E. N. Hegseth, K. M. Kovacs, C. Wiencke et al., The marine ecosystem of Kongsfjorden, vol.21, pp.167-208, 2002.

G. P. Hoskin, Alterations in Lipid Metabolism of Molluscs Due to Dietary Changes, Invertebrate Models for Biochemical Research, vol.184, pp.149-166, 1978.

B. L. Ingram, J. C. Ingle, and M. E. Conrad, Stable isotope record of late Holocene salinity and river discharge in San Francisco Bay, California. Earth and Planetary Science Letters, vol.141, pp.237-247, 1996.

T. F. Stocker, D. Qin, G. K. Plattner, M. Tignor, S. K. Allen et al., Climate Change 2014: Synthesis Report. Contribution of Working Groups I, II and III to the Fifth Assessment Report of the Intergovernmental Panel on Climate Change, The Physical Science Basis. Contribution of Working Group I to the Fifth Assessment Report of the Intergovernmental Panel on Climate Change, 2013.

C. Israelson, B. Buchardt, S. Funder, and H. W. Hubberten, Oxygen and carbon-isotope composition of Quaternary bivalve shells as a water mass indicator: Last interglacial and Holocene, Palaeogeography Palaeoclimatology Palaeoecology, vol.111, pp.119-134, 1994.

L. C. Ivany, B. H. Wilkinson, K. C. Lohmann, E. R. Johnson, B. J. Mcelroy et al., Intraannual isotopic variation in Venericardia bivalves: Implications for early eocene temperature, seasonality, and salinity on the US Gulf Coast, Journal of Sedimentary Research, vol.74, pp.7-19, 2004.

S. Iverson, M. L. Springer-jakobsson, A. Mayer, B. Coakley, J. A. Dowdeswell et al., Tracing Aquatic Food Webs Using Fatty Acids: From Qualitative Indicators to Quantitative Determination, vol.39, p.12609, 2009.

D. S. Jones, Sclerochronology-Reading the record of the Molluscan shell, American Scientist, vol.71, pp.384-391, 1983.

D. S. Jones and W. D. Allmon, Records of upwelling, seasonality and growth in stableisotope profiles of Pliocene Mollusk shells from Florida, Lethaia, vol.28, pp.61-74, 1995.

U. Karsten, C. Schlie, J. Wolfel, and B. Becker, Benthic diatoms in Arctic Seas-Ecological Functions and Adaptations, Polarforschung, vol.81, pp.77-84, 2012.

M. Kedra, C. Moritz, E. S. Choy, C. David, R. Degen et al., Status and trends in the structure of Arctic benthic food webs, vol.34, 2015.
URL : https://hal.archives-ouvertes.fr/hal-01161465

J. R. Kelly and R. E. Scheibling, Fatty acids as dietary tracers in benthic food webs, Marine Ecology Progress Series, vol.446, pp.1-22, 2012.

B. K. Khim, D. E. Krantz, and J. Brigham-grette, Stable isotope profiles of Last Interglacial (Pelukian Transgression) mollusks and paleoclimate implications in the Bering Strait Region, Quaternary Science Reviews, vol.20, pp.463-483, 2001.

B. K. Khim, D. E. Krantz, L. W. Cooper, and J. M. Grebmeier, Seasonal discharge of estuarine freshwater to the western Chukchi Sea shelf identified in stable isotope profiles of mollusk shells, Journal of Geophysical Research-Oceans, vol.108, 2003.

J. S. Kim, K. S. Woo, and W. Hong, High resolution geochemical investigation of the bivalve shells (Glycymeris sp.) from shell mounds in Jeju Island, Korea: Late Holocene paleoclimatic implications related to East Asian Monsoon climate, Quaternary International, vol.392, pp.3-12, 2016.

M. Klages, A. Boetius, J. P. Christensen, H. Deubel, D. Piepenburg et al., The Benthos of Arctic Seas and its Role for the Organic Carbon Cycle at the Seafloor, The Organic Carbon Cycle in the Arctic Ocean, 2004.

R. T. Klein, K. C. Lohmann, and C. W. Thayer, Bivalve skeletons record sea-surface temperature and delta O-18 via Mg/Ca and O-18/O-16 ratios, Geology, vol.24, pp.415-418, 1996.

J. S. Klingensmith, Distribution of methylene and nonmethylene-interrupted dienoic fatty-acids in polar lipids and triacylglycerols of selected tissues of the hardshell clam (Mercenaria mercenaria), Lipids, vol.17, pp.976-981, 1982.

M. H. Klunder, D. Hippler, R. Witbaard, and D. Frei, Laser ablation analysis of bivalve shells-archives of environmental information, Geol Surv Den Greenl Bull, pp.89-92, 2008.

D. Krause-jensen and C. M. Duarte, Expansion of vegetated coastal ecosystems in the future Arctic, Frontiers in Marine Science, vol.1, 2014.

D. Krause-jensen, M. Kühl, P. Christensen, and J. Borum, (eds) Carbon cycling in Arctic marine ecosystems: Case study Young Sound, Meddelelser om Grønland Bioscience, vol.58, 2007.

D. Krause-jensen, N. Marba, B. Olesen, M. K. Sejr, P. B. Christensen et al., Seasonal sea ice cover as principal driver of spatial and temporal variation in depth extension and annual production of kelp in Greenland, Global Change Biology, vol.18, pp.2981-2994, 2012.

D. Krawczyk, K. Arendt, T. Juul-pedersen, M. Sejr, M. Blicher et al., Spatial and temporal distribution of planctonic protists in the East Greenland fjord and offshore waters, Marine Ecology Progress Series, vol.538, pp.99-116, 2015.

C. Lalande, B. Moriceau, A. Leynaert, and N. Morata, Spatial and temporal variability in export fluxes of biogenic matter in Kongsfjorden, Polar Biology, pp.1-14, 2016.
URL : https://hal.archives-ouvertes.fr/hal-01916502

C. E. Lazareth, La coquille des mollusques : mémoire de l'environnement, 2006.

C. E. Lazareth, G. Lasne, and L. Ortlieb, Growth anomalies in Protothaca thaca (Mollusca, Veneridae) shells as markers of ENSO conditions, Climate Research, vol.30, pp.263-269, 2006.
URL : https://hal.archives-ouvertes.fr/ird-01697388

C. E. Lazareth, E. Vander-putten, L. Andre, and F. Dehairs, High-resolution trace element profiles in shells of the mangrove bivalve Isognomon ephippium: a record of environmental spatio-temporal variations?, Estuarine Coastal and Shelf Science, vol.57, pp.1103-1114, 2003.

C. E. Lee, Global phylogeography of a cryptic copepod species complex and reproductive isolation between genetically proximate "populations, Evolution, vol.54, pp.2014-2027, 2000.

L. Legendre, S. F. Ackley, G. S. Dieckmann, B. Gulliksen, R. Horner et al., Ecology of sea ice biota .2. Global significance, Polar Biology, vol.12, pp.429-444, 1992.

J. Legezynska, M. Kedra, and W. Walkusz, Identifying trophic relationships within the high Arctic benthic community: how much can fatty acids tell?, Marine Biology, vol.161, pp.821-836, 2014.

C. Lehane and J. Davenport, Ingestion of mesozooplankton by three species of bivalve, Mytilus edulis, Cerastoderma edule and Aequipecten opercularis, Journal of the Marine Biological Association of the United Kingdom, vol.82, pp.615-619, 2002.

C. Lehane and J. Davenport, Ingestion of bivalve larvae by Mytilus edulis: experimental and field demonstrations of larviphagy in farmed blue mussels, Marine Biology, vol.145, pp.101-107, 2004.

F. Leroy, T. Meziane, P. Riera, and T. Comtet, Seasonal Variations in Maternal Provisioning of Crepidula fornicata (Gastropoda): Fatty Acid Composition of Females, Embryos and Larvae, PloS one, vol.8, 2013.
URL : https://hal.archives-ouvertes.fr/hal-01218825

E. Leu, J. E. Soreide, D. O. Hessen, S. Falk-petersen, and J. Berge, Consequences of changing sea-ice cover for primary and secondary producers in the European Arctic shelf seas: Timing, quantity, and quality, Progress in Oceanography, vol.90, pp.18-32, 2011.

R. W. Lewis, Temperature and pressure effects on fatty acids of some marine ectotherms, Comparative Biochemistry and Physiology, vol.6, 1962.

A. Lorrain, Y. M. Paulet, L. Chauvaud, N. Savoye, A. Donval et al., Differential delta C-13 and delta N-15 signatures among scallop tissues: implications for ecology and physiology, Journal of Experimental Marine Biology and Ecology, vol.275, pp.47-61, 2002.

A. Lorrain, N. Savoye, L. Chauvaud, Y. M. Paulet, and N. Naulet, Decarbonation and preservation method for the analysis of organic C and N contents and stable isotope ratios of low-carbonated suspended particulate material, Analytica Chimica Acta, vol.491, pp.125-133, 2003.
URL : https://hal.archives-ouvertes.fr/hal-00452450

M. Che, L. , L. Campion-alsumard, T. Boury-esnault, N. Payri et al., Biodegradation of shells of the black pearl oyster, Pinctada margaritifera var cumingii, by microborers and sponges of French Polynesia, Marine Biology, vol.126, pp.509-519, 1996.

M. C. Marcano, T. D. Frank, S. B. Mukasa, K. C. Lohmann, and M. Taviani, Diagenetic incorporation of Sr into aragonitic bivalve shells: implications for chronostratigraphic and palaeoenvironmental interpretations, Depositional Record, vol.1, pp.38-52, 2015.

F. Marin and G. Luquet, Molluscan shell proteins, Comptes Rendus Palevol, vol.3, pp.469-492, 2004.
URL : https://hal.archives-ouvertes.fr/hal-00197133

L. Marincovich, K. B. Barinov, and A. E. Oleinik, The Astarte (Bivalvia : Astartidae) that document the earliest opening of Bering Strait, Journal of Paleontology, vol.76, pp.239-245, 2002.

R. Marsh, B. Petrie, C. R. Weidman, R. R. Dickson, J. W. Loder et al., The 1882 tilefish kill-a cold event in shelf waters off the north-eastern United States?, Fisheries Oceanography, vol.8, pp.39-49, 1999.

. Masson-delmotte-v-;-le-vrai, . Le-faux.-le, P. Pommier, J. H. Mccutchan, W. M. Lewis et al., Variation in trophic shift for stable isotope ratios of carbon, nitrogen, and sulfur, Oikos, vol.102, pp.378-390, 2003.

K. W. Mcmahon, W. G. Ambrose, B. J. Johnson, M. Sun, G. R. Lopez et al., Benthic community response to ice algae and phytoplankton in Ny Ålesund, Marine Ecology Progress Series, vol.310, pp.1-14, 2006.

B. C. Mcmeans, N. Rooney, M. T. Arts, and A. T. Fisk, Food web structure of a coastal Arctic marine ecosystem and implications for stability, Marine Ecology Progress Series, vol.482, 2013.

N. D. Mctigue, P. Bucolo, Z. F. Liu, and K. H. Dunton, Pelagic-benthic coupling, food webs, and organic matter degradation in the Chukchi Sea: Insights from sedimentary pigments and stable carbon isotopes, Limnology and Oceanography, vol.60, pp.429-445, 2015.

T. Meziane and M. Tsuchiya, Organic matter in a subtropical mangrove-estuary subjected to wastewater discharge: Origin and utilisation by two macrozoobenthic species, Journal of Sea Research, vol.47, pp.1-11, 2002.

O. Monroig, D. R. Tocher, and J. C. Navarro, Biosynthesis of polyunsaturated fatty acids in marine invertebrates: recent advances in molecular mechanisms, Marine drugs, vol.11, pp.3998-4018, 2013.

J. M. Mortillaro, G. Abril, P. Moreira-turcq, R. L. Sobrinho, M. Perez et al., Fatty acid and stable isotope (delta C-13, delta N-15) signatures of particulate organic matter in the lower Amazon River: Seasonal contrasts and connectivity between floodplain lakes and the mainstem, Organic Geochemistry, vol.42, pp.1159-1168, 2011.

T. Mueller-lupp, H. Erlenkeuser, and H. A. Bauch, Seasonal and interannual variability of Siberian river discharge in the Laptev Sea inferred from stable isotopes in modern bivalves, Boreas, vol.32, pp.292-303, 2003.

T. Muller-lupp and H. Bauch, Linkage of Arctic atmospheric circulation and Siberian shelf hydrography: A proxy validation using delta(18)O records of bivalve shells, Glob Planet Change, vol.48, pp.175-186, 2005.

C. J. Mundy, M. Gosselin, Y. Gratton, K. Brown, V. Galindo et al., Role of environmental factors on phytoplankton bloom initiation under landfast sea ice in Resolute Passage, Canada. Marine Ecology Progress Series, vol.497, 2014.

S. A. Murzina, Z. A. Nefedova, S. Falk-petersen, H. Hop, T. R. Ryokolainen et al., Lipids in the daubed shanny (Teleostei: Leptoclinus maculatus) in Svalbard waters, Polar Biology, vol.36, pp.1619-1631, 2013.

M. Najdek, D. Ezgeta-balic, M. Blazina, M. Crncevic, and M. Peharda, Potential food sources of Glycymeris nummaria (Mollusca: Bivalvia) during the annual cycle indicated by fatty acid analysis, Scientia Marina, vol.80, pp.123-129, 2016.

G. E. Napolitano and R. G. Ackman, Anatomical distributions and temporal variations of lipid classes in sea scallops Placopecten magellanicus (Gmelin) from Georges Bank, Comparative Biochemistry and Physiology B-Biochemistry & Molecular Biology, vol.103, pp.645-650, 1992.

N. , Nuuk Ecological Research Operations 7th Annual Report, 2013.

C. Nerot, A. Lorrain, J. Grall, D. P. Gillikin, J. M. Munaron et al., Stable isotope variations in benthic filter feeders across a large depth gradient on the continental shelf, Estuarine Coastal and Shelf Science, vol.96, pp.228-235, 2012.
DOI : 10.1016/j.ecss.2011.11.004

URL : https://hal.archives-ouvertes.fr/hal-00660783

C. Nerot, T. Meziane, G. Schaal, J. Grall, A. Lorrain et al., Spatial changes in fatty acids signatures of the great scallop Pecten maximus across the Bay of Biscay continental shelf, Continental Shelf Research, vol.109, pp.1-9, 2015.

W. Oschmann, Sclerochronology: editorial. International Journal of Earth Sciences, vol.98, pp.1-2, 2009.

L. E. Oxtoby, J. T. Mathis, L. W. Juranek, and M. J. Wooller, Estimating stable carbon isotope values of microphytobenthos in the Arctic for application to food web studies, Polar Biology, vol.39, pp.473-483, 2015.

M. Paar, A. Voronkov, H. Hop, T. Brey, I. Bartsch et al., Temporal shift in biomass and production of macrozoobenthos in the macroalgal belt at Hansneset, Kongsfjorden, after 15 years, Polar Biology Parent GJ, vol.3, pp.101-109, 2008.
URL : https://hal.archives-ouvertes.fr/hal-01243387

A. C. Parnell, R. Inger, S. Bearhop, and A. L. Jackson, Source Partitioning Using Stable Isotopes: Coping with Too Much Variation, PLoS ONE, vol.5, issue.3, p.9672, 2010.

Y. Paulet, A. Lorrain, J. Richard, and S. Pouvreau, Experimental shift in diet ?13C: A potential tool for ecophysiological studies in marine bivalves, Organic Geochemistry, vol.37, pp.1359-1370, 2006.
URL : https://hal.archives-ouvertes.fr/hal-00452408

A. J. Pazos, J. L. Sanchez, G. Roman, M. L. Perez-paralle, and M. Abad, Seasonal changes in lipid classes and fatty acid composition in the digestive gland of Pecten maximus, Comparative Biochemistry and Physiology B-Biochemistry & Molecular Biology, vol.134, pp.367-380, 2003.

M. Peharda, D. Ezgeta-balic, J. Davenport, N. Bojanic, O. Vidjak et al., Differential ingestion of zooplankton by four species of bivalves (Mollusca) in the Mali, Marine Biology, vol.159, pp.881-895, 2012.

V. Perez, F. Olivier, R. Tremblay, U. Neumeier, J. Thebault et al., Trophic resources of the bivalve, Venus verrucosa, the Chausey archipelago, vol.26, pp.229-239, 2013.
URL : https://hal.archives-ouvertes.fr/hal-00946527

F. Pernet, R. Tremblay, C. Gionet, and T. Landry, Lipid remodeling in wild and selectively bred hard clams at low temperatures in relation to genetic and physiological parameters, Journal of Experimental Biology, vol.209, pp.4663-4675, 2006.

G. H. Petersen, Studies on some Arctic and Baltic Astarte species (Bivalvia, Mollusca), Bioscience, vol.52, 2000.

B. Peterson and B. Fry, Stable Isotopes in Ecosystem Studies, Annual Review of Ecology and Systematics, vol.18, pp.293-320, 1987.

B. J. Peterson, R. M. Holmes, J. W. Mcclelland, C. J. Vorosmarty, R. B. Lammers et al., Increasing river discharge to the Arctic Ocean, Science, vol.298, pp.2171-2173, 2002.

C. H. Peterson, Quantitative allometry of gamete production by Mercenaria mercenaria into old-age, Marine Ecology Progress Series, vol.29, pp.93-97, 1986.
DOI : 10.3354/meps029093

URL : http://doi.org/10.3354/meps029093

A. K. Pettersen, G. M. Turchini, S. Jahangard, B. A. Ingram, and C. Sherman, Effects of different dietary microalgae on survival, growth, settlement and fatty acid composition of blue mussel (Mytilus galloprovincialis) larvae, Aquaculture, vol.309, pp.115-124, 2010.

D. Piepenburg, Recent research on Arctic benthos: common notions need to be revised, Polar Biology, vol.28, pp.733-755, 2005.
DOI : 10.1007/s00300-005-0013-5

D. Piepenburg, P. Archambault, W. Ambrose, A. Blanchard, B. Bluhm et al., Towards a pan-Arctic inventory of the species diversity of the macro-and megabenthic fauna of the Arctic shelf seas, Marine Biodiversity, vol.41, pp.51-70, 2011.

S. E. Portilla, B. F. Branco, and J. T. Tanacredi, and 22:6n-3, on mortality of juvenile Mercenaria mercenaria during the approach to winter, Aquaculture International, vol.20, pp.1357-1376, 2015.

D. M. Post, Using stable isotopes to estimate trophic position: Models, methods, and assumptions, Ecology, vol.83, pp.703-718, 2002.
DOI : 10.2307/3071875

URL : http://onlinelibrary.wiley.com/doi/10.1890/0012-9658(2002)083[0703:USITET]2.0.CO;2/pdf

E. Prato, A. Danieli, M. Maffia, and F. Biandolino, Lipid and fatty acid compositions of Mytilus galloprovincialis cultured in the Mar Grande of Taranto (Southern Italy): feeding strategies and trophic relationships, Zoological Studies, vol.49, pp.211-219, 2010.

A. Y. Proshutinsky and M. A. Johnson, Two circulation regimes of the wind driven Arctic Ocean, Journal of Geophysical Research-Oceans, vol.102, pp.12493-12514, 1997.
DOI : 10.1029/97jc00738

URL : http://onlinelibrary.wiley.com/doi/10.1029/97JC00738/pdf

. R-core-team, R: A language and environment for statistical computing. R Foundation for Statistical Computing, 2014.

P. Renaud, T. Løkken, L. Jørgensen, J. Berge, and B. Johnson, Macroalgal detritus and food-web subsidies along an Arctic fjord depth-gradient, Frontiers in Marine Science, vol.2, pp.1-15, 2015.
DOI : 10.3389/fmars.2015.00031

URL : https://www.frontiersin.org/articles/10.3389/fmars.2015.00031/pdf

P. E. Renaud, M. K. Sejr, B. A. Bluhm, B. Sirenko, and I. H. Ellingsen, The future of Arctic benthos: Expansion, invasion, and biodiversity, Progress in Oceanography, vol.139, pp.244-257, 2015.

P. E. Renaud, M. Tessmann, A. Evenset, and G. N. Christensen, Benthic food-web structure of an Arctic fjord, Marine Biology Research, vol.7, pp.13-26, 2011.

N. Reuss and L. K. Poulsen, Evaluation of fatty acids as biomarkers for a natural plankton community. A field study of a spring bloom and a post-bloom period off West Greenland, Marine Biology, vol.141, pp.423-434, 2002.

D. Rhoads and R. Lutz, Molluscs as archives of environmental change, Oceanography and Marine Biology: an Annual Review, vol.39, pp.103-164, 1980.

C. A. Richardson, D. J. Crisp, and N. W. Runham, Tidally deposited growth bands in the shell of the common cockle, Cerastoderma edule, Malacologia, vol.18, pp.277-290, 1979.

C. A. Richardson, D. J. Crisp, and N. W. Runham, Factors influencing shell deposition during a tidal cycle in the inter-tidal bivalve Cerastoderma edule, Journal of the Marine Biological Association of the United Kingdom, vol.61, pp.465-476, 1981.

K. Richardson and T. Cedhagen, Quantifying Pelagic-Benthic coupling in the North Sea: Are we asking the right questions, Burning Issues of North Sea Ecology, p.31, 2001.

E. A. Robinson, G. A. Blagoev, P. Hebert, and S. J. Adamowicz, Prospects for using DNA barcoding to identify spiders in species-rich genera, Zookeys, vol.16, pp.27-46, 2009.

C. Royer, J. Thebault, L. Chauvaud, and F. Olivier, , 2013.

N. Brittany and . France, Palaeogeography Palaeoclimatology Palaeoecology, vol.373, pp.123-132

S. Rysgaard and M. K. Sejr, Vertical flux of particulate organic matter in a High Arctic fjord: Relative importance of terrestrial and marine sources, Meddelelser om Grønland Bioscience, vol.58, 2007.

S. Rysgaard and R. N. Glud, Carbon cycling and climate change: Predictions for a High Arctic marine ecosystem (Young Sound, NE Greenland). Carbon cycling in Arctic marine ecosystems: Case study Young Sound, Meddelelser om Grønland Bioscience, vol.58, 2007.

, Carbon cycling in Arctic marine ecosystems: Case study Young Sound, Meddelelser om Grønland Bioscience, vol.58, 2007.

S. Rysgaard, T. G. Nielsen, and B. W. Hansen, Seasonal variation in nutrients, pelagic primary production and grazing in a high-Arctic coastal marine ecosystem, Marine Ecology Progress Series, vol.179, pp.13-25, 1999.

E. Sakshaug, Primary and Secondary Production in the Arctic Seas. The Organic Carbon Cycle in the Arctic Ocean, 2004.

A. Saleuddin, The gonads and reproductive cycle of Astarte sulcata (Da Costa) and sexuality in A. elliptica (Brown), Proceedings of the Malacological Society of London, vol.86, pp.141-147, 1964.

A. Saleuddin, The mode of life and functional anatomy of Astarte spp. (Eulamellibranchia), Journal of Molluscan Studies, vol.36, pp.229-257, 1965.

G. Schomburg, Gas chromatographie Grundlagen-Praxis-Kapillartechnik 2nd ed, 1987.

B. R. Schone, Arctica islandica (Bivalvia): A unique paleoenvironmental archive of the northern North Atlantic Ocean, Glob Planet Change, vol.111, pp.199-225, 2013.

B. R. Schone, A. Castro, J. Fiebig, S. D. Houk, W. Oschmann et al., Sea surface water temperatures over the period 1884-1983 reconstructed from oxygen isotope ratios of a bivalve mollusk shell (Arctica islandica, 2004.

, Palaeogeography Palaeoclimatology Palaeoecology, vol.212, pp.215-232

B. R. Schone, J. Fiebig, M. Pfeiffer, R. Gless, J. Hickson et al., Climate records from a bivalved Methuselah (Arctica islandica, Mollusca; Iceland), Palaeogeography Palaeoclimatology Palaeoecology, vol.228, pp.130-148, 2005.

B. R. Schone and D. P. Gillikin, Unraveling environmental histories from skeletal diaries-Advances in sclerochronology, Palaeogeography Palaeoclimatology Palaeoecology, vol.373, pp.1-5, 2013.

B. R. Schone, J. Lega, K. W. Flessa, D. H. Goodwin, and D. L. Dettman, Reconstructing daily temperatures from growth rates of the intertidal bivalve mollusk Chione cortezi (northern Gulf of California, Mexico), Palaeogeography Palaeoclimatology Palaeoecology, vol.184, pp.131-146, 2002.

B. R. Schone, W. Oschmann, J. Rossler, A. Castro, S. D. Houk et al., North Atlantic Oscillation dynamics recorded in shells of a long-lived bivalve mollusk, Geology, vol.31, pp.1037-1040, 2003.

B. R. Schone, Z. J. Zhang, P. Radermacher, J. Thebault, D. E. Jacob et al., Sr/Ca and Mg/Ca ratios of ontogenetically old, long-lived bivalve shells (Arctica islandica) and their function as paleotemperature proxies, Palaeogeography Palaeoclimatology Palaeoecology, vol.302, pp.52-64, 2011.
URL : https://hal.archives-ouvertes.fr/hal-00675854

J. D. Scourse, A. D. Wanamaker, C. Weidman, J. Heinemeier, P. J. Reimer et al., The marine radiocarbon bomb pulse across the temperate North Atlantic: a compilation of delta c-14 time histories from Arctica Islandica growth increments, Radiocarbon, vol.54, pp.165-186, 2012.

M. K. Sejr, M. E. Blicher, and S. Rysgaard, Sea ice cover affects inter-annual and geographic variation in growth of the Arctic cockle Clinocardium ciliatum (Bivalvia) in Greenland, Marine Ecology Progress Series, vol.389, pp.149-158, 2009.

M. K. Sejr, K. T. Jensen, and S. Rysgaard, Annual growth bands in the bivalve Hiatella arctica validated by a mark-recapture study in NE Greenland, Polar Biology, vol.25, pp.794-796, 2002.

N. I. Selin, Shell form, growth and life span of Astarte arctica and A. borealis (Mollusca : Bivalvia) from the subtidal zone of northeastern Sakhalin, Russian Journal of Marine Biology, vol.33, pp.232-237, 2007.

O. A. Sherwood and E. N. Edinger, Ages and growth rates of some deep-sea gorgonian and antipatharian corals of Newfoundland and Labrador, Canadian Journal of Fisheries and Aquatic Sciences, vol.66, pp.142-152, 2009.

J. Simstich, I. Harms, M. J. Karcher, H. Erlenkeuser, V. Stanovoy et al., Recent freshening in the Kara Sea (Siberia) recorded by stable isotopes in Arctic bivalve shells, Journal of Geophysical Research-Oceans, vol.110, 2005.

J. E. Soreide, M. L. Carroll, H. Hop, W. G. Ambrose, E. N. Hegseth et al., Sympagic-pelagic-benthic coupling in Arctic and Atlantic waters around Svalbard revealed by stable isotopic and fatty acid tracers, Marine Biology Research, vol.9, pp.831-850, 2013.

J. E. Soreide, H. Hop, M. L. Carroll, S. Falk-petersen, and E. N. Hegseth, Seasonal food web structures and sympagic-pelagic coupling in the European Arctic revealed by stable isotopes and a two-source food web model, Progress in Oceanography, vol.71, pp.59-87, 2006.

J. E. Soreide, E. Leu, J. Berge, M. Graeve, and S. Falk-petersen, Timing of blooms, algal food quality and Calanus glacialis reproduction and growth in a changing Arctic, Global Change Biology, vol.16, pp.3154-3163, 2010.

R. Spencer, G. R. Aiken, K. D. Butler, M. M. Dornblaser, R. G. Striegl et al., Utilizing chromophoric dissolved organic matter measurements to derive export and reactivity of dissolved organic carbon exported to the Arctic Ocean: A case study of the Yukon River, Alaska. Geophysical Research Letters, vol.36, 2009.

R. A. Stead, N. B. Richoux, S. V. Pereda, and R. J. Thompson, Influence of an intermittent food supply on energy storage by the subpolar deposit feeder Yoldia hyperborea (Bivalvia: Nuculanidae), Polar Biology, vol.36, pp.1333-1345, 2013.

H. A. Stecher, D. E. Krantz, C. J. Lord, G. W. Luther, and K. W. Bock, Profiles of strontium and barium in Mercenaria mercenaria and Spisula solidissima shells, Geochimica Et Cosmochimica Acta, vol.60, pp.3445-3456, 1996.

H. Svendsen, A. Beszczynska-moller, J. O. Hagen, B. Lefauconnier, V. Tverberg et al., The physical environment of Kongsfjorden-Krossfjorden, an Arctic fjord system in Svalbard, Polar Research, vol.21, pp.133-166, 2002.

R. K. Takesue and A. Van-geen, Mg/Ca, Sr/Ca, and stable isotopes in modern and holocene Protothaca staminea shells from a northern California coastal upwelling region, Geochimica Et Cosmochimica Acta, vol.68, pp.3845-3861, 2004.

K. Tamura, G. Stecher, D. Peterson, A. Filipski, and S. Kumar, MEGA6: Molecular Evolutionary Genetics Analysis version 6.0, Molecular Biology and Evolution, vol.30, pp.2725-2729, 2013.

A. G. Tansley, The use and abuse of vegetational concepts and terms, Ecology, vol.16, pp.284-307, 1935.

J. Thebault and L. Chauvaud, Li/Ca enrichments in great scallop shells (Pecten maximus) and their relationship with phytoplankton blooms, Palaeogeography Palaeoclimatology Palaeoecology, vol.373, pp.108-122, 2013.
URL : https://hal.archives-ouvertes.fr/hal-00670785

J. Thebault, L. Chauvaud, J. Clavier, R. Fichez, and E. Morize, Evidence of a 2-day periodicity of striae formation in the tropical scallop Comptopallium radula using calcein marking, Marine Biology, vol.149, pp.257-267, 2006.
URL : https://hal.archives-ouvertes.fr/hal-00449282

J. Thébault, L. Chauvaud, L. 'helguen, S. Clavier, J. Barats et al., Barium and molybdenum records in bivalve shells: geochemical proxies for phytoplankton dynamics in coastal environments?, Limnology and Oceanography, vol.54, pp.1002-1014, 2009.

D. Thomas, G. Fogg, P. Convey, C. Fritsen, J. Gili et al., The Biology of Polar Regions, Biogeochemistry of Sea Ice, 2008.
DOI : 10.1093/acprof:oso/9780199298112.001.0001

G. Thorson, The larval development, growth and metabolism of Arctic marine bottom invertebrates, p.100, 1936.

L. L. Tieszen, T. W. Boutton, K. G. Tesdahl, and N. A. Slade, Fractionation and turnover of stable carbon isotopes in animal tissues. Implications for delta c13 analysis of diet, Oecologia, vol.57, pp.32-37, 1983.

H. Toland, B. Perkins, N. Pearce, F. Keenan, and M. J. Leng, A study of sclerochronology by laser ablation ICP-MS, Journal of Analytical Atomic Spectrometry, vol.15, pp.1143-1148, 2000.

M. E. Torres, D. Zima, K. K. Falkner, R. W. Macdonald, M. O'brien et al., Hydrographic Changes in Nares Strait (Canadian Arctic Archipelago) in Recent Decades Based on delta O-18 Profiles of Bivalve Shells, Arctic, vol.64, pp.45-58, 2011.

K. Trutschler and C. Samtleben, Shell growth of Astarte elliptica (Bivalvia) from Kiel bay (Western Baltic sea), Marine Ecology Progress Series, vol.42, pp.155-162, 1988.

H. C. Urey, The thermodynamic properties of isotopic substances, Journal of the Chemical Society, pp.562-581, 1947.

M. Vihtakari, Bivalves as indicators of environmental perturbations related to climate and ocean acidification. Faculty of Biosciences, Fisheries and Economics, 2014.

R. J. Veefkind, Carbon isotope ratios and composition of fatty acids: tags and trophic markers of pelagic organisms, vol.291, 2003.

J. Von-oertzen, Cycles and rates of reproduction of six Baltic Sea bivalves of different zoogeographical origin, Marine Biology, vol.14, pp.143-149, 1972.

A. D. Wanamaker, S. Hetzinger, and J. Halfar, Reconstructing mid-to high-latitude marine climate and ocean variability using bivalves, coralline algae, and marine sediment cores from the Northern Hemisphere, Palaeogeography Palaeoclimatology Palaeoecology, vol.302, pp.1-9, 2011.

S. W. Wang, S. M. Budge, R. R. Gradinger, K. Iken, and M. J. Wooller, Fatty acid and stable isotope characteristics of sea ice and pelagic particulate organic matter in the Bering Sea: tools for estimating sea ice algal contribution to Arctic food web production, Oecologia, vol.174, pp.699-712, 2014.

S. W. Wang, S. M. Budge, K. Iken, R. R. Gradinger, A. M. Springer et al., Importance of sympagic production to Bering Sea zooplankton as revealed from fatty acid-carbon stable isotope analyses, Marine Ecology Progress Series, vol.518, pp.31-50, 2015.

S. W. Wang, A. M. Springer, S. M. Budge, L. Horstmann, L. T. Quakenbush et al., Carbon sources and trophic relationships of ice seals during recent environmental shifts in the Bering Sea, Ecological Applications, vol.26, pp.830-845, 2016.

P. Wassmann, Retention versus export food chains: processes controlling sinking loss from marine pelagic systems, Hydrobiologia, vol.363, pp.29-57, 1998.

P. Wassmann, I. Andreassen, M. Reigstad, and D. Slagstad, Pelagic-benthic coupling in the Nordic Seas: The role of episodic events, Marine Ecology-Pubblicazioni Della Stazione Zoologica Di Napoli I, vol.17, pp.447-471, 1996.

P. Wassmann, C. M. Duarte, S. Agusti, and M. K. Sejr, Footprints of climate change in the Arctic marine ecosystem, Global Change Biology, vol.17, pp.1235-1249, 2011.

P. Wassmann and M. Reigstad, Future Arctic Ocean Seasonal Ice Zones and Implications for Pelagic-Benthic Coupling, Oceanography, vol.24, pp.220-231, 2011.

A. Weber, R. Witbaard, and S. Van-steenpaal, Patterns of growth and undetectable growth lines of Astarte sulcata (Bivalvia) in the Faroe-Shetland Channel, Burning Issues of North Sea Ecology, p.31, 2001.

J. Weems, K. Iken, R. Gradinger, and M. J. Wooller, Carbon and nitrogen assimilation in the Bering Sea clams Nuculana radiata and Macoma moesta, Journal of Experimental Marine Biology and Ecology, vol.430, pp.32-42, 2012.

G. Wefer, W. H. Berger, J. Buma, and G. Fischer, A shell-derived time history of bomb c-14 on Georges Bank and its Labrador Sea implications, Journal of Geophysical Research-Oceans, vol.98, pp.14577-14588, 1993.

J. M. Weslawski and J. J. Wiktor, Increase in biodiversity in the arctic rocky littoral, Sorkappland, Svalbard, after 20 years of climate warming, Marine Biodiversity, vol.40, pp.123-130, 2010.

F. J. Wessels and D. A. Hahn, Carbon 13 discrimination during lipid biosynthesis varies with dietary concentration of stable isotopes: implications for stable isotope analyses, Functional Ecology, vol.24, pp.1017-1022, 2010.

H. Wessels, W. Hagen, C. Wiencke, and U. Karsten, Trophic interactions between macroalgae and herbivores from Kongsfjorden (Svalbard) Dans : Wiencke C (ed) The coastal ecosystem of Kongsfjorden, Svalbard Synopsis of biological research performed at the Koldewey Station in the years, Ber. Polarforsch. Meeresforsch, p.492, 1991.

R. Witbaard, Tree of the sea. The use of the internal growth lines in the shell of Arctica islandica (Bivalvia, Mollusca) for the retrospective assessment of marine environmental change. Rijksuniversiteit Groningen Witbaard R, Duineveld GCA, de Wilde P (1999) Geographical differences in growth rates of Arctica islandica (Mollusca : Bivalvia) from the North Sea and adjacent waters, Journal of the Marine Biological Association of the United Kingdom, vol.79, pp.907-915, 1997.

J. Woelfel, R. Schumann, F. Peine, A. Flohr, A. Kruss et al., ): biomass and potential primary production along the shore line, Microphytobenthos of Arctic Kongsfjorden, vol.33, pp.1239-1253, 2010.

Q. Xu and H. S. Yang, Food sources of three bivalves living in two habitats of Jiaozhou Bay, Journal of Shellfish Research, vol.26, pp.561-567, 2007.

. , Zackenberg Ecological Research Operations 19th Annual Report, 2013.

M. L. Zettler, Ecological and morphological features of the bivalve Astarte borealis (Schumacher, 1817) in the Baltic Sea near its geographical range, Journal of Shellfish Research, vol.21, pp.33-40, 2002.

N. V. Zhukova, The pathway of the biosynthesis of non-methylene-interrupted dienoic fatty-acids in mollusks, Comparative Biochemistry and Physiology BBiochemistry & Molecular Biology, vol.100, pp.801-804, 1991.