C. Albérola, C. Millot, and J. Et-font, On the seasonal and mesoscale variabilities of the Northern Current during the PRIMO-0 experiment in the western Mediterranean Sea, Oceanol. Acta, vol.18, issue.2, pp.163-192, 1995.

V. Andersen and L. Et-prieur, One-month study in the open NW Mediterranean Sea (DYNAPROC experiment): overview of the hydrobiogeochemical structures and effects of wind events, Deep-Sea Res. PT, vol.1, issue.3, pp.47-397, 1995.

D. Antoine, A. Morel, H. R. Gordon, V. F. Banzon, and R. H. Et-evans, Bridging ocean color observations of the 1980s and 2000s in search of long-term trends, Journal of Geophysical Research, vol.7, issue.15
DOI : 10.1029/2004JC002620

F. Auclair, S. Casitas, and P. Et-marsaleix, Application of an Inverse Method to Coastal Modeling, Journal of Atmospheric and Oceanic Technology, vol.17, issue.10, pp.1368-1391, 2000.
DOI : 10.1175/1520-0426(2000)017<1368:AOAIMT>2.0.CO;2

F. Auclair, P. Marsaleix, and P. Et-de-mey, Space-time structure and dynamics of the forecast error in a coastal circulation model of the Gulf of Lions, Dynamics of Atmospheres and Oceans, vol.36, issue.4, pp.309-346, 2003.
DOI : 10.1016/S0377-0265(02)00068-4

URL : https://hal.archives-ouvertes.fr/hal-00136965

B. Avril, DOC dynamics in the northwestern Mediterranean Sea (DYFAMED site), Deep Sea Research Part II: Topical Studies in Oceanography, vol.49, issue.11
DOI : 10.1016/S0967-0645(02)00033-4

N. Hoepffner, Variations in the light absorption coefficients of phytoplankton, nonalgal particles, and dissolved organic matter in coastal waters around Europe, J. Geophys

J. P. Béthoux, P. Morin, and D. P. Et-ruiz-pino, Temporal trends in nutrient ratios: chemical evidence of Mediterranean ecosystem changes driven by human activity, Deep Sea Research Part II: Topical Studies in Oceanography, vol.49, issue.11, p.49, 2002.
DOI : 10.1016/S0967-0645(02)00024-3

G. Billen and J. Et-garnier, River basin nutrient delivery to the coastal sea: Assessing its potential to sustain new production of non-siliceous algae, Marine Chemistry, vol.106, issue.1-2, pp.148-160, 2007.
DOI : 10.1016/j.marchem.2006.12.017

J. C. Et-orr, Potential impact of climate change on marine export production, Global Biogeochem. Cy, vol.15, issue.1, pp.81-99, 2001.

E. Bosc, A. Bricaud, and D. Et-antoine, Seasonal and interannual variability in algal biomass and primary production in the Mediterranean Sea, as derived from 4 years of SeaWiFS observations, Global Biogeochemical Cycles, vol.106, issue.3, p.18, 2004.
DOI : 10.1029/2003GB002034

H. L. Bryden, J. Candela, and T. H. Et-kinder, Exchange through the Strait of Gibraltar, Progress in Oceanography, vol.33, issue.3, pp.201-248, 1994.
DOI : 10.1016/0079-6611(94)90028-0

W. J. Cai, Sinks or Sites of Terrestrial Carbon Incineration?, Annual Review of Marine Science, vol.3, issue.1, pp.123-145, 2011.
DOI : 10.1146/annurev-marine-120709-142723

M. Canals, P. Puig, X. Durrieu-de-madron, S. Heussner, A. Palanques et al., Flushing submarine canyons, Nature, vol.47, issue.3, pp.354-357, 2006.
DOI : 10.1038/nature05271

C. Cassou, Du changement climatique aux régimes de temps : l'Oscillation Nord Atlantique, pp.21-32, 2004.

C. Cathalot, C. Rabouille, N. Tisnerat-laborde, P. Kerhervé, K. Bowles et al., Continental shelf particles: a major contributor of pre-aged organic carbon in deltaic areas, Nature, 2010.

A. Et-lebaron and P. , Dynamic Characteristics of Prochlorococcus and Synechococcus Consumption by Bacterivorous Nanoflagellates, Microbial Ecol, vol.43, issue.3, pp.341-352, 2002.

J. R. Christian, M. R. Lewis, and D. M. Et-karl, Vertical fluxes of carbon, nitrogen, and phosphorus in the North Pacific Subtropical Gyre near Hawaii, Journal of Geophysical Research: Oceans, vol.8, issue.16, pp.15667-15677, 1997.
DOI : 10.1029/97JC00369

J. R. Christian, Biogeochemical cycling in the oligotrophic ocean: Redfield and non-Redfield models, Limnology and Oceanography, vol.50, issue.2, pp.646-657, 2005.
DOI : 10.4319/lo.2005.50.2.0646

P. Conan and C. Et-millot, Variability of the northern current off Marseilles, western Mediterranean Sea : from february to june 1992, Oceanol. Acta, vol.18, issue.2, pp.193-205, 1995.

A. Conversi, S. Piontkovski, and S. Et-hameed, Seasonal and interannual dynamics of Calanus finmarchicus in the Gulf of Maine (Northeastern US shelf) with reference to the North Atlantic Oscillation, Deep-Sea Res, PT, vol.2, issue.48, pp.1-3, 2001.

G. Copin-montégut and B. Et-avril, Vertical distribution and temporal variation of dissolved organic carbon in the North-Western Mediterranean Sea, Deep Sea Research Part I: Oceanographic Research Papers, vol.40, issue.10, pp.40-1963, 1993.
DOI : 10.1016/0967-0637(93)90041-Z

B. Coste, Rôle des apports nutritifs minéraux rhodaniens sur la production organique des eaux du Golfe du Lion, pp.727-740, 1974.

M. J. Dagg, R. Benner, S. Lohrenz, and D. Et-lawrence, Transformation of dissolved and particulate materials on continental shelves influenced by large rivers: plume processes, Continental Shelf Research, vol.24, issue.7-8, pp.833-858, 2004.
DOI : 10.1016/j.csr.2004.02.003

F. Diaz, P. Raimbault, and P. Et-conan, Small-scale study of primary productivity during spring in a Mediterranean coastal area (Gulf of Lions), Continental Shelf Research, vol.20, issue.9, pp.975-996, 2000.
DOI : 10.1016/S0278-4343(00)00006-6

F. Diaz, P. Raimbault, B. Boudjellal, N. Garcia, and T. Et-moutin, Early spring phosphorus limitation of primary productivity in a NW Mediterranean coastal zone (Gulf of Lions), Marine Ecology Progress Series, vol.211
DOI : 10.3354/meps211051

F. Diaz, J. J. Naudin, C. Courties, P. Rimmelin, and L. Et-oriol, Biogeochemical and ecological functioning of the low-salinity water lenses in the region of the Rhone River freshwater influence, NW Mediterranean Sea, Continental Shelf Research, vol.28, issue.12, pp.28-1511, 2008.
DOI : 10.1016/j.csr.2007.08.009

E. M. Dickman, J. M. Newell, M. J. Gonzalez, and M. J. Vanni, Light, nutrients, and food-chain length constrain planktonic energy transfer efficiency across multiple trophic levels, Proceedings of the National Academy of Sciences, vol.105, issue.47, pp.105-18408, 2008.
DOI : 10.1073/pnas.0805566105

URL : http://www.ncbi.nlm.nih.gov/pmc/articles/PMC2587603

D. Ortenzio, F. Et-ribera-d-'alcala, and M. , On the trophic regimes of the Mediterranean Sea: a satellite analysis, Biogeosciences, vol.6, issue.2, pp.139-148, 2009.
DOI : 10.5194/bg-6-139-2009

K. Drinkwater, A. Belgrano, A. Borja, A. Conversi, and C. Et-greene, the response of marine ecosystems to climate variability associated with the North Atlantic Oscillation
DOI : 10.1029/134GM10

J. Hurrell, Y. Kushnir, G. Ottersen, and M. Visbeck, The North Atlantic Oscillation: climatic significance and environmental impact, pp.211-234, 2003.
DOI : 10.1029/GM134

H. W. Ducklow, D. K. Steinberg, and K. O. Et-buesseler, Upper Ocean Carbon Export and the Biological Pump, Oceanography, vol.14, issue.4, 2001.
DOI : 10.5670/oceanog.2001.06

C. Dufau-julliand, P. Marsaleix, A. Petrenko, and I. Et-dekeyser, Threedimensional modeling of the Gulf of Lion's hydrodynamics (northwest Mediterranean) during January 1999 (MOOGLI3 Experiment) and late winter 1999: Western Mediterranean Intermediate Water's (WIW's) formation and its cascading over the shelf break, J

R. Dugdale and J. Et-goering, UPTAKE OF NEW AND REGENERATED FORMS OF NITROGEN IN PRIMARY PRODUCTIVITY1, Limnology and Oceanography, vol.12, issue.2, pp.196-206, 1967.
DOI : 10.4319/lo.1967.12.2.0196

X. Durrieu-de-madron, B. Ferré, G. Le-corre, C. Grenz, P. Conan et al., Trawling-induced resuspension and dispersal of muddy sediments and dissolved elements in the Gulf of Lion (NW Mediterranean), Cont. Shelf
DOI : 10.1016/j.csr.2005.08.002

URL : https://hal.archives-ouvertes.fr/hal-00092784

R. W. Eppley and B. J. Et-peterson, Particulate organic matter flux and planktonic new production in the deep ocean, Nature, vol.20, issue.5740, pp.282-677, 1979.
DOI : 10.1007/BF00394908

C. Estournel, V. Kondrachoff, P. Marsaleix, and R. Vehil, The plume of the Rhone: numerical simulation and remote sensing, Continental Shelf Research, vol.17, issue.8, pp.899-924, 1997.
DOI : 10.1016/S0278-4343(96)00064-7

URL : https://hal.archives-ouvertes.fr/hal-00160813

C. Estournel, P. Broche, P. Marsaleix, J. L. Devenon, F. Auclair et al., The Rhone River Plume in Unsteady Conditions: Numerical and Experimental Results, Estuarine, Coastal and Shelf Science, vol.53, issue.1, pp.25-38, 2001.
DOI : 10.1006/ecss.2000.0685

URL : https://hal.archives-ouvertes.fr/hal-00160500

C. Estournel, X. Durrieu-de-madron, P. Marsaleix, F. Auclair, C. Julliand et al., Observation and modeling of the winter coastal oceanic circulation in the Gulf of Lion under wind conditions influenced by the continental orography (FETCH experiment), Journal of Geophysical Research, vol.6, issue.3
DOI : 10.1029/2001JC000825

P. G. Falkowski and C. S. Davis, Natural proportions, Nature, vol.431, issue.7005, pp.431-131, 2004.
DOI : 10.1073/pnas.0404222101

M. J. Fasham, H. W. Ducklow, and S. M. Et-mckelvie, A nitrogen-based model of plankton dynamics in the oceanic mixed layer, Journal of Marine Research, vol.48, issue.3, pp.591-639, 1990.
DOI : 10.1357/002224090784984678

M. Fieux, Formation d'eau dense sur le plateau continental du golfe du lion., La formation des eaux océaniques profondes, Colloques Internationaux du C, N.R.S, 1974.

J. M. Fromentin and B. Et-planque, Calanus and environment in the eastern North Atlantic. II. Influence of the North Atlantic Oscillation on C. finmarchicus and C. helgolandicus, Marine Ecology Progress Series, vol.134, pp.1-3, 1996.
DOI : 10.3354/meps134111

S. Gao and Y. P. Wang, Changes in material fluxes from the Changjiang River and their implications on the adjoining continental shelf ecosystem, Continental Shelf Research, vol.28, issue.12, pp.1490-1500, 2008.
DOI : 10.1016/j.csr.2007.02.010

C. Garcia-comas, L. Stemmann, F. Ibanez, L. Berline, M. G. Mazzocchi et al., Zooplankton long-term changes in the NW Mediterranean Sea: Decadal periodicity forced by winter hydrographic conditions related to large-scale atmospheric changes?, Journal of Marine Systems, vol.87, issue.3-4, pp.3-4, 2011.
DOI : 10.1016/j.jmarsys.2011.04.003

J. C. Gascard, Mediterranean deep water formation, baroclinic eddies and ocean eddies, Oceanol. Acta, vol.1, issue.3, pp.315-330, 1978.

J. Gatti, A. Petrenko, J. L. Devenon, Y. Leredde, and C. Et-ulses, The Rhone river dilution zone present in the northeastern, 2003.
URL : https://hal.archives-ouvertes.fr/hal-00138666

W. Gentleman, A. Leising, B. Frost, S. Strom, and J. Et-murray, Functional responses for zooplankton feeding on multiple resources: a review of assumptions and biological dynamics, Deep Sea Research Part II: Topical Studies in Oceanography, vol.50, issue.22-26, pp.22-26, 2003.
DOI : 10.1016/j.dsr2.2003.07.001

E. Goberville, G. Beaugrand, B. Sautour, P. Tréguer, S. Et et al., Climate-driven changes in coastal marine systems of western Europe, Marine Ecology Progress Series, vol.408, pp.129-148, 2010.
DOI : 10.3354/meps08564

URL : https://hal.archives-ouvertes.fr/hal-00634519

A. Goffart, J. H. Hecq, and L. Et-legendre, Changes in the development of the winter-spring phytoplankton bloom in the Bay of Calvi (NW Mediterranean) over the last two decades: a response to changing climate?, Marine Ecology Progress Series, vol.236, pp.45-60, 2002.
DOI : 10.3354/meps236045

F. Gomez and G. Et-gorsky, Annual microplankton cycles in Villefranche Bay, Ligurian Sea, NW Mediterranean, Journal of Plankton Research, vol.25, issue.4, pp.323-339, 2003.
DOI : 10.1093/plankt/25.4.323

URL : http://plankt.oxfordjournals.org/cgi/content/short/25/4/323

C. H. Greene, A. J. Pershing, A. Conversi, B. Planque, C. Hannah et al., Trans-Atlantic responses of Calanus finmarchicus populations to basin-scale forcing associated with the North Atlantic Oscillation, Progress in Oceanography, vol.58, issue.2-4, pp.2-4, 2003.
DOI : 10.1016/j.pocean.2003.08.009

M. Grégoire, K. Soetaert, N. Nezlin, and A. Et-kostianov, Modeling the nitrogen cycling and plankton productivity in the Black Sea using a three-dimensional interdisciplinary model, Journal of Geophysical Research, vol.11, issue.2, 2004.
DOI : 10.1029/2001JC001014

M. Guarracino, B. Barnier, P. Marsaleix, X. Durrieu-de-madron, A. Monaco et al., Transfer of particulate matter from the northwestern Mediterranean continental margin: Variability and controlling factors, Journal of Marine Research, vol.64, issue.2, pp.195-220, 2006.
DOI : 10.1357/002224006777606498

URL : https://hal.archives-ouvertes.fr/hal-00138670

S. Guerzoni, R. Chester, F. Dulac, B. Herut, M. D. Loÿe-pilot et al., The role of atmospheric deposition in the biogeochemistry of the Mediterranean Sea, Progress in Oceanography, vol.44, issue.1-3, pp.1-3, 1999.
DOI : 10.1016/S0079-6611(99)00024-5

T. B. Hallett, T. Coulson, J. G. Pilkington, T. H. Clutton-brock, J. M. Pemberton et al., Why large-scale climate indices seem to predict ecological processes better than local weather, Nature, vol.105, issue.6995, pp.430-71, 2004.
DOI : 10.1046/j.1365-2656.1999.00366.x

M. Herrmann, S. Somot, F. Sevault, C. Estournel, and M. Et-déqué, Modeling the deep convection in the northwestern Mediterranean Sea using an eddy-permitting and an eddy-resolving model: Case study of winter, J. Geophys. Res. -Oceans, issue.C4, p.113, 1986.
URL : https://hal.archives-ouvertes.fr/hal-00195049

M. Herrmann, Prix Prud'hommeLa formation d'eau dense en Méditerranée : interactions d'échelles, changement climatique et écosystèmes, La Météorologie, vol.70, pp.1-10, 2010.
DOI : 10.4267/2042/34573

L. Houpert, Etude du renouvellement des eaux profondes en Méditerranée nordoccidentale pendant la période, Rapport de Master 2 Océan Atmosphère Climat Observations Spatiales, 2007.

V. V. Ivanov, G. I. Shapiro, J. M. Huthnance, D. L. Aleynik, and P. N. Et-golovin, Cascades of dense water around the world ocean, Progress in Oceanography, vol.60, issue.1, pp.47-98, 2004.
DOI : 10.1016/j.pocean.2003.12.002

H. Jones and J. Et-marshall, Restratification after Deep Convection, Journal of Physical Oceanography, vol.27, issue.10, pp.2276-2287, 1997.
DOI : 10.1175/1520-0485(1997)027<2276:RADC>2.0.CO;2

C. A. Katsman, M. A. Spall, and R. S. Et-pickart, Boundary Current Eddies and Their Role in the Restratification of the Labrador Sea*, Journal of Physical Oceanography, vol.34, issue.9, pp.1967-1983, 2004.
DOI : 10.1175/1520-0485(2004)034<1967:BCEATR>2.0.CO;2

P. Kerhervé, M. Minagawa, S. Heussner, and A. Monaco, Stable isotopes (13C/12C and 15N/14N) insettling organic matter of the northwestern Mediterranean Sea: biogeochemical implications, Oceanologica Acta, vol.24, pp.24-77, 2001.
DOI : 10.1016/S0399-1784(00)01111-7

T. Kiorboe, Turbulence, Phytoplankton Cell Size, and the Structure of Pelagic Food Webs, Adv. Mar. Biol, vol.29, pp.1-72, 1993.
DOI : 10.1016/S0065-2881(08)60129-7

G. Krahmann, Saisonale und zwischenja¨hrliche Variabilita¨t im westlichen Mittelmeer, Ph.D. dissertation, Institut für Meere-skunde, 1997.

H. Lacombe, J. C. Gascard, J. Gonella, and J. P. Et-béthoux, Response of the Mediterranean to the water and energy fluxes across its surface, on seasonal and interannual scales, Oceanol. Acta, vol.4, issue.2, pp.247-255, 1981.

A. Lamy, C. Millot, and M. Et-molines, Bottom Pressure and Sea Level Measurements in the Gulf of Lions, Journal of Physical Oceanography, vol.11, issue.3, pp.394-410, 1981.
DOI : 10.1175/1520-0485(1981)011<0394:BPASLM>2.0.CO;2

K. D. Leaman, The formation of western Mediterranean Deep Water, Seasonal and Interannual Variability of the Western Mediterranean Sea, Coastal and Estuarine Studies, 1994.
DOI : 10.1029/CE046p0227

K. Leblanc, B. Quéginer, N. Garcia, P. Rimmelin, and P. Et-raimbault, Silicon cycle in the NW Mediterranean Sea: seasonal study of a coastal oligotrophic site, Oceanologica Acta, vol.26, issue.4, pp.339-355, 2003.
DOI : 10.1016/S0399-1784(03)00035-5

L. Legendre, Hydrodynamic control of marine phytoplankton : the paradox of stability, Ecohydrodynamics : Proceedings of the 12th International Liège Colloquium on Ocean Hydrodynamics, pp.191-207, 1981.

L. Legendre and F. Et-rassoulzadegan, Food-web mediated export of biogenic carbon in oceans:hydrodynamic control, Marine Ecology Progress Series, vol.145, pp.179-193, 1996.
DOI : 10.3354/meps145179

S. Legg and J. Et-marshall, The influence of the ambient flow on the spreading of convected water masses, Journal of Marine Research, vol.56, issue.1, pp.107-139, 1998.
DOI : 10.1357/002224098321836136

M. Lévy, L. Mémery, and G. Et-madec, The onset of the Spring Bloom in the MEDOC area: mesoscale spatial variability, Deep Sea Research Part I: Oceanographic Research Papers, vol.46, issue.7, pp.46-1137, 1999.
DOI : 10.1016/S0967-0637(98)00105-8

M. Lévy, P. Klein, and A. M. Et-tréguier, Impact of sub-mesoscale physics on production and subduction of phytoplankton in an oligotrophic regime, Journal of Marine Research, vol.59, issue.4, pp.535-565, 2001.
DOI : 10.1357/002224001762842181

M. Lévy, Mesoscale variability of phytoplankton and of new production: Impact of the large-scale nutrient distribution, Journal of Geophysical Research, vol.41, issue.C9, p.3358, 2003.
DOI : 10.1029/2002JC001577

P. Licandro, F. Ibanez, and M. Et-etienne, in the northwestern Mediterranean Sea: Relationships with hydroclimatic variability, Limnology and Oceanography, vol.51, issue.4, pp.1832-1848, 2006.
DOI : 10.4319/lo.2006.51.4.1832

P. Licandro and P. Et-icardi, Basin scale distribution of zooplankton in the Ligurian Sea (north-western Mediterranean) in late autumn, Hydrobiologia, vol.60, issue.61, pp.17-40, 2009.
DOI : 10.1007/s10750-008-9523-9

M. Livre-blanc-du and . Group, Marine Ecosystems response to climatic and anthropogenic forcings in the Mediterranean, Soumis à Progress in Oceanography, 2010.

S. E. Lohrenz, D. G. Redalje, W. J. Cai, J. Acker, and M. Dagg, A retrospective analysis of nutrients and phytoplankton productivity in the Mississippi River plume, Continental Shelf Research, vol.28, issue.12, pp.1466-1475, 2008.
DOI : 10.1016/j.csr.2007.06.019

W. Ludwig, E. Dumont, M. Meybeck, and S. Et-heussner, River discharges of water and nutrients to the Mediterranean and Black Sea: Major drivers for ecosystem changes during past and future decades?, Progress in Oceanography, vol.80, issue.3-4, pp.3-4, 2009.
DOI : 10.1016/j.pocean.2009.02.001

G. Madec, M. Chartier, and M. Et-crepon, The effect of thermohaline forcing variability on deep water formation in the western Mediterranean Sea: a high-resolution three-dimensional numerical study, Dynamics of Atmospheres and Oceans, vol.15, issue.3-5, pp.3-5, 1991.
DOI : 10.1016/0377-0265(91)90024-A

P. Marsaleix, C. Estournel, V. Kondrachoff, and R. Vehil, A numerical study of the formation of the Rh??ne River plume, Journal of Marine Systems, vol.14, issue.1-2, pp.99-115, 1998.
DOI : 10.1016/S0924-7963(97)00011-0

P. Marsaleix, F. Auclair, J. Floor, M. Herrmann, C. Estournel et al., Energy conservation issues in sigma-coordinate free-surface ocean models, Ocean Modelling, pp.61-89, 2008.

J. Marshall and F. Schott, Open-ocean convection: Observations, theory, and models, Reviews of Geophysics, vol.28, issue.20, pp.1-64, 1999.
DOI : 10.1029/98RG02739

URL : http://citeseerx.ist.psu.edu/viewdoc/summary?doi=10.1.1.467.7642

E. Martinez, D. Antoine, F. Ortenzio, and B. Et-gentili, Climate-Driven Basin-Scale Decadal Oscillations of Oceanic Phytoplankton, Science, vol.326, issue.5957, pp.326-1253, 2009.
DOI : 10.1126/science.1177012

J. C. Marty, E. Nicolas, J. C. Miquel, and S. W. Fowler, Particulate fluxes of organic compounds and their relationship to zooplankton fecal pellets in the northwestern Mediterranean Sea, Marine Chemistry, vol.46, issue.4, pp.387-405, 1994.
DOI : 10.1016/0304-4203(94)90034-5

J. C. Marty, J. Chiavérini, M. D. Pizay, and B. Et-avril, Seasonal and interannual dynamics of nutrients and phytoplankton pigments in the western Mediterranean Sea at the DYFAMED time-series station, Deep-Sea Res. PT, p.49, 1965.

J. C. Marty and J. Et-chiavérini, Hydrological changes in the Ligurian Sea (NW Mediterranean, DYFAMED site) during 1995???2007 and biogeochemical consequences, Biogeosciences, vol.7, issue.7, pp.2117-2128, 2010.
DOI : 10.5194/bg-7-2117-2010

D. Mcgill, A budget for dissolved nutrients salts in the Mediterranean Sea, Cahiers Océanographiques, pp.543-554, 1969.

T. D. Dickey, J. Mcneil, A. F. Michaels, and A. H. Et-knap, Influence of mesoscale eddies on new production in the Sargasso Sea, Nature, vol.394, pp.263-266, 1998.

C. Mertens and F. Schott, Interannual Variability of Deep-Water Formation in the Northwestern Mediterranean, Journal of Physical Oceanography, vol.28, issue.7, pp.1410-1424, 1998.
DOI : 10.1175/1520-0485(1998)028<1410:IVODWF>2.0.CO;2

C. Millot, Circulation in the Western Mediterranean Sea, Journal of Marine Systems, vol.20, issue.1-4, pp.423-442, 1999.
DOI : 10.1016/S0924-7963(98)00078-5

URL : https://hal.archives-ouvertes.fr/hal-00153954

C. Millot, J. Candela, J. L. Fuda, and Y. Et-tber, Large warming and salinification of the Mediterranean outflow due to changes in its composition, Deep Sea Research Part I: Oceanographic Research Papers, vol.53, issue.4, pp.656-666, 2006.
DOI : 10.1016/j.dsr.2005.12.017

J. C. Miquel, S. W. Fowler, J. Larosa, and P. Et-buatmenard, Dynamics of the downward flux of particles and carbon in the open northwestern Mediterranean Sea, Deep Sea Research Part I: Oceanographic Research Papers, vol.41, issue.2, pp.41-243, 1994.
DOI : 10.1016/0967-0637(94)90002-7

J. C. Molinero, F. Ibanez, S. Souissi, M. Chifflet, and P. Et-nival, Phenological changes in the Northwestern Mediterranean copepods Centropages typicus and Temora stylifera linked to climate forcing, Oecologia, vol.85, issue.Sciences de la, pp.640-649, 2005.
DOI : 10.1007/s00442-005-0130-4

J. C. Molinero, F. Ibanez, S. Souissi, E. Buecher, S. Dallot et al., Climate control on the long-term anomalous changes of zooplankton communities in the Northwestern Mediterranean, Global Change Biology, vol.26, issue.0, pp.11-26, 2008.
DOI : 10.3354/meps298021

. Mediterranean, Influence of mesoscale instabilities on the biomass and production of phytoplankton and bacterioplankton, Deep-Sea Res, PT, vol.1, issue.482, pp.405-437, 2001.

A. Morel, A. Bricaud, J. M. André, and J. Et-pelaez-udlet, Spatial/temporal evolution of the Rhône plume as seen by CZCS imagery-consequence upon the primary production in the Gulf of Lions, 1990.

A. Morel and J. M. Et-andré, Pigment distribution and primary production in the western Mediterranean as derived and modeled from coastal zone color scanner observations, Journal of Geophysical Research, vol.9, issue.C7, pp.12685-12698, 1991.
DOI : 10.1029/91JC00788

T. Moutin, P. Raimbault, H. Golterman, and B. Et-coste, The input of nutrients by the Rh??ne river into the Mediterranean Sea: Recent observations and comparison with earlier data, pp.373-374, 1998.
DOI : 10.1007/978-94-011-5266-2_19

T. Moutin, T. F. Thingstad, F. Van-wambeke, D. Marie, G. Slawyk et al., Does competition for nanomolar phosphate supply explain the predominance of the cyanobacterium Synechococcus?, Limnol. Oceanogr, issue.5, pp.47-1562, 2002.

T. Nagata and D. L. Et-kirchman, Release of dissolved organic matter by heterotrophic protozoa: Implications for microbial foodwebs, Arch. Hydrobiol, vol.35, pp.99-109, 1992.

J. J. Naudin, G. Cauwet, M. J. Chrétiennot-dinet, B. Deniaux, J. L. Devenon et al., River Discharge and Wind Influence Upon Particulate Transfer at the Land???Ocean Interaction: Case Study of the Rhone River Plume, Estuarine, Coastal and Shelf Science, vol.45, issue.3, pp.303-316, 1997.
DOI : 10.1006/ecss.1996.0190

J. J. Naudin, G. Cauwet, C. Fajon, L. Oriol, S. Terzic et al., Effect of mixing on microbial communities in the Rhone River plume, Journal of Marine Systems, vol.28, issue.3-4, pp.3-4, 2001.
DOI : 10.1016/S0924-7963(01)00004-5

K. Niewiadomska, H. Claustre, L. Prieur, and F. Et-d-'ortenzio, Submesoscale physical-biogeochemical coupling across the Ligurian current (northwestern Mediterranean) using a bio-optical glider, Limnology and Oceanography, vol.53, issue.5part2, pp.2210-2225, 2008.
DOI : 10.4319/lo.2008.53.5_part_2.2210

M. Pahlow and U. Et-riebesell, Temporal Trends in Deep Ocean Redfield Ratios, Science, vol.287, issue.5454, pp.287-831, 2000.
DOI : 10.1126/science.287.5454.831

I. Palomera, M. P. Olivar, J. Salat, A. Sabatés, M. Coll et al., Small pelagic fish in the NW Mediterranean Sea: An ecological review, Progress in Oceanography, vol.74, issue.2-3, pp.377-396, 2007.
DOI : 10.1016/j.pocean.2007.04.012

F. Et-jahoda and M. , Fin whales summering in the Ligurian Sea: distribution, encounter rate, mean group size and relation to physiographic variables, Journal of Cetacean Research and Management, vol.7, issue.2, pp.137-145, 2005.

K. Pearson, Mathematical contributions to the theory of evolution. III. Regression, heredity and panmixia, Philos. Trans. Royal Soc. London, Serie A, issue.187, pp.253-318, 1896.

A. Petrenko, Variability of circulation features in the Gulf of Lion NW Mediterranean Sea. Importance of inertial currents, Oceanologica Acta, vol.26, issue.4, pp.323-338, 2003.
DOI : 10.1016/S0399-1784(03)00038-0

S. Pinca and S. Et-dallot, Meso-and macrozooplankton composition patterns related to hydrodynamic structures in the Ligurian Sea (Trophos-2 experiment, 1986.

L. Polimene, N. Pinardi, M. Zavatarelli, and S. Et-colella, The Adriatic Sea ecosystem seasonal cycle: Validation of a three-dimensional numerical model, Journal of Geophysical Research, vol.41, issue.C1, 2006.
DOI : 10.1029/2005JC003260

L. Prieur and A. Et-sournia, Almofront-1): an interdisciplinary study of the Almeria-Oran geostrophic front, SW Mediterranean Sea, J. Marine Syst, vol.5, pp.3-5, 1991.

I. Puillat, I. Taupier-lepage, and C. Et-millot, Algerian Eddies lifetime can near 3 years, Journal of Marine Systems, vol.31, issue.4
DOI : 10.1016/S0924-7963(01)00056-2

URL : http://citeseerx.ist.psu.edu/viewdoc/summary?doi=10.1.1.502.598

M. Pujo-pay, P. Conan, F. Joux, L. Oriol, J. J. Naudin et al., Impact of phytoplankton and bacterial production on nutrient and DOM uptake in the Rh??ne River plume (NW Mediterranean), Marine Ecology Progress Series, vol.315, pp.43-54, 2006.
DOI : 10.3354/meps315043

Z. F. Qiu, A. M. Doglioli, Z. Y. Hu, P. Marsaleix, and F. Et-carlotti, The influence of hydrodynamic processes on zooplankton transport and distributions in the North Western Mediterranean: Estimates from a Lagrangian model, Ecological Modelling, vol.221, issue.23, pp.221-2816, 2010.
DOI : 10.1016/j.ecolmodel.2010.07.025

URL : https://hal.archives-ouvertes.fr/hal-00560868

R. Quadrelli, V. Pavan, and F. Et-molteni, Wintertime variability of Mediterranean precipitation and its links with large-scale circulation anomalies, Climate Dynamics, vol.17, issue.5-6, pp.5-6, 2001.
DOI : 10.1007/s003820000121

M. Ribera-d-'alcala, F. Conversano, F. Corato, P. Licandro, O. Mangoni et al., Seasonal patterns in plankton communities in a pluriannual time series at a coastal Mediterranean site, Gulf of Naples), Sci. Mar, pp.68-65, 2004.

M. Rixen, J. M. Beckers, S. Levitus, J. Antonov, T. Boyer et al., The Western Mediterranean Deep Water: A proxy for climate change, Geophysical Research Letters, vol.20, issue.3, p.12608, 2005.
DOI : 10.1007/s00382-003-0304-x

E. J. Rohling and H. L. Et-bryden, Man-Induced Salinity and Temperature Increases in Western Mediterranean Deep Water, Journal of Geophysical Research, vol.97, issue.C7, pp.11191-11198, 1992.
DOI : 10.1029/92JC00767

A. Rubio, V. Taillandier, and P. Garreau, Reconstruction of the Mediterranean northern current variability and associated cross-shelf transport in the Gulf of Lions from satellitetracked drifters and model outputs, J. Marine Syst, pp.78-63, 2009.

A. Sabates, J. Salat, and M. Et-maso, Spatial heterogeneity of fish larvae across a meandering current in the northwestern Mediterranean, Deep Sea Research Part I: Oceanographic Research Papers, vol.51, issue.4, pp.545-557, 2004.
DOI : 10.1016/j.dsr.2003.11.003

R. Santoleri, V. Banzon, S. Marullo, E. Napolitano, F. Ortenzio et al., Yearto-year variability of the phytoplankton bloom in the southern Adriatic Sea, 1998.

J. L. Sarmiento, R. Slater, R. Barber, L. Bopp, S. C. Doney et al., Response of ocean ecosystems to climate warming, Global Biogeochemical Cycles, vol.14, issue.4, 2004.
DOI : 10.1029/2003GB002134

G. Sarthou, K. R. Timmermans, S. Blain, and P. Et-tréguer, Growth physiology and fate of diatoms in the ocean: a review, Journal of Sea Research, vol.53, issue.1-2, pp.25-42, 2005.
DOI : 10.1016/j.seares.2004.01.007

URL : https://hal.archives-ouvertes.fr/hal-00456424

D. P. Schrag and R. B. Et-alley, PHYSICS: Ancient Lessons for Our Future Climate, Science, vol.306, issue.5697, pp.306-821, 2004.
DOI : 10.1126/science.1103025

U. Send, J. Font, and C. Et-mertens, Recent observation indicates convection' role in deep water circulation, Eos, Transactions American Geophysical Union, vol.75, issue.7, pp.61-65, 1996.
DOI : 10.1029/96EO00040

U. Siegenthaler and J. L. Et-sarmiento, Atmospheric carbon dioxide and the ocean, Nature, vol.365, issue.6442, pp.119-125, 1993.
DOI : 10.1038/365119a0

J. H. Simpson, Physical processes in the ROFI regime, Journal of Marine Systems, vol.12, issue.1-4, pp.3-15, 1997.
DOI : 10.1016/S0924-7963(96)00085-1

I. Siokou-frangou, U. Christaki, M. G. Mazzocchi, M. Montresor, M. Ribera-d-'alcala et al., Plankton in the open Mediterranean Sea: a review, Biogeosciences, vol.7, issue.5, pp.1543-1586, 2010.
DOI : 10.5194/bg-7-1543-2010

URL : https://hal.archives-ouvertes.fr/hal-00767891

N. Skliris and A. Et-lascaratos, Impacts of the Nile River damming on the thermohaline circulation and water mass characteristics of the Mediterranean Sea, Journal of Marine Systems, vol.52, issue.1-4, pp.1-4, 2004.
DOI : 10.1016/j.jmarsys.2004.02.005

F. Wulff, Humans, Hydrology, and the Distribution of Inorganic Nutrient Loading to the Ocean, Bioscience, vol.53, issue.3, pp.235-245, 2003.

A. Humborg, C. Eriksson, H. Et-hannerz, and F. , River Nutrient Loads and Catchment Size, Biogeochemistry, vol.75, issue.1, pp.83-107, 2005.

R. O. Smith, H. L. Bryden, and K. Stansfield, Observations of new western Mediterranean deep water formation using Argo floats 2004&ndash;2006, Ocean Science, vol.4, issue.2, pp.133-149, 2004.
DOI : 10.5194/os-4-133-2008

K. Soetaert, J. J. Middelburg, P. M. Herman, and K. Et-buis, On the coupling of benthic and pelagic biogeochemical models, Earth-Science Reviews, vol.51, issue.1-4, pp.1-4, 2000.
DOI : 10.1016/S0012-8252(00)00004-0

S. Somot, F. Sevault, and M. Et-déqué, Transient climate change scenario simulation of the Mediterranean Sea for the twenty-first century using a high-resolution ocean circulation model, Climate Dynamics, vol.66, issue.10, pp.851-879, 2006.
DOI : 10.1007/s00382-006-0167-z

A. Sournia, La production primaire planctonique en Méditerranée, Essai de mise à jour, N° Spécial, vol.5, 1973.

L. Stemmann, G. Gorsky, J. C. Marty, M. Picheral, and J. C. Et-miquel, Four-year study of large-particle vertical distribution (0?1000 m) in the NW Mediterranean in relation to hydrology, phytoplankton, and vertical flux, Deep-Sea Res. PT, vol.2, issue.11, pp.49-1243, 2002.

N. G. Yoccoz and B. Et-adlandsvik, Studying climate effects on ecology through the use of climate indices: the North Atlantic Oscillation, El Nino Southern Oscillation and beyond, P, Roy. Soc. Lond. B Bio, vol.270, pp.2087-2096, 1529.

H. U. Sverdrup, On Conditions for the Vernal Blooming of Phytoplankton, Journal du Conseil International pour l'Exploration de la Mer, pp.287-295, 1953.
DOI : 10.1093/icesjms/18.3.287

T. Tanaka and F. Et-rassoulzadegan, Full-depth profile (0?2000 m) of bacteria, heterotrophic nanoflagellates and ciliates in the NW Mediterranean Sea: Vertical partitioning of microbial trophic structures, Deep-Sea Res, PT, vol.2, issue.11, pp.49-2093, 2002.

I. Taupier-letage and C. Et-millot, Surface circulation in the Algerian basin during, Oceanol. Acta, vol.9, pp.79-85, 1984.

I. Taupier-letage, I. Puillat, C. Millot, and P. Et-raimbault, Biological response to mesoscale eddies in the Algerian Basin, Journal of Geophysical Research, vol.167, issue.34, p.3245, 2003.
DOI : 10.1029/1999JC000117

P. Testor and J. C. Et-gascard, Large-Scale Spreading of Deep Waters in the Western Mediterranean Sea by Submesoscale Coherent Eddies, Journal of Physical Oceanography, vol.33, issue.1, pp.75-87, 2003.
DOI : 10.1175/1520-0485(2003)033<0075:LSSODW>2.0.CO;2

P. Testor and J. C. Et-gascard, Post-convection spreading phase in the Northwestern Mediterranean Sea, Deep Sea Research Part I: Oceanographic Research Papers, vol.53, issue.5, pp.869-893, 2006.
DOI : 10.1016/j.dsr.2006.02.004

URL : https://hal.archives-ouvertes.fr/hal-00122309

T. F. Thingstad and F. Et-rassoulzadegan, Nutrient limitations, microbial food webs, and biological C-pumps: suggested interactions in a P-limited Mediterranean, Marine Ecology Progress Series, vol.117
DOI : 10.3354/meps117299

T. F. Thingstad, U. L. Zweifel, and F. Et-rassoulzadegan, P limitation of heterotrophic bacteria and phytoplankton in the northwest Mediterranean, Limnology and Oceanography, vol.43, issue.1, pp.88-94, 1998.
DOI : 10.4319/lo.1998.43.1.0088

T. F. Thingstad and F. Et-rassoulzadegan, Conceptual models for the biogeochemical role of the photic zone microbial food web, with particular reference to the Mediterranean Sea, Progress in Oceanography, vol.44, issue.1-3, pp.1-3, 1999.
DOI : 10.1016/S0079-6611(99)00029-4

H. Thomsen, Nitrate and phosphate contents of Mediterranean waters, Danish Oceanographic Expedition, 1931.

C. M. Turley, The changing Mediterranean Sea ??? a sensitive ecosystem?, Progress in Oceanography, vol.44, issue.1-3, pp.387-400, 1999.
DOI : 10.1016/S0079-6611(99)00033-6

M. H. Tusseau-vuillemin, L. Mortier, and C. Et-herbaut, Modeling nitrate fluxes in an open coastal environment (Gulf of Lions): Transport versus biogeochemical processes, Journal of Geophysical Research: Oceans, vol.5, issue.76
DOI : 10.1029/97JC03709

URL : https://hal.archives-ouvertes.fr/hal-00154210

C. Ulses, Dynamique océanique et transport de la matière particulaire dans le Golfe du Lion : crue, tempête et période hivernale, Thèse, 2005.

C. Ulses, C. Estournel, J. Bonnin, X. Durrieu-de-madron, and P. Et-marsaleix, Impact of storms and dense water cascading on shelf-slope exchanges in the Gulf of Lion (NW Mediterranean), Journal of Geophysical Research, vol.52, issue.9???11, p.113, 2008.
DOI : 10.1029/2006JC003795

URL : https://hal.archives-ouvertes.fr/hal-00562356

C. Ulses, 3D modelling of the NW Mediterranean biogeochemistry with a multinutrient and multi-plankton functional type model coupled to hydrodynamic model, 2011.

P. Vandromme, L. Stemmann, L. Berline, S. Gasparini, L. Mousseau et al., Zooplankton communities fluctuations from 1995 to 2005 in the Bay of Villefranche-sur-Mer, Biogeosciences Discuss, pp.9175-9207, 2010.

S. M. Vicente-serrano and J. I. Et-lopez-moreno, Nonstationary influence of the North Atlantic Oscillation on European precipitation, Journal of Geophysical Research, vol.111, issue.D17, p.113, 2008.
DOI : 10.1029/2008JD010382

M. Vichi, W. May, and A. Et-navarra, Response of a complex ecosystem model of the northern Adriatic Sea to a regional climate change scenario, Climate Research, vol.24, issue.2, pp.141-159, 2003.
DOI : 10.3354/cr024141

J. D. Warren, D. A. Demer, D. E. Mcgehee, D. Ment, R. Et-borsani et al., Zooplankton in the Ligurian Sea: Part II. Exploration of their physical and biological forcing functions during summer 2000, Journal of Plankton Research, vol.26, issue.12, pp.1419-1427, 2000.
DOI : 10.1093/plankt/fbh129

M. J. Zhou, K. L. Shen, and R. C. Et-yu, Responses of a coastal phytoplankton community to increased nutrient input from the Changjiang (Yangtze) River, Continental Shelf Research, vol.28, issue.12, pp.1483-1489, 2008.
DOI : 10.1016/j.csr.2007.02.009

C. Figure, Moyennes annuelles de température (°C) dans la couche de surface (0-100 m) de 1976 à Les lignes horizontales pointillées correspondent aux moyennes sur les souspériodes 1976, 1989.