Abstract : The volcanic system of La Reunion is made of the coalescent Piton des Neiges and Piton de la Fournaise edifices. Its formation, associated with the activity of a mantle plume, is characterized by phases of submarine and subaerial growth, followed by a period of destruction, as described in the classical model proposed for Hawaiian Island volcanoes. Previous studies showed that the subaerial activity of La Reunion would have started around 2.2 Ma ago. Geochemical analyses carried out on the products associated with subaerial and submarine volcanism revealed their transitional chemical nature and their peculiar homogeneous isotopic compositions (87Sr/86Sr, 143Nd/144Nd et 176Hf/177Hf) for a hotspot derived magmatism. In this study, we show that some unusual submarine lavas were recently identified below the basal series of Piton de la Fournaise volcano (dredged on the North-East Rift Zone of la Fournaise) and below the subaerial lavas outcropping at the base of the differentiated series of Rivière des Remparts, indicate that the evolution of La Reunion system is probably more complex than previously suggested. The question of the nature and origin of these lavas, together with their implication on the formation of La Reunion, is an issue of major interest in order to better constrain the global evolution of the volcanic system. As a consequence, new petrological, geochemical and geochronological investigations were conducted on the submarine part of La Reunion edifice. They revealed the existence of a geochemically exceptional group of lavas. Its composition is clearly different from common samples of La Reunion and presents (1) an enrichment in incompatible elements and (2) tends toward an enriched EM endmember. These chemical specificities reveal that the source of La Reunion magmatic products is (1) an heterogeneous source affected by (2) variable melting degrees. Moreover, two samples of this new group dated at 3.77 (0,08) and 3.34 (0,07) Ma extend considerably the period of activity of the island. These rocks are the oldest samples ever dated at La Reunion. The nature and stratigraphical location of subaerial and submarine samples from Piton des Neiges and Piton de la Fournaise imply a new interpretation of the global evolution of the island. Unlike previous studies, our results indicate that :(1) The differentiated subaerial lavas from Rivière des Remparts, due to their stratigraphical and geographical location underlying Piton de la Fournaise, and by analogy with the Piton des Neiges, cannot be linked to the activity of Piton de la Fournaise. (2) The isotope signature of the submarine lava group from the North-East Rift Zone of Piton de la Fournaise, reveals the heterogeneous character of the mantle source under La Reunion Island, that can be modelled as embedded eclogite in a matrix of spinel lherzolite, (3) The age of this submarine group and its location under the eastern flank of Piton de la Fournaise, imply a more complex model of evolution of La Reunion. Therefore, our geochemical and geochronological data are new arguments suggesting the existence of a third volcanic center on the island. This assumption confirms the previous petrological and geophysical evidence supporting the existence of this volcano at the East of La Reunion, commonly called Les Alizés volcano.